Supplementary material accompanying

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Electronic Supplementary Material (on-line archive) accompanying:
Index of Alien Impact (IAI): A method for evaluating potential ecological impact of alien plant
species
Appendix 1. Ecological Traits for 61 Alien and 2 Cryptogenic Species.
Electronic Supplement - Index of Alien Impact – Appendix 1
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Appendix 1. Ecological Traits for 61 Alien and 2 Cryptogenic Species.
Table A1 References documenting which of 24 ecological traits (listed in Table 1 in the journal
article) were present for each of 61 alien and 2 cryptogenic* species found along 35 wadeable streams
of the John Day River basin. Taxonomy follows the PLANTS database (USDA and NRCS 2007).
References are listed by first author and publication year; full citations are listed alphabetically
following this table. For detailed descriptions of ecological traits, see Table 1 in the journal article.
Traits
Species names and references documenting the presence of
specific ecological traits associated with each species.
Acroptilon repens (L.) DC.
Strongly clonal
Large propagule crop
Beck 2002, Carpenter 1998a, Cranston 2002, CWMA 2002,
Hickman 1993, Parker 1972, Prather 2002, SAFRR 2000,
Taylor 1990, UNR 2002, Watson 1980, Whitson 1999, Zouhar
2001a
Carpenter 1998a, Watson 1980, Zouhar 2001a
Small seeds/fruits
Drought tolerant
Carpenter 1998a, Hickman 1993, Parker 1972, SAFRR 2000,
Watson 1980, Zouhar 2001a
Beck 2002, Carpenter 1998a, Taylor 1990, Watson 1980
Wide moisture regime
Beck 2002, Carpenter 1998a, CWMA 2002, Parker 1972,
Taylor 1990, UNR 2002, Zouhar 2001a
Flooding/saturation tolerant
Alkaline or saline tolerant
Grazing tolerant or increaser
Increases post-fire
Beck 2002, Carpenter 1998a, CWMA 2002, Parker 1972, UNR
2002, Zouhar 2001a
Beck 2002, Zouhar 2001a
Carpenter 1998a, Parker 1972, Zouhar 2001a
Carpenter 1998a, Zouhar 2001a
Excretes salts or toxins
Beck 2002, Carpenter 1998a, Stermitz 2003, Watson 1980,
Zouhar 2001a
Forms monocultures or near
monocultures
Beck 2002, Carpenter 1998a, Cranston 2002, SAFRR 2000,
UNR 2002, Watson 1980, Whitson 1999, Zouhar 2001a
Invades in absence of human
disturbance
Carpenter 1998a
Agrostis capillaris L.
Strongly clonal
Large propagule crop
Small seeds/fruits
Clampitt 1987, Cook 1998, Hickman 1993, Hill (no date),
Hitchcock 1969, Peat 2005, USDA-NRCS 2004, Weber 2003,
Whitson 1999
Boedeltje 2003, Peat 2005, USDA-NRCS 2004
Grime 1981, Hickman 1993, Hitchcock 1969, Peat 2005
Dispersal over time
Grime 1979, Pakeman 1998, Peat 2005, Thompson 1993, Xiong
2001
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Traits
Species names and references documenting the presence of
specific ecological traits associated with each species.
Plasticity
Drought tolerant
Wide moisture regime
Batson 1998, Cook 1998, Grime 1979, Hitchcock 1969, USDANRCS 2004, Weber 2003
Cook 1998, Jackson 1982
Cook 1998, USDA-NRCS 2004
Flooding/saturation tolerant
Alkaline or saline tolerant
Batson 1998, Hill (no date), Taylor 1983, USDA-NRCS 2004
Peat 2005, USDA-NRCS 2004
Wide nutrient or soil texture
ranges
Cook 1998, Hill (no date), Tilman 1999, Weber 2003
Grazing tolerant or increaser
Increases post-fire
Alters soil stability
Grime 1975, Grime 1979, Hill (no date), Hellström 2003,
Valassis 1957
USDA-NRCS 2004, Wilson 1999
Weber 2003
Forms monocultures or near
monocultures
Hill (no date), Jackson 1982, Tilman 1999, USDA-NRCS 2004,
Weber 2003
Invades in absence of human
disturbance
Clampitt 1987, Jackson 1982
Agrostis gigantea Roth
Strongly clonal
Large propagule crop
Small seeds/fruits
Dispersal over time
Plasticity
Wide moisture regime
Carey 1995, Casler 2003, Hickman 1993, Peat 2005, Taylor
1990, USDA-NRCS 2002a, USDA-NRCS 2004
Peat 2005, USDA-NRCS 2004
Grime 1981, Hickman 1993, USDA-NRCS 2002a
Carey 1995, Oosting 1940, Peat 2005
Casler 2003, USDA-NRCS 2004
Carey 1995, USDA-NRCS 2002a, USDA-NRCS 2004
Flooding/saturation tolerant
Wide light regime
Alkaline or saline tolerant
Wide nutrient or soil texture
Grazing tolerant or increaser
Increases post-fire
Alters soil stability
Carey 1995, Frenkel 1978, Leege 1981, USDA-NRCS 2004
Carey 1995
Casler 2003, Frenkel 1978
Carey 1995, USDA-NRCS 2002a, USDA-NRCS 2004
Carey 1995, Clary 1995, Leege 1981
Carey 1995, Swan 1970, USDA-NRCS 2004
Carey 1995, Casler 2003, USDA-NRCS 2002a
Forms monocultures or near
monocultures
Carey 1995
Invades in absence of human
disturbance
Frenkel 1978
Alopecurus pratensis L.
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Traits
Species names and references documenting the presence of
specific ecological traits associated with each species.
Strongly clonal
Large propagule crop
Small seeds/fruits
Plasticity
Drought tolerant
Wide moisture regime
Flooding/saturation tolerant
Wide light regime
Alkaline or saline tolerant
Wide nutrient or soil texture
Grazing tolerant or increaser
Crowe 1997, Hitchcock 1969, PU 2002, USDA-NRCS 2004
Boedeltje 2003, USDA-NRCS 2004
Hitchcock 1969, PU 2002
PU 2002
Morisawa 1999, PU 2002, USDA-NRCS 2004
Morisawa 1999, PU 2002, USDA-NRCS 2004
Clary 1995, Hitchcock 1969, Morisawa 1999, PU 2002
Morisawa 1999, PU 2002
Morisawa 1999, PU 2002
Morisawa 1999, PU 2002
Clary 1995, Morisawa 1999
Forms monocultures or near
monocultures
Small seeds/fruits
Animal dispersal
Drought tolerant
Wide nutrient or soil texture
PU 2002
Alyssum alyssoides (L.) L.
Hickman 1993, Hitchcock 1969, Peat 2005
Peat 2005
Hitchcock 1969, Whitson 1999
Peat 2005
Anthriscus caucalis Bieb.
Large propagule crop
Small seeds/fruits
Animal dispersal
Plasticity
Wide moisture regime
Flooding/saturation tolerant
Wide light regime
Wide nutrient or soil texture
Hawkes 1989, Hickman 1993, Hickman 1993, Hitchcock 1969,
Peat 2005
Peat 2005
Hawkes 1989, Hitchcock 1969, Peat 2005, Whitson 1999
Hawkes 1989, Hitchcock 1969
Hickman 1993
Wilson 1999
Arctium minus Bernh.
Gross 1980
Hickman 1993
Gross 1980, Taylor 1990, Whitson 1999
Gross 1980
Gross 1980
Gross 1980
Gross 1980
Gross 1980
Forms monocultures or near
monocultures
Gross 1980
Small seeds/fruits
Dispersal over time
Animal dispersal
Flooding/saturation tolerant
Wide light regime
Increases post-fire
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Traits
Species names and references documenting the presence of
specific ecological traits associated with each species.
