CHAPTER 11 - Gas Exchange Review questions ANSWERS 1 It is necessary for organisms to exchange oxygen and carbon dioxide with their external environment to provide the necessary requirements for cellular respiration and photosynthesis. During cellular respiration oxygen is required for the conversion of glucose into carbon dioxide and water. Energy is released as a result of this conversion. The carbon dioxide produced by this process will cause the tissue to become more acidic if it is not removed. Increased acidity would damage the structure of many of the important molecules thus affecting their function. In photosynthesis, carbon dioxide is required from the external environment. Carbon dioxide and water are converted into glucose and oxygen. The oxygen is released into the external environment. 2 Diffusion is the process by which gases move across an exchange surface. The direction and rate of diffusion are dependent upon the concentration gradient of the substance being transferred across the exchange surface. Molecules will move to a region of lower concentration and they will move more quickly if there is large concentration gradient. 3 Algae do not have any stomata for gaseous exchange and carbon dioxide diffuses across their exterior surface into the photosynthetic cells. 4 Cellular respiration Photosynthesis Takes place in all organisms Only takes place in plants Takes place all the time Only happens in sunlight or under artificial lights Produces energy Requires energy 5 During the day two processes occur i.e. photosynthesis and respiration. Because the rate of photosynthesis is usually faster than the rate of respiration, carbon dioxide produced in respiration is not sufficient to supply enough carbon dioxide for photosynthesis and it diffuses into the leaf. At night only one process i.e. respiration and this requires oxygen. 6 Take a potted plant and paint a section of the underside of a leaf with Vaseline or a substance impermeable to gases. Leave the plant in bright sunlight for a couple of hours. Remove the leaf and test it for starch. The area that was painted should be negative. 7a Leaves have stomata whose opening and closing is controlled by guard cells. Stems have lenticels which are a group of loosely packed cells through which air passes freely to the cells beneath. Just behind the growing points of roots are root hairs. Oxygen readily diffuses into the film of moisture surrounding the root hairs, and then into the roots themselves. 7b air stomata air spaces dissolves in moisture surrounding the cell membrane passes through the membrane 8a They have chloroplasts which undergo photosynthesis and produce glucose which is used in respiration to make energy for movement i.e. opening and closing. The glucose produced by photosynthesis causes water to move into the guard cells by osmosis. The cells become turgid and the stomata open. They also have a thickened inner cell wall with cellulose fibres that prevent the cells expanding in width. Therefore they expand lengthways and this buckles the guard cells and opens the stoma. 8b If the water supply is abundant the guard cells will be full of water (turgid) and the stomata will be open. If water is lost the guard cells lose their turgor and the stomata close. In bright sunlight photosynthesis will take place in the guard cells. As the concentration of glucose in the cells increases water will move in, increasing the turgor and the stomata will open. If the concentration of carbon dioxide is low inside the leaf the stomata will open so that photosynthesis can continue. 8c Land plants need to conserve water. The rays of the sun are very drying and they strike the tops of the leaves. By not having any stomata in the upper surface water loss is minimised. Aquatic plants need to float on top of the water without coming water-logged and having stomata only on the top of the leaf allows free exchange of gases between the leaf and the atmosphere. 8d The soil has poor drainage and becomes saturated with water. This reduces the amount of oxygen available to the root cells. The roots cannot take up oxygen which is needed for respiration and the plant could die. 9 There could be special structures in their stems above the level of the water which allow exchange of gases. They also could have large intercellular spaces in their stems to allow rapid diffusion. 10 Comparison of lungs and fish gills: Feature Large surface area Thin and moist Good ventilation Good blood supply Protection/support Features Large surface area Thin and moist Good ventilation Good blood supply Protection/support Lungs Greater gas exchange per unit of time To allow gases to pass through easily. Gas can only pass across an exchange surface if it is in solution, thus the surface must be moist. It is difficult for gases to pass through thick barriers To provide an adequate supply of the gas being transferred or the gas will become scarce and the rate of exchange will drop Inadequate blood flow means oxygen accumulates and so diffusion slows down Respiratory membranes are delicate and as such require protection from physical damage and, in terrestrial animals, protection from water loss Fish gills Fish gills are composed of a series of gill arches each of which has many finger-like gill filaments. Each of these has a row of closely packed, flat, leaf-like lamellae across which gas exchange takes place. The lamellae provide a large surface area for gas exchange Lamellae are thin and moist Gills are ventilated with oxygen by the movement of water over the gills Lamellae contain many blood vessels The gills are covered by the operculum, which protects the fragile gills 11a Gills are not suitable for breathing air because they are external respiratory surfaces. Being external, gills would be prone to water loss due to evaporation of water from the respiratory surface. Also, when gills are out of water the filaments clump together and the surface area is reduced. 