Strongly clonal
Large propagule crop
Small seeds/fruits
Specialized dispersal
Water dispersal
Arrhenatherum elatius (L.) Beauv. ex J.& K. Presl
Wilson 1998, Wilson 2001
Boedeltje 2003, Grime 1979, USDA-NRCS 2004
Weber 2003, Weiller 1995Weber 2003, Whitson 1999
Boedeltje 2003
Plasticity
Drought tolerant
Wide light regime
Wide nutrient or soil texture
Grazing tolerant or increaser
Increases post-fire
Alters soil stability
FAO 2005, Hickman 1993, USDA-NRCS 2004, Weber 2003,
Wilson 2001
FAO 2005, Weiller 1995USDA-NRCS 2004
FAO 2005, USDA-NRCS 2002b, USDA-NRCS 2004
Grime 1979
USDA-NRCS 2004
Hickman 1993, USDA-NRCS 2002b, Weber 2003
Forms monocultures or near
monocultures
Weber 2003, Wilson 1998, Wilson 2001
Invades in absence of human
disturbance
Drought tolerant
Wide nutrient or soil texture
Grazing tolerant or increaser
Large propagule crop
Animal dispersal
Drought tolerant
Wide moisture regime
Flooding/saturation tolerant
Wide nutrient or soil texture
Grazing tolerant or increaser
Increases post-fire
Alters hydrology
Alters nutrient cycling
Strongly clonal
Large propagule crop
Animal dispersal
Weber 2003, Wilson 2001
Bromus briziformis Fisch. & C.A. Mey.
Robocker 1973, USDA-NRCS 2004
USDA-NRCS 2004
Hitchcock 1969, Taylor 1990
Bromus racemosus L.
Howard 1994
Howard 1994
FAO 2005, Howard 1994, Howard 1998
Howard 1994
Howard 1994
FAO 2005, Howard 1994, Howard 1998
Green 1995, Howard 1994, Howard 1998
Howard 1998
Howard 1994
Howard 1994
Bromus inermis Leyss. ssp. inermis
Bush 2002a, Hickman 1993, Howard 1996, Sather 1987,
USDA-NRCS 2004, Weber 2003
Sather 1987, USDA-NRCS 2004
Sather 1987
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Traits
Species names and references documenting the presence of
specific ecological traits associated with each species.
Plasticity
Howard 1996, Sather 1987, USDA-NRCS 2004, Weber 2003
Drought tolerant
Wide moisture regime
Flooding/saturation tolerant
Alkaline or saline tolerant
Grazing tolerant or increaser
Increases post-fire
Alters soil stability
Bush 2002a, Howard 1996, Sather 1987, USDA-NRCS 2004
Howard 1996, Sather 1987, USDA-NRCS 2004
Howard 1996, Sather 1987
Howard 1996, Sather 1987, Weber 2003
Howard 1996
Howard 1996, Sather 1987, USDA-NRCS 2004
Bush 2002a, Hickman 1993, Howard 1996
Forms monocultures or near
monocultures
Large propagule crop
Dispersal over time
Animal dispersal
Plasticity
Bush 2002a, Howard 1996, Sather 1987, USDA-NRCS 2004,
Weber 2003
Bromus sterilis L.
Cooper 2000, Kleeman 2004, Kon 1988
Cooper 2000, Kleeman 2004
Clapham 1987, Cooper 2000, Kleeman 2004
Kleeman 2004, Kon 1988
Drought tolerant
Wide moisture regime
Wide light regime
Wide nutrient or soil texture
Grazing tolerant or increaser
Alters hydrology
Alters nutrient cycling
Clapham 1987, Cooper 2000, Kie 1988, Kleeman 2004, PFAF
1997-2001, YCRCD 1997-2004
UCIPM 2004
Theaker 1995
Cooper 2000, Kleeman 2004, PFAF 1997-2001
Grime 1979
Kleeman 2004
Cooper 2000, Dastgheib 2003, Kleeman 2004
Forms monocultures or near
monocultures
Clapham 1987, Kie 1988
Bromus tectorum L.
Large propagule crop
Dispersal over time
Wind dispersal
Carpenter 1999, Evans 2001, Pellant 1996, Upadhyaya 1986,
Weber 2003, Young 2000, Zouhar 2003a
Pellant 1996
APRS 2000, Carpenter 1999, Upadhyaya 1986, Young 2000
Animal dispersal
APRS 2000, Carpenter 1999, Neese 2000, Upadhyaya 1986,
Young 2000
Plasticity
Carpenter 1999, Evans 2001, Pellant 1996, Upadhyaya 1986,
Weber 2003
Drought tolerant
APRS 2000, Carpenter 1999, Evans 2001, Neese 2000, Pellant
1996, Upadhyaya 1986, Weber 2003, Young 2000
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Traits
Species names and references documenting the presence of
specific ecological traits associated with each species.
Wide nutrient or soil texture
APRS 2000, Carpenter 1999, Upadhyaya 1986, Zouhar 2003a
Grazing tolerant or increaser
Carpenter 1999, Neese 2000, Taylor 1990, Upadhyaya 1986,
Zouhar 2003a
Increases post-fire
APRS 2000, Carpenter 1999, Neese 2000, Pellant 1996,
Upadhyaya 1986, Young 2000, Zouhar 2003a
Alters hydrology
Alters nutrient cycling
APRS 2000, Carpenter 1999, Evans 2001, Neese 2000, Pellant
1996, Upadhyaya 1986, Weber 2003, Whitson 1999, Young
2000, Zouhar 2003a
Carpenter 1999, Evans 2001, Pellant 1996
Alters fire regime
Alters soil stability
Carpenter 1999, Evans 2001, Neese 2000, Pellant 1996,
Upadhyaya 1986, Weber 2003, Whitson 1999, Young 2000,
Zouhar 2003a
Carpenter 1999, Pellant 1996, Upadhyaya 1986
Forms monocultures or near
monocultures
Carpenter 1999, Evans 2001, Neese 2000, Pellant 1996,
Upadhyaya 1986, Young 2000, Zouhar 2003a
Invades in absence of human
disturbance
Carpenter 1999, Evans 2001, Pellant 1996, Upadhyaya 1986,
Weber 2003
Centaurea solstitialis L.
Large propagule crop
DiTomaso 2000, DiTomaso 2001, Gerlach 2000, Sheley 1999,
Weber 2003, Zouhar 2002a
Small seeds/fruits
DiTomaso 2000, DiTomaso 2001, Gerlach 2000, Hickman
1993, Prather 2002, Weber 2003, Zouhar 2002a
Dispersal over time
DiTomaso 2000, DiTomaso 2001, Gerlach 2000, Zouhar 2002a
Wind dispersal
DiTomaso 2000, DiTomaso 2001, Sheley 1999, Weber 2003,
Zouhar 2002a
Animal dispersal
Plasticity
DiTomaso 2000, DiTomaso 2001, Gerlach 2000, Sheley 1999,
Zouhar 2002a
Weber 2003, Zouhar 2002a
Drought tolerant
Grazing tolerant or increaser
Increases post-fire
DiTomaso 1999, DiTomaso 2000, DiTomaso 2001, Gerlach
2000, Prather 2002, Sheley 1999, Weber 2003, Zouhar 2002a
DiTomaso 2000, DiTomaso 2001, Gerlach 2000
DiTomaso 1999, DiTomaso 2001, Zouhar 2002a
Alters hydrology
Alters fire regime
DiTomaso 2000, DiTomaso 2001, Gerlach 2000, Weber 2003,
Zouhar 2002a
Zouhar 2002a
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Traits
Species names and references documenting the presence of
specific ecological traits associated with each species.
Excretes salts or toxins
Zouhar 2002a
Forms monocultures or near
monocultures
DiTomaso 2001, Gerlach 2000, Sheley 1999, Weber 2003,
Zouhar 2002a
Invades in absence of human
disturbance
Strongly clonal
Large propagule crop
Small seeds/fruits
Dispersal over time
Plasticity
Drought tolerant
Wide moisture regime
Flooding/saturation tolerant
Wide nutrient or soil texture
Grazing tolerant or increaser
Invades in absence of human
disturbance
DiTomaso 1999, DiTomaso 2001, Weber 2003, Zouhar 2002a
Centaurea L.
Cerastium fontanum Baumg. ssp. vulgare (Hartman) Greuter &
Burdet
Hickman 1993, Hitchcock 1969, Peat 2005, Taylor 1990
Boedeltje 2003, Peat 2005
Hickman 1993, Hitchcock 1969, Peat 2005
Peat 2005
Hitchcock 1969
Peat 2005
Peat 2005
Hickman 1993
Peat 2005
Hellström 2003, Pakeman 2004
Ryan 2003
Cirsium arvense (L.) Scop.
Strongly clonal
APRS 2000, Bayer 2000, Crowe 1997, Hickman 1993, Moore
1975, Nuzzo 1997, Prather 2002, Stritzke (no date), Taylor
1990, Weber 2003, Whitson 1999, Zouhar 2001b
Large propagule crop
APRS 2000, Bayer 2000, Moore 1975, Nuzzo 1997, Weber
2003, Zouhar 2001b
Small seeds/fruits
Dispersal over time
APRS 2000, Bayer 2000, Hickman 1993, Moore 1975, Nuzzo
1997, Prather 2002, Weber 2003, Whitson 1999
Bayer 2000, Moore 1975, Nuzzo 1997
Wind dispersal
Animal dispersal
Andersen 1995, APRS 2000, Bayer 2000, Hickman 1993,
Moore 1975, Nuzzo 1997, Prather 2002, Taylor 1990, Weber
2003, Zouhar 2001b
Andersen 1995
Water dispersal
Plasticity
Drought tolerant
APRS 2000, Bayer 2000, Boedeltje 2003, Moore 1975, Nuzzo
1997, Zouhar 2001b
Bayer 2000, Moore 1975, Nuzzo 1997, Weber 2003
APRS 2000, Bayer 2000, Moore 1975, Prather 2002
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Traits
Species names and references documenting the presence of
specific ecological traits associated with each species.
Wide moisture regime
APRS 2000, Bayer 2000, Moore 1975, Nuzzo 1997, Prather
2002, Zouhar 2001b
Flooding/saturation tolerant
Alkaline or saline tolerant
Wide nutrient or soil texture
Grazing tolerant or increaser
Increases post-fire
Alters hydrology
Alters nutrient cycling
Excretes salts or toxins
APRS 2000, Bayer 2000, Nuzzo 1997, Prather 2002, Weber
2003, Zouhar 2001b
APRS 2000, Moore 1975, Nuzzo 1997
APRS 2000, Nuzzo 1997, Zouhar 2001b
Bayer 2000, Crowe 1997, Nuzzo 1997, Zouhar 2001b
Zouhar 2001b
Bayer 2000, Moore 1975
Bayer 2000, Moore 1975
APRS 2000, Bayer 2000, Moore 1975, Nuzzo 1997
Forms monocultures or near
monocultures
Bayer 2000, Moore 1975, Nuzzo 1997, Prather 2002, Weber
2003
Invades in absence of human
disturbance
Large propagule crop
APRS 2000, Bayer 2000, Nuzzo 1997, Weber 2003, Zouhar
2001b
Cirsium vulgare (Savi) Ten.
APRS 2000, Beck 1999, Randall 2000, Zouhar 2002b
Small seeds/fruits
Beck 1999, Hickman 1993, Randall 2000, Weber 2003, Zouhar
2002b
Wind dispersal
Animal dispersal
Water dispersal
Drought tolerant
Wide moisture regime
Alkaline or saline tolerant
Wide nutrient or soil texture
APRS 2000, Beck 1999, Hickman 1993, Randall 2000, Taylor
1990, Weber 2003, Zouhar 2002b
Beck 1999
Beck 1999, Weber 2003
Beck 1999, Weber 2003, Zouhar 2002b
Beck 1999, Zouhar 2002b
Randall 2000, Zouhar 2002b
Beck 1999, Zouhar 2002b
Grazing tolerant or increaser
Increases post-fire
Alters hydrology
Alters nutrient cycling
Beck 1999, Bullock 1994, Randall 2000, Medlin (no date),
Zouhar 2002b
Zouhar 2002b, Zouhar 2002c
Randall 2000
Randall 2000, Zouhar 2002b
Forms monocultures or near
monocultures
Randall 2000, Weber 2003
Invades in absence of human
disturbance
Large propagule crop
Randall 2000, Weber 2003, Zouhar 2002b
Cynoglossum officinale L.
APRS 2000, Upadhyaya 1988, Zouhar 2002c
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Traits
Species names and references documenting the presence of
specific ecological traits associated with each species.
Specialized dispersal
Upadhyaya 1988, Zouhar 2002c
Animal dispersal
Drought tolerant
Wide moisture regime
Wide light regime
Wide nutrient or soil texture
Grazing tolerant or increaser
Alters hydrology
Alters nutrient cycling
APRS 2000, Taylor 1990, Upadhyaya 1988, Whitson 1999,
Zouhar 2002c
APRS 2000, Upadhyaya 1988, Zouhar 2002c
Zouhar 2002c
Zouhar 2002c
Upadhyaya 1988
APRS 2000, Upadhyaya 1988, Zouhar 2002c
APRS 2000, Upadhyaya 1988
APRS 2000, Upadhyaya 1988
Forms monocultures or near
monocultures
APRS 2000, Upadhyaya 1988, Zouhar 2002c
Invades in absence of human
disturbance
Strongly clonal
Large propagule crop
Small seeds/fruits
Wind dispersal
Animal dispersal
Water dispersal
Zouhar 2002c
Dactylis glomerata L.
Hickman 1993, Sullivan 1992a
Boedeltje 2003, USDA-NRCS 2004, Weber 2003
Bush 2000, Bush 2003, Weber 2003
Weber 2003
Weber 2003
Boedeltje 2003, Weber 2003
Plasticity
Wide moisture regime
Flooding/saturation tolerant
Wide light regime
Wide nutrient or soil texture
Grazing tolerant or increaser
Increases post-fire
Alters soil stability
Bush 2000, Bush 2003, Grime 1979, Haubensak 1999a, Sullivan
1992a, USDA-NRCS 2004, Weber 2003
Bush 2000, Bush 2003, USDA-NRCS 2004
Weber 2003
Sullivan 1992a, USDA-NRCS 2004
Haubensak 1999a
Grime 1979, Lacey 2002, Sullivan 1992a
Sullivan 1992a, USDA-NRCS 2004
Bush 2000, Bush 2003, Sullivan 1992a
Forms monocultures or near
monocultures
Large propagule crop
Small seeds/fruits
Dispersal over time
Specialized dispersal
Water dispersal
Bush 2000, Bush 2003, Weber 2003
Dipsacus fullonum L.
Weber 2003, Werner 1975
Weber 2003, Werner 1975
Werner 1975
Werner 1975
Weber 2003, Werner 1975
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Traits
Species names and references documenting the presence of
specific ecological traits associated with each species.
Wide moisture regime
Flooding/saturation tolerant
Werner 1975
Weber 2003, Werner 1975, Whitson 1999, Howard 1992
Forms monocultures or near
monocultures
Weber 2003, Werner 1975
Invades in absence of human
disturbance
Dispersal over time
Specialized dispersal
Animal dispersal
Plasticity
Weber 2003
Erodium cicutarium (L.) L'Hér. ex Ait.
Howard 1992, Schiffman 2004
APRS 2000, Hawkes 1989, Howard 1992, Schiffman 2004
Howard 1992, Schiffman 2004, Taylor 1990
Hawkes 1989, Weber 2003
Drought tolerant
Wide moisture regime
Wide light regime
Grazing tolerant or increaser
Increases post-fire
APRS 2000, Hawkes 1989, Howard 1992, Schiffman 2004,
Taylor 1990, Weber 2003
Howard 1992, Schiffman 2004
Howard 1992, Schiffman 2004
Howard 1992, Schiffman 2004
Howard 1992, Schiffman 2004
Forms monocultures or near
monocultures
Hawkes 1989, Schiffman 2004, Weber 2003
Invades in absence of human
disturbance
Schiffman 2004, Weber 2003
Euphorbia esula L.
Strongly clonal
Anonymous 2005, APRS 2000, Best 1980, Biesboer 1996,
Kreps 2000, Prather 2002, Simonin 2000, Taylor 1990, Weber
2003, Whitson 1999
Large propagule crop
Anonymous 2005, APRS 2000, Best 1980, Biesboer 1996,
Taylor 1990
Small seeds/fruits
Dispersal over time
APRS 2000, Best 1980, Hickman 1993, Hitchcock 1969, Kreps
2000, Simonin 2000, Weber 2003
Kreps 2000, Simonin 2000
Specialized dispersal
Anonymous 2005, APRS 2000, Best 1980, Biesboer 1996,
Kreps 2000, Prather 2002, Simonin 2000, Whitson 1999
Animal dispersal
Water dispersal
Plasticity
Anonymous 2005, APRS 2000, Biesboer 1996, Kreps 2000,
Simonin 2000
Biesboer 1996, Simonin 2000
APRS 2000, Weber 2003
Drought tolerant
Anonymous 2005, APRS 2000, Best 1980, Biesboer 1996,
Kreps 2000, Prather 2002, Taylor 1990, Weber 2003
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Traits
Species names and references documenting the presence of
specific ecological traits associated with each species.