11b Lungs rely on the ventilating medium being able to exit the body by the same route it entered. Water is a heavy medium compared with air and a great deal of energy would be required to move water across the respiratory surface in two directions. Also, water contains less oxygen than air, and it would not be possible to maintain an effective exchange gradient for oxygen in the lungs. 12 They have highly efficient mechanisms of moving water over their respiratory surfaces i.e. they move large volumes of water with little dissolved oxygen. Most fish actively ventilate their gills by means of a double-action pump of the mouth and opercular cavities, crayfish use the rhythmic beating of specialised appendages and some fish such as sharks and tuna use ram ventilation. The respiratory surfaces have a large surface area exposed to the respiratory medium and a countercurrent flow system between the blood and respiratory medium ensures maximum oxygen uptake. 13a Water-ventilating animals use dissolved oxygen levels and air-ventilating animals use dissolved carbon dioxide levels. 13b For water ventilators carbon dioxide removal is easy but oxygen is the crucial factor because it is in short supply. For air ventilators, oxygen is abundant in air but carbon dioxide is only slowly removed by diffusion and tends to accumulate in body fluids lowering the pH to dangerous levels. 14 Frog Insect Advantages and disadvantag es Air is drawn into the mouth through nostrils and then is pushed into the lungs by raising and lowering the floor of the mouth One pair of lungs Oxygen moves into the circulatory system by which it is distributed to the body cells The skin is also used as a respiratory surface. It will only function if it is kept moist Oxygen reaches the cells as the circulatory system carries it there Exchange surface can more easily dry out, especially the skin Rate of breathing can be adjusted according to the needs of the organism for oxygen Air moves by diffusion through the spiracles and trachea Spiracles and trachea in every segment. Spiracles can close to reduce moisture loss Oxygen is exchanged directly with the cells where it will be used Oxygen reaches the cells rapidly Exchange surface is deep within the body therefore there is little chance of it drying out Rate of gas exchange depends on the diffusion gradient. To some extent this will be modified by the level of activity of the organism. The cells that are using oxygen at the fastest rate will gain oxygen at the fastest rate 15 The functions of various parts of the respiratory system are as follows: • Nasal passages filter, moisten and warm air. • The trachea, bronchi and bronchioles make up the passage along which air travels. • The trachea and bronchi are lined with ciliated cells and secrete mucus. Particles of dust or bacteria are trapped by the mucus and swept by the cilia back up to the pharynx and swallowed. • Alveoli are the site of gas exchange. 16a During inhalation: • the diaphragm contracts downwards • the ribs are raised outwards by contraction of the intercostal muscles • the chest cavity increases in volume. During exhalation: • the diaphragm relaxes and moves up • the intercostal muscles relax and the ribs lower • the chest cavity decreases in volume. 16b Raising the ribs and lowering the diaphragm during inhalation increases the volume of the chest cavity and draws air into the lungs. Relaxing these muscles during exhalation reduces the volume of the chest cavity and forces air out of the lungs. 17a Respiratory pigments make it possible to carry much more oxygen than could be carried by the fluid (plasma) alone. They increase the oxygen-carrying capacity of the blood. 17b Respiratory pigments provide energy savings for an animal by increasing the oxygen-carrying capacity of the blood. An increased oxygen-carrying capacity results in a reduction in the amount of energy that must be spent pumping the blood. It also means that the animal can have a much smaller volume of blood, and that it can be pumped more slowly. 18 Breathing carbon monoxide can be fatal because it binds irreversibly to the same position on haemoglobin as oxygen, thus preventing haemoglobin molecules from carrying oxygen. Death can occur if the oxygen-carrying capacity of the blood is reduced below tolerable limits. 19a About 7% of the carbon dioxide is transported in the plasma. About 23% of carbon dioxide transported in blood combines with haemoglobin molecules forming carbaminohaemoglobin. The carbon dioxide binds to a different site on the haemoglobin molecule to the site where oxygen binds. About 70% of carbon dioxide transported in the blood exists as hydrogen carbonate ions and is transported in plasma. 19b The amount of carbon dioxide dissolved in blood plasma must be kept low because when it combines with water in the plasma carbon dioxide forms carbonic acid which results in a decrease in the pH of the plasma. Low pH can affect the structure of proteins (e.g. enzymes) thus affecting their function. 20a Asthma: The space through which air can travel in the airways (bronchioles) is reduced. Emphysema:The alveoli break down reducing the surface area across which gases can be exchanged. Pneumonia: The alveoli become filled with fluid. Anaemia: There is less haemoglobin in the blood. 20b Asthma: The amount of air that can be exchanged between the lungs and the atmosphere is limited. Emphysema: Gas exchange may be reduced to one quarter of normal. Pneumonia: The surface area of the alveoli in contact with the air is reduced and gas exchange becomes less efficient. Anaemia: The smaller amount of haemoglobin reduces the ability of the blood to carry oxygen to the cells.