Wide moisture regime
Anonymous 2005, APRS 2000, Best 1980, Biesboer 1996,
Kreps 2000, Prather 2002
Flooding/saturation tolerant
Wide light regime
Wide nutrient or soil texture
APRS 2000, Best 1980, Biesboer 1996, Kreps 2000, Prather
2002
Anonymous 2005, APRS 2000, Best 1980, Kreps 2000
APRS 2000, Biesboer 1996
Grazing tolerant or increaser
Increases post-fire
Alters hydrology
Alters nutrient cycling
Alters fire regime
Alters soil stability
Anonymous 2005, Best 1980, Hirsch 1998, Kreps 2000, Prather
2002
APRS 2000, Simonin 2000
Anonymous 2005, Kreps 2000
Anonymous 2005
Simonin 2000
Hirsch 1998
Excretes salts or toxins
Anonymous 2005, APRS 2000, Best 1980, Biesboer 1996,
Kreps 2000
Forms monocultures or near
monocultures
Anonymous 2005, APRS 2000, Best 1980, Biesboer 1996,
Hirsch 1998, Kreps 2000, Weber 2003
Invades in absence of human
disturbance
Anonymous 2005, Best 1980, Biesboer 1996, Hirsch 1998,
Kreps 2000, Simonin 2000, Weber 2003
Festuca rubra L.*
Strongly clonal
Large propagule crop
Plasticity
Drought tolerant
Wide moisture regime
Flooding/saturation tolerant
Wide light regime
Crowe 1997, Hickman 1993, USDA-NRCS 2004, Walsh 1995a
USDA-NRCS 2004
Hickman 1993, USDA-NRCS 2004, Walsh 1995a
USDA-NRCS 2004, Walsh 1995a
Frenkel 1978, USDA-NRCS 2004, Walsh 1995a
Campbell 1988, Frenkel 1978, Taylor 1983, Walsh 1995a
USDA-NRCS 2002c, USDA-NRCS 2004
Alkaline or saline tolerant
Wide nutrient or soil texture
Grazing tolerant or increaser
Increases post-fire
Alters soil stability
Excretes salts or toxins
Campbell 1988, Frenkel 1978, Hickman 1993, Taylor 1983,
USDA-NRCS 2004, Walsh 1995a
Fransen 2001, USDA-NRCS 2004, Walsh 1995a
Fransen 2001, Grime 1979, Hellström 2003, Walsh 1995a
Walsh 1995a, Wilson 1999, Wilson 1999
Crowe 1997, USDA-NRCS 2002c, Walsh 1995a
Walsh 1995a
Invades in absence of human
disturbance
Frenkel 1978
Holcus lanatus L.
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Traits
Species names and references documenting the presence of
specific ecological traits associated with each species.
Strongly clonal
Large propagule crop
Small seeds/fruits
Dispersal over time
Wind dispersal
Plasticity
Hawkes 1989, Pitcher 1988, Thompson 1988
Pitcher 1988, Thompson 1988, Weber 2003
Hickman 1993, Thompson 1988, USDA-NRCS 2004
Grime 1979, Pitcher 1988, Thompson 1988
Thompson 1988, Weber 2003
Grime 1979, Thompson 1988
Wide moisture regime
Flooding/saturation tolerant
Wide light regime
Alkaline or saline tolerant
Cook 1998, Pitcher 1988, Thompson 1988, USDA-NRCS 2004
Pitcher 1988, Taylor 1983, Thompson 1988, Weber 2003
Pitcher 1988, USDA-NRCS 2004
Thompson 1988
Wide nutrient or soil texture
Pitcher 1988, Thompson 1988, USDA-NRCS 2004, Weber
2003
Grazing tolerant or increaser
Alters soil stability
Excretes salts or toxins
Grime 1979, Hawkes 1989, Pitcher 1988, Thompson 1988,
USDA-NRCS 2004
Thompson 1988, Weber 2003
Grime 1979, Pitcher 1988
Forms monocultures or near
monocultures
Pitcher 1988, Thompson 1988, Weber 2003
Invades in absence of human
disturbance
Large propagule crop
Wind dispersal
Animal dispersal
Plasticity
Drought tolerant
Wide light regime
Wide nutrient or soil texture
Grazing tolerant or increaser
Forms monocultures or near
monocultures
Strongly clonal
Large propagule crop
Small seeds/fruits
Dispersal over time
Weber 2003
Hordeum murinum L. ssp. glaucum (Steud.) Tzvelev
Dean 1990
Weber 2003
Dean 1990, Weber 2003
Dean 1990, Maron 2004
Dean 1990, Weber 2003
Dean 1990
Dean 1990
Dean 1990, Grime 1979
Dean 1990, Weber 2003
Hypericum perforatum L.
APRS 2000, Maron 2004, Taylor 1990, Weber 2003, Whitson
1999
APRS 2000, Weber 2003, Whitson 1999
Hickman 1993, Weber 2003
APRS 2000, Grime 1979
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Traits
Species names and references documenting the presence of
specific ecological traits associated with each species.
Water dispersal
Drought tolerant
Grazing tolerant or increaser
Increases post-fire
APRS 2000, Weber 2003
APRS 2000
APRS 2000, Taylor 1990
APRS 2000, Weber 2003, Wilson 1999
Forms monocultures or near
monocultures
Large propagule crop
Small seeds/fruits
Dispersal over time
Weber 2003
Lactuca serriola L.
Weaver 2003, Whitson 1999
Hickman 1993, Weaver 2003, Wilken 1998a
Hickman 1993, Taylor 1990, Weaver 2003, Wilken 1998a
Weaver 2003, Whitson 1999, Wilken 1998a
Weaver 2003
Weaver 2003
Lacey 2002, Weaver 2003
Lepidium campestre (L.) Ait. f.
Hickman 1993, Hitchcock 1969, Peat 2005
Peat 2005
Peat 2005
Hitchcock 1969
Peat 2005
Leucanthemum vulgare Lam.
Alvarez 2000, Clements 2004, Cranston 2002, Olson 1997,
Olson 1999, OMAF 2002, Rutledge 1998, Taylor 1990,
Whitson 1999, WSNWCB 2002
Alvarez 2000, Clements 2004, Mitch 2000, Olson 1997, Olson
1999, Rutledge 1998, WSNWCB 2002
Alvarez 2000, Clements 2004, Hickman 1993
Alvarez 2000, Mitch 2000
Wind dispersal
Olson 1999, WSNWCB 2002
Large propagule crop
Small seeds/fruits
Wind dispersal
Plasticity
Drought tolerant
Wide moisture regime
Grazing tolerant or increaser
Small seeds/fruits
Dispersal over time
Animal dispersal
Drought tolerant
Wide nutrient or soil texture
Strongly clonal
Wide moisture regime
Alvarez 2000, Andersen 1995, Clements 2004, Mitch 2000,
Olson 1999, WSNWCB 2002
Clements 2004
Clements 2004, Mitch 2000, Olson 1999
Alvarez 2000, Clements 2004, Mitch 2000, Olson 1999,
WSNWCB 2002
Wide light regime
Alkaline or saline tolerant
Alvarez 2000, Clements 2004, USDA-NRCS 2004
Clements 2004
Animal dispersal
Plasticity
Drought tolerant
Electronic Supplement - Index of Alien Impact – Appendix 1
14
Grazing tolerant or increaser
Alters nutrient cycling
Alters soil stability
Species names and references documenting the presence of
specific ecological traits associated with each species.
Clements 2004, Mitch 2000, Olson 1999, USDA-NRCS 2004,
WSNWCB 2002
Alvarez 2000, Clements 2004, Cranston 2002, Mitch 2000,
Olson 1997, Olson 1999, Taylor 1990, USDA-NRCS 2004,
WSNWCB 2002
Olson 1999
Clements 2004, Olson 1999, WSNWCB 2002
Forms monocultures or near
monocultures
Alvarez 2000, Clements 2004, CWMA 2002, Olson 1999,
OMAF 2002, Taylor 1990, WSNWCB 2002
Traits
Wide nutrient or soil texture
Invades in absence of human
disturbance
Strongly clonal
Alvarez 2000, Rutledge 1998
Linaria dalmatica (L.) P. Mill.
Butler 1994, Carpenter 1998b, Lajeunesse 1999, Prather 2002,
Rutledge 1998, Taylor 1990, Whitson 1999, WSNWCB 2002,
Zouhar 2003b
Large propagule crop
Butler 1994, Carpenter 1998b, Gates 1960, Lajeunesse 1999,
Prather 2002, WSNWCB 2002, Zouhar 2003b
Dispersal over time
Carpenter 1998b, Hickman 1993, Lajeunesse 1999, Zouhar
2003b
Butler 1994, Carpenter 1998b, Lajeunesse 1999, Rutledge 1998,
WSNWCB 2002, Zouhar 2003b
Specialized dispersal
Lajeunesse 1999
Wind dispersal
Butler 1994, Carpenter 1998b, WSNWCB 2002, Zouhar 2003b
Animal dispersal
Butler 1994, Lajeunesse 1999, Zouhar 2003b
Water dispersal
Plasticity
Drought tolerant
Flooding/saturation tolerant
Wide light regime
Wide nutrient or soil texture
Lajeunesse 1999, Zouhar 2003b
Carpenter 1998b, Lajeunesse 1999, Zouhar 2003b
Butler 1994, Carpenter 1998b, Gates 1960, Lajeunesse 1999,
Prather 2002, Rutledge 1998, Taylor 1990, Whitson 1999,
WSNWCB 2002, Zouhar 2003b
Carpenter 1998b, Rutledge 1998, Zouhar 2003b
Zouhar 2003b
Gates 1960, Lajeunesse 1999
Grazing tolerant or increaser
Beck 2002, Butler 1994, Carpenter 1998b, Lajeunesse 1999,
Rutledge 1998, Zouhar 2003b
Small seeds/fruits
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Traits
Species names and references documenting the presence of
specific ecological traits associated with each species.
Increases post-fire
Alters hydrology
Alters soil stability
Carpenter 1998b, Lajeunesse 1999, Zouhar 2003b
Carpenter 1998b, Lajeunesse 1999
Lajeunesse 1999, Zouhar 2003b
Forms monocultures or near
monocultures
Invades in absence of human
disturbance
Strongly clonal
Carpenter 1998b, Lajeunesse 1999, Rutledge 1998, Whitson
1999, WSNWCB 2002, Zouhar 2003b
Beck 2002, Lajeunesse 1999, Rutledge 1998, Taylor 1990,
Zouhar 2003b
Schedonorus phoenix (Scop.) Holub (formerly, Lolium
arundinaceum (Schreb.) S.J. Darbyshire
Batcher 2000, Gibson 2001, USDA-NRCS 2004, Walsh 1995b,
Weber 2003
Large propagule crop
Boedeltje 2003, USDA-NRCS 2004
Animal dispersal
Gibson 2001
Plasticity
Gibson 2001, Hensen 2001, USDA-NRCS 2002d, USDANRCS 2004, Walsh 1995b, Weber 2003
Drought tolerant
Wide moisture regime
Flooding/saturation tolerant
Batcher 2000, Gibson 2001, Walsh 1995b
Batcher 2000, Gibson 2001, Walsh 1995b, Weber 2003
Batcher 2000, Gibson 2001, Walsh 1995b, Weber 2003
Wide light regime
Alkaline or saline tolerant
Wide nutrient or soil texture
Grazing tolerant or increaser
Increases post-fire
Batcher 2000, USDA-NRCS 2004, Walsh 1995b
Gibson 2001, USDA-NRCS 2004, Walsh 1995b, Weber 2003
Batcher 2000, Gibson 2001, Hensen 2001, USDA-NRCS 2002d,
USDA-NRCS 2004, Walsh 1995b
Gibson 2001, Walsh 1995b
Batcher 2000, Gibson 2001, USDA-NRCS 2004, Walsh 1995b
Alters soil stability
Batcher 2000, Hensen 2001, USDA-NRCS 2002d, Walsh
1995b, Weber 2003
Excretes salts or toxins
Forms monocultures or near
monocultures
Batcher 2000, Gibson 2001, Hensen 2001, USDA-NRCS 2004,
Walsh 1995b
Batcher 2000, Gibson 2001, Hensen 2001, USDA-NRCS 2002d,
Walsh 1995b, Weber 2003
Invades in absence of human
disturbance
Batcher 2000, Walsh 1995b
Lotus corniculatus L.
Strongly clonal
Haubensak 1999b, Taylor 1990, Turkington 1980
Electronic Supplement - Index of Alien Impact – Appendix 1
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Traits
Species names and references documenting the presence of
specific ecological traits associated with each species.
Small seeds/fruits
Turkington 1980, Weber 2003
Dispersal over time
Specialized dispersal
Animal dispersal
Plasticity
Wide moisture regime
Flooding/saturation tolerant
Wide light regime
Alkaline or saline tolerant
Wide nutrient or soil texture
Turkington 1980
Turkington 1980
Turkington 1980
Turkington 1980, USDA-NRCS 2004, Weber 2003
Haubensak 1999b, Turkington 1980
Haubensak 1999b, Taylor 1983, Turkington 1980, Weber 2003
Taylor 1983, Turkington 1980, Sullivan 1992c
Turkington 1980
Turkington 1980
Grazing tolerant or increaser
Increases post-fire
Haubensak 1999b, Turkington 1980
Weber 2003
Haubensak 1999b, Turkington 1980, USDA-NRCS 2004,
Weber 2003
Alters nutrient cycling
Forms monocultures or near
monocultures
Large propagule crop
Turkington 1980, Weber 2003
Medicago lupulina L.
Turkington 1979, USDA-NRCS 2004
Small seeds/fruits
Hickman 1993, Turkington 1979
Dispersal over time
Animal dispersal
Water dispersal
Plasticity
Drought tolerant
Wide moisture regime
Wide nutrient or soil texture
Grazing tolerant or increaser
Alters nutrient cycling
Turkington 1979
Turkington 1979
Turkington 1979
Turkington 1979
Gebhart 1993, Turkington 1979, USDA-NRCS 2004
Turkington 1979, USDA-NRCS 2004
Turkington 1979
Grime 1979
Gebhart 1993, USDA-NRCS 2004
Medicago sativa L. ssp. sativa
Sullivan 1992b
Hickman 1993
Sullivan 1992b
Sullivan 1992b, USDA-NRCS 2002e, USDA-NRCS 2004
Gebhart 1993, Sullivan 1992b, Taylor 1990, USDA-NRCS 2004
Strongly clonal
Small seeds/fruits
Dispersal over time
Plasticity
Drought tolerant
Electronic Supplement - Index of Alien Impact – Appendix 1
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Traits
Species names and references documenting the presence of
specific ecological traits associated with each species.
Wide moisture regime
Alkaline or saline tolerant
Wide nutrient or soil texture
Sullivan 1992b, USDA-NRCS 2004
Sullivan 1992b
Sullivan 1992b, USDA-NRCS 2004
Grazing tolerant or increaser
Increases post-fire
Alters nutrient cycling
Alters soil stability
Sullivan 1992b
Sullivan 1992b, USDA-NRCS 2004
Gebhart 1993, Sullivan 1992b, USDA-NRCS 2004
Sullivan 1992b
Melilotus officinalis (L.) Lam.
Turkington 1978, USDA-NRCS 2004
Eckardt 1987, Hickman 1993, Turkington 1978, Weber 2003
Eckardt 1987, Turkington 1978, Weber 2003
Turkington 1978
Eckardt 1987, Sullivan 1992c, Turkington 1978
Sullivan 1992c, Turkington 1978
Eckardt 1987, Sullivan 1992c, Turkington 1978, Weber 2003
Turkington 1978
Eckardt 1987, Gebhart 1993, Sullivan 1992c, Turkington 1978,
USDA-NRCS 2004
Eckardt 1987, Sullivan 1992c, USDA-NRCS 2004
Eckardt 1987, Sullivan 1992c
Eckardt 1987, Sullivan 1992c, Turkington 1978, Weber 2003
Large propagule crop
Small seeds/fruits
Dispersal over time
Specialized dispersal
Wind dispersal
Animal dispersal
Water dispersal
Plasticity
Drought tolerant
Wide moisture regime
Flooding/saturation tolerant
Alkaline or saline tolerant
Wide nutrient or soil texture
Grazing tolerant or increaser
Increases post-fire
Alters nutrient cycling
Alters soil stability
Excretes salts or toxins
Sullivan 1992c, Turkington 1978, Weber 2003
Lacey 2002
Eckardt 1987, Sullivan 1992c, USDA-NRCS 2004, Weber 2003
Eckardt 1987, Gebhart 1993, Sullivan 1992c, Taylor 1990,
Turkington 1978, USDA-NRCS 2004, Weber 2003
Sullivan 1992c, Turkington 1978, Whitson 1999
Sullivan 1992c
Forms monocultures or near
monocultures
Eckardt 1987, Turkington 1978, Weber 2003
Invades in absence of human
disturbance
Strongly clonal
Eckardt 1987, Sullivan 1992c
Mentha ×piperita L. (pro sp.) [aquatica x spicata]
Hickman 1993, Leonard 2001
Small seeds/fruits
Hickman 1993
Electronic Supplement - Index of Alien Impact – Appendix 1
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Traits
Plasticity
Flooding/saturation tolerant
Wide light regime
Forms monocultures or near
monocultures
Large propagule crop
Small seeds/fruits
Dispersal over time
Species names and references documenting the presence of
specific ecological traits associated with each species.
Leonard 2001
Leonard 2001
Leonard 2001
Leonard 2001
Onopordum acanthium L.
Beck 1999, NRE 2002, Parent 2004, Piper 1984, WSNWCB
2002
Beck 1999, CWMA 2002, Hickman 1993, NRE 2002, Piper
1984, Whitson 1999
Wind dispersal
Beck 1999, Parent 2004, Peat 2005
Beck 1999, Hickman 1993, NRE 2002, Parent 2004, Piper 1984,
WSNWCB 2002
Animal dispersal
Beck 1999, NRE 2002, Parent 2004, Piper 1984, WSNWCB
2002
Drought tolerant
Beck 1999, NRE 2002, Parent 2004, Piper 1984, WSNWCB
2002
Beck 1999, CWMA 2002, Julian 2002, Parent 2004, Piper 1984,
Prather 2002, Taylor 1990, UNR 2002, WSNWCB 2002
Wide moisture regime
Beck 1999, Julian 2002, Parent 2004, Prather 2002, Taylor
1990, UNR 2002, WSNWCB 2002
Flooding/saturation tolerant
Beck 1999, Julian 2002, Parent 2004, Prather 2002, UNR 2002,
WSNWCB 2002
Wide nutrient or soil texture
WSNWCB 2002
Grazing tolerant or increaser
Beck 1999, CWMA 2002, Julian 2002, NRE 2002, Parent 2004,
UNR 2002, Whitson 1999, WSNWCB 2002
Forms monocultures or near
monocultures
CWMA 2002, Julian 2002, Parent 2004, Prather 2002, UNR
2002, Whitson 1999, WSNWCB 2002
Water dispersal
Invades in absence of human
disturbance
Strongly clonal
Julian 2002, Taylor 1990
Phalaris arundinacea L.*
APRS 2000, Hickman 1993, Lyons 1998, Naglich 1994, Snyder
1992, Stannard 2002, Taylor 1990, USDA-NRCS 2002f,
USDA-NRCS 2004, Weber 2003, Whitson 1999
Large propagule crop
Boedeltje 2003, USDA-NRCS 2004
Electronic Supplement - Index of Alien Impact – Appendix 1
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Traits
Species names and references documenting the presence of
specific ecological traits associated with each species.
Small seeds/fruits
Wind dispersal
Hickman 1993, Lyons 1998, Stannard 2002, USDA-NRCS 2004
Moffat 2004
Water dispersal
APRS 2000, Boedeltje 2003, Naglich 1994
Lyons 1998, Stannard 2002, USDA-NRCS 2002f, USDANRCS 2004, Weber 2003
Plasticity
Drought tolerant
Wide moisture regime
Flooding/saturation tolerant
Wide light regime
Lyons 1998, Naglich 1994
APRS 2000, Lyons 1998, Naglich 1994, Snyder 1992, USDANRCS 2004
APRS 2000, Hickman 1993, Lyons 1998, Naglich 1994, Snyder
1992, Stannard 2002, Taylor 1990, USDA-NRCS 2002f,
USDA-NRCS 2004, Weber 2003, Whitson 1999, Wilson 1999
Alkaline or saline tolerant
Lyons 1998, Naglich 1994
Lyons 1998, Snyder 1992, USDA-NRCS 2002f, USDA-NRCS
2004
Grazing tolerant or increaser
Lyons 1998, Snyder 1992, Stannard 2002, USDA-NRCS 2002f
Increases post-fire
Alters hydrology
APRS 2000, Snyder 1992, Stannard 2002, USDA-NRCS 2004,
Wilson 1999
APRS 2000, Lyons 1998
Alters soil stability
APRS 2000, Lyons 1998, Naglich 1994, Snyder 1992, USDANRCS 2002f, Weber 2003
Forms monocultures or near
monocultures
APRS 2000, Lyons 1998, Naglich 1994, Snyder 1992, Stannard
2002, Taylor 1990, USDA-NRCS 2004, Weber 2003
Invades in absence of human
disturbance
APRS 2000, Lyons 1998, Stannard 2002
Phleum pratense L.
Strongly clonal
Esser 1993a
Large propagule crop
Esser 1993a, USDA-NRCS 2004
Small seeds/fruits
Bush 2002b, Esser 1993a, Hickman 1993, USDA-NRCS 2004
Electronic Supplement - Index of Alien Impact – Appendix 1
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Traits
Species names and references documenting the presence of
specific ecological traits associated with each species.
Wind dispersal
Animal dispersal
Plasticity
Esser 1993a
Esser 1993a
Bush 2002b, Esser 1993a, USDA-NRCS 2004
Wide moisture regime
Flooding/saturation tolerant
Wide light regime
Wide nutrient or soil texture
Grazing tolerant or increaser
Increases post-fire
Alters soil stability
Excretes salts or toxins
Forms monocultures or near
monocultures
Invades in absence of human
disturbance
APRS 2000, Esser 1993a, Leege 1981
APRS 2000, Esser 1993a, Leege 1981
APRS 2000, Esser 1993a
Bush 2002b, Esser 1993a, USDA-NRCS 2004
Crowe 1997, Esser 1993a, Green 1995, Leege 1981
Esser 1993a, USDA-NRCS 2004
Esser 1993a, Weber 2003
Esser 1993a
Weber 2003
Strongly clonal
Esser 1993a, Weber 2003
Plantago lanceolata L.
Cavers 1980, Clapham 1987, Weber 2003
Large propagule crop
Cavers 1980
Small seeds/fruits
Dispersal over time
Animal dispersal
Plasticity
Cavers 1980, Clapham 1987, Hickman 1993, Weber 2003,
Whitson 1999
Cavers 1980, Grime 1979, Tonsor 1993
Cavers 1980, Clapham 1987, Weber 2003
Cavers 1980, Clapham 1987, van der Toorn 1988, Weber 2003
Drought tolerant
Wide moisture regime
Flooding/saturation tolerant
Cavers 1980
Cavers 1980, Clapham 1987
Weber 2003, Hitchcock 1969
Wide light regime
Alkaline or saline tolerant
Wide nutrient or soil texture
Forms monocultures or near
monocultures
Invades in absence of human
disturbance
Cavers 1980
Cavers 1980, Clapham 1987
Cavers 1980, Clapham 1987
Large propagule crop
Weber 2003
van der Toorn 1988
Plantago major L.
APRS 2000, Hawthorn 1974
Electronic Supplement - Index of Alien Impact – Appendix 1
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Traits
Species names and references documenting the presence of
specific ecological traits associated with each species.
Small seeds/fruits
Hawthorn 1974, Hickman 1993, USDA-NRCS 2004, van der
Toorn 1988
Dispersal over time
Animal dispersal
Plasticity
Hawthorn 1974
APRS 2000, Hawthorn 1974
Hawthorn 1974, Hickman 1993
Drought tolerant
Wide moisture regime
Hawthorn 1974, USDA-NRCS 2004
Hawthorn 1974, USDA-NRCS 2004
APRS 2000, Hawthorn 1974, USDA-NRCS 2004, van der
Toorn 1988
APRS 2000, Hawthorn 1974, USDA-NRCS 2004
Hawthorn 1974, USDA-NRCS 2004
USDA-NRCS 2004
Wide light regime
Wide nutrient or soil texture
Grazing tolerant or increaser
Increases post-fire
Strongly clonal
Large propagule crop
Specialized dispersal
Drought tolerant
Wide moisture regime
Wide light regime
Wide nutrient or soil texture
Grazing tolerant or increaser
Increases post-fire
Alters soil stability
Invades in absence of human
disturbance
Poa bulbosa L.
Harrison (no date)
Harrison (no date)
Harrison (no date), Hawkes 1989, Hickman 1993, Taylor 1990,
Whitson 1999
Harrison (no date), USDA-NRCS 2004
APRS 2000, USDA-NRCS 2004
APRS 2000, USDA-NRCS 2004, Hickman 1993
Harrison (no date), USDA-NRCS 2004
Harrison (no date)
USDA-NRCS 2004
APRS 2000, Harrison (no date)
Large propagule crop
Harrison (no date)
Poa pratensis L. and Poa compressa L.
APRS 2000, Bush 2002c, Crowe 1997, Green 1995, Hickman
1993, Sather 1996, Taylor 1990, Uchytil 1993, USDA-NRCS
2004, Weber 2003, Zouhar 2003c
APRS 2000, Bush 2002c, Sather 1996, Uchytil 1993, USDANRCS 2004
Small seeds/fruits
Dispersal over time
APRS 2000, Bush 2002c, Hickman 1993, USDA-NRCS 2004
Sather 1996
Strongly clonal
Electronic Supplement - Index of Alien Impact – Appendix 1
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Traits
Species names and references documenting the presence of
specific ecological traits associated with each species.
Plasticity
Bush 2002c, Green 1995, Taylor 1990, USDA-NRCS 2004
Drought tolerant
Bush 2002c, Uchytil 1993
Wide moisture regime
Flooding/saturation tolerant
APRS 2000, Bush 2002c, Sather 1996, Uchytil 1993
APRS 2000, Sather 1996, Uchytil 1993, Weber 2003
APRS 2000, Bush 2002c, Sather 1996, Uchytil 1993, USDANRCS 2004
Hickman 1993
USDA-NRCS 2004
APRS 2000, Clary 1995, Crowe 1997, Green 1995, Hellström
2003, Sather 1996, Uchytil 1993
Wide light regime
Alkaline or saline tolerant
Wide nutrient or soil texture
Grazing tolerant or increaser
Increases post-fire
Alters soil stability
Forms monocultures or near
monocultures
Invades in absence of human
disturbance
Uchytil 1993, USDA-NRCS 2004
APRS 2000, Bush 2002c, Uchytil 1993, Weber 2003
Bush 2002c, Sather 1996, USDA-NRCS 2004, Weber 2003
Large propagule crop
Sather 1996, Uchytil 1993
Polygonum convolvulus L.
Hume 1983
Small seeds/fruits
Hickman 1993, Hitchcock 1969, Hume 1983, Whitson 1999
Dispersal over time
Water dispersal
Wide moisture regime
Wide nutrient or soil texture
Small seeds/fruits
Dispersal over time
Wind dispersal
Animal dispersal
Hume 1983
Andersen 1995, Hume 1983
Hume 1983
Hume 1983
Potentilla recta L.
Endress 2005, Rice 1991, Werner 1976, Zouhar 2003c
Endress 2005, Hickman 1993, Rice 1999, Werner 1976, Zouhar
2003c
Endress 2005, Grime 1979, Zouhar 2003c
Endress 2005
Endress 2005
Water dispersal
Plasticity
Endress 2005
Werner 1976
Large propagule crop
Electronic Supplement - Index of Alien Impact – Appendix 1
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Traits
Drought tolerant
Wide moisture regime
Wide light regime
Wide nutrient or soil texture
Grazing tolerant or increaser
Increases post-fire
Alters soil stability
Forms monocultures or near
monocultures
Invades in absence of human
disturbance
Species names and references documenting the presence of
specific ecological traits associated with each species.
Endress 2005, Rice 1991, Werner 1976, Zouhar 2003c
Endress 2005, Rice 1991, Rice 1999, Zouhar 2003c
Endress 2005, Zouhar 2003c
Rice 1999, Werner 1976
Endress 2005, Rice 1991, Rice 1999, Werner 1976, WSNWCB
2002, Zouhar 2003c
Endress 2005, Zouhar 2003c
Werner 1976, WSNWCB 2002
Endress 2005, Rice 1999, Werner 1976, WSNWCB 2002
Strongly clonal
Endress 2005, Rice 1999, Whitson 1999, Zouhar 2003c
Rumex acetosella L.
Esser 1995, Hickman 1993, Taylor 1990, Weber 2003, Whitson
1999, Wilken 1998b
Small seeds/fruits
Weber 2003, Whitson 1999, Wilken 1998b
Wind dispersal
Animal dispersal
Esser 1995, Weber 2003
Esser 1995, Weber 2003
Plasticity
Wide moisture regime
Flooding/saturation tolerant
Wide light regime
Wide nutrient or soil texture
Grazing tolerant or increaser
Increases post-fire
Forms monocultures or near
monocultures
Invades in absence of human
disturbance
Weber 2003, Wilken 1998b
Esser 1995, Weber 2003
Esser 1995, Weber 2003
Esser 1995
Esser 1995, Whitson 1999, Wilken 1998b
Esser 1995, Leege 1981
Esser 1995, Wilken 1998b
Esser 1995, Weber 2003
Large propagule crop
Esser 1995, Weber 2003
Rumex crispus L.
Boedeltje 2003, Weber 2003
Small seeds/fruits
Hickman 1993, Weber 2003, Wilken 1998c
Dispersal over time
Wind dispersal
Water dispersal
Grime 1979, Weber 2003, Wilken 1998c
Moffat 2004, Weber 2003, Whitson 1999
Andersen 1995, Boedeltje 2003, Weber 2003, Whitson 1999
Electronic Supplement - Index of Alien Impact – Appendix 1
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Traits
Plasticity
Flooding/saturation tolerant
Wide light regime
Alkaline or saline tolerant
Grazing tolerant or increaser
Alters soil stability
Forms monocultures or near
monocultures
Large propagule crop
Small seeds/fruits
Dispersal over time
Specialized dispersal
Wind dispersal
Drought tolerant
Wide moisture regime
Alkaline or saline tolerant
Wide nutrient or soil texture
Grazing tolerant or increaser
Increases post-fire
Invades in absence of human
disturbance
Species names and references documenting the presence of
specific ecological traits associated with each species.
Weber 2003, Wilken 1998c
Weber 2003, Wilken 1998c
Weber 2003
Wilken 1998c
Crowe 1997, Wilken 1998c
Wilken 1998c
Weber 2003
Sisymbrium altissimum L.
APRS 2000, Howard 2003, Peat 2005, Whitson 1999
Hickman 1993, Hitchcock 1969, Peat 2005, Whitson 1999
Peat 2005
APRS 2000, Howard 2003, Taylor 1990, Whitson 1999
Howard 2003, Peat 2005, Taylor 1990, Whitson 1999
Howard 2003
Howard 2003
Howard 2003
Howard 2003
Howard 2003, Zollinger 1999
Howard 2003
Small seeds/fruits
APRS 2000, Howard 2003
Sonchus asper (L.) Hill
Boedeltje 2003, Clapham 1987, Holm 1997, Hutchinson 1984
Clapham 1987, Hickman 1993, Holm 1997, Hutchinson 1984,
Wilken 1998d
Wind dispersal
Animal dispersal
Clapham 1987, Hickman 1993, Holm 1997, Hutchinson 1984,
Taylor 1990, Wilken 1998d, Zollinger 1999
Holm 1997, Hutchinson 1984, Zollinger 1999
Water dispersal
Boedeltje 2003, Holm 1997
Plasticity
Holm 1997, Hutchinson 1984
Alkaline or saline tolerant
Wide nutrient or soil texture
Holm 1997, Hutchinson 1984
Clapham 1987, Hutchinson 1984
Taraxacum officinale G.H. Weber ex Wiggers
Boedeltje 2003, Esser 1993b, Stewart-Wade 2002, USDANRCS 2004
Large propagule crop
Large propagule crop
Electronic Supplement - Index of Alien Impact – Appendix 1
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Wind dispersal
Species names and references documenting the presence of
specific ecological traits associated with each species.
Haubensak 1999c, Hickman 1993, Stewart-Wade 2002, USDANRCS 2004, Whitson 1999
Esser 1993b, Stewart-Wade 2002
Esser 1993b, Haubensak 1999c, Stewart-Wade 2002, Taylor
1990, Whitson 1999
Animal dispersal
Stewart-Wade 2002
Water dispersal
Boedeltje 2003, Stewart-Wade 2002
Haubensak 1999c, Stewart-Wade 2002, Taylor 1990, Whitson
1999
Traits
Small seeds/fruits
Dispersal over time
Plasticity
Drought tolerant
Wide moisture regime
Flooding/saturation tolerant
Stewart-Wade 2002, USDA-NRCS 2004
Esser 1993b, Leege 1981, Stewart-Wade 2002, USDA-NRCS
2004
Esser 1993b, Leege 1981
Wide light regime
Wide nutrient or soil texture
Stewart-Wade 2002, USDA-NRCS 2004
Esser 1993b, Stewart-Wade 2002, USDA-NRCS 2004
Grazing tolerant or increaser
Increases post-fire
Excretes salts or toxins
Invades in absence of human
disturbance
Crowe 1997, Esser 1993b, Green 1995, Leege 1981, StewartWade 2002, USDA-NRCS 2004
Esser 1993b, Stewart-Wade 2002
Stewart-Wade 2002
Esser 1993b, Haubensak 1999c, Stewart-Wade 2002
Strongly clonal
Large propagule crop
Thinopyrum intermedium (Host) Barkworth & D.R. Dewey
Hickman 1993, Hull 1971, Ogle 2003, USDA-NRCS 2004
USDA-NRCS 2004
Drought tolerant
Wide moisture regime
Alkaline or saline tolerant
Wide nutrient or soil texture
Harrison (no date), Hull 1971, Ogle 2003, USDA-NRCS 2004
Ogle 2003
Ogle 2003, USDA-NRCS 2004
USDA-NRCS 2004
Increases post-fire
Alters soil stability
Forms monocultures or near
monocultures
Invades in absence of human
disturbance
Ogle 2003, USDA-NRCS 2004
Harrison (no date), Ogle 2003
Harrison (no date), Ogle 2003
Harrison (no date), Ogle 2003
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Traits
Species names and references documenting the presence of
specific ecological traits associated with each species.
Large propagule crop
Tragopogon dubius Scop.
USDA-NRCS 2004
Wind dispersal
Hickman 1993, Hitchcock 1969, Taylor 1990
Drought tolerant
Wide nutrient or soil texture
Grazing tolerant or increaser
APRS 2000, Hitchcock 1969, Taylor 1990, USDA-NRCS 2004
USDA-NRCS 2004
Lacey 2002, USDA-NRCS 2004
Trifolium dubium Sibthorp
Strongly clonal
Large propagule crop
Small seeds/fruits
Dispersal over time
Peat 2005
Peat 2005
FAO 2005, Grime 1979
Wilson 1999
FAO 2005
Trifolium hybridum L.
USDA-NRCS 2004
USDA-NRCS 2004
USDA-NRCS 2004
APRS 2000, USDA-NRCS 2004
USDA-NRCS 2004
Trifolium pratense L.
USDA-NRCS 2004
APRS 2000
USDA-NRCS 2004
Hellström 2003
USDA-NRCS 2004
Trifolium repens L.
Coladonato 1993, Crowe 1997, Grime 1979, Taylor 1990,
Turkington 1983, USDA-NRCS 2004, Weber 2003
Turkington 1983
Turkington 1983, USDA-NRCS 2004, Weber 2003
Coladonato 1993, Turkington 1983
Wind dispersal
Animal dispersal
Water dispersal
Plasticity
Wide moisture regime
Coladonato 1993
Coladonato 1993, Moffat 2004, Turkington 1983, Weber 2003
Coladonato 1993
Turkington 1983, USDA-NRCS 2004, Weber 2003
Coladonato 1993
Dispersal over time
Grazing tolerant or increaser
Increases post-fire
Alters nutrient cycling
Large propagule crop
Small seeds/fruits
Wide moisture regime
Flooding/saturation tolerant
Alters nutrient cycling
Large propagule crop
Dispersal over time
Plasticity
Grazing tolerant or increaser
Alters nutrient cycling
Electronic Supplement - Index of Alien Impact – Appendix 1
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Traits
Wide light regime
Wide nutrient or soil texture
Grazing tolerant or increaser
Alters nutrient cycling
Forms monocultures or near
monocultures
Species names and references documenting the presence of
specific ecological traits associated with each species.
Coladonato 1993, USDA-NRCS 2004
Coladonato 1993, Turkington 1983, USDA-NRCS 2004
Crowe 1997, Grime 1979, Turkington 1983, Weber 2003
Coladonato 1993, Turkington 1983, USDA-NRCS 2004, Weber
2003
Animal dispersal
Turkington 1983, Weber 2003
Ventenata dubia (Leers) Coss. & Durieu
Hitchcock 1969
Drought tolerant
Martin 2003, Old 1987, Whitson 1999
Grazing tolerant or increaser
Martin 2003, Old 1987, Whitson 1999
Verbascum blattaria L.
Gross 1978, Whitson 1999
Gross 1978
Gross 1978
Gross 1978
Gross 1978
Gross 1978
Large propagule crop
Small seeds/fruits
Dispersal over time
Specialized dispersal
Plasticity
Wide moisture regime
Forms monocultures or near
monocultures
Large propagule crop
Small seeds/fruits
Dispersal over time
Specialized dispersal
Plasticity
Drought tolerant
Wide moisture regime
Grazing tolerant or increaser
Increases post-fire
Forms monocultures or near
monocultures
Invades in absence of human
disturbance
Strongly clonal
Gross 1978
Verbascum thapsus L.
Gross 1978, Hoshovsky 1986, Pitcairn 2000, Whitson 1999
Gross 1978, Hoshovsky 1986, Pitcairn 2000, Whitson 1999
Gross 1978, Pitcairn 2000, Weber 2003
Gross 1978
Gross 1978
Gross 1978, Hoshovsky 1986, Pitcairn 2000, Weber 2003
Hoshovsky 1986, Pitcairn 2000, Weber 2003
Gross 1978, Hoshovsky 1986, Pitcairn 2000, Whitson 1999
Hoshovsky 1986, Pitcairn 2000
Gross 1978, Weber 2003
Pitcairn 2000, Weber 2003
Veronica anagallis-aquatica L.
Hickman 1993, Peat 2005
Electronic Supplement - Index of Alien Impact – Appendix 1
28
Traits
Species names and references documenting the presence of
specific ecological traits associated with each species.
Large propagule crop
Hitchcock 1969
Small seeds/fruits
Wind dispersal
Animal dispersal
Plasticity
Flooding/saturation tolerant
Hickman 1993, Hitchcock 1969
Peat 2005
Peat 2005
Boeger 2003
Boeger 2003, Hickman 1993, Hitchcock 1969, Peat 2005
Vulpia bromoides (L.) S.F. Gray
Dowling 2004, Haubensak 1999d
Haubensak 1999d
Haubensak 1999d, Hickman 1993, Hitchcock 1969
Hitchcock 1969
Hitchcock 1969, Weber 2003
Hitchcock 1969
Haubensak 1999d
Haubensak 1999d
Large propagule crop
Dispersal over time
Drought tolerant
Wide moisture regime
Flooding/saturation tolerant
Alkaline or saline tolerant
Increases post-fire
Excretes salts or toxins
Forms monocultures or near
monocultures
Invades in absence of human
disturbance
Weber 2003
Weber 2003
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