EXCRETORY SYSTEM Regulate the chemical composition of body fluids by removing metabolic wastes and retaining the proper amounts of water, salts, and nutrients. Components of this system in vertebrates liver, lungs, skin. the kidneys-urinary system Liver is involved in detoxification of toxic materials that are consumed by animals while lungs remove carbon dioxide and the skin removes excess water and salt through sweating. THE URINARY SYSTEM Consists of: Two kidneys, Two ureters A urinary Bladder The Urethra THE KIDNEYS Vertebrates have paired kidneys Located in the dorsal part of abdominal cavity ventral to lumber vertebrae. In most animals right kidney slightly cranial to left. In ruminants left kidney is loosely attached to the body wall as it is pushed to right side when rumen gets filled. Kidneys regulate body fluid levels primarily and remove wastes as a secondary one. FUNCTIONS: Maintain volume of extracellular fluid Maintain ionic balance in extracellular fluid Regulate pH and osmotic concentration of extracellular fluid. Excrete metabolic waste as urea, ammonia, uric acid URINARY SYSTEM Kidneys Blood vessel Ureter Urinary bladder Urethra ANATOMY OF KIDNEYS Medial border of the kidney is concave and has depressionHilus -arteries and nerves enter - the ureters, lymphatics,veins leave kidneys. Renal pelvis, -Origin of ureter inside kidney, which is expanded receives urine from collecting tubules. Medulla- Part of kidney surrounding the renal pelvis. Cortex-the outer part surrounding the medulla. Glomeruli, proximal and distal convoluted tubules are located in medulla. The outer most layer covering the kidney is made up of thin connective tissue called renal capsule. Kidney identification Bovine kidneys: Bean shape Lobulated-distinct lobules, visible externally. Multiple pappilae. Equine kidneys smooth without lobulations. right bean shaped left - heart shaped Pigs Long and narrow bean shaped smooth externally but lobulated internally. Multiple papillae when cut. Sheep, goat, dog, cat- bean shaped, smooth without lobulations. The Nephron Is the functional unit of urinary system Consists of 1. Cup-shaped capsule -Bowman’s capsule 2. Glomerulus- capillary tuft. 3. Together they are called-renal corpuscle. 4. Renal tubule. The tubule is divided into proximal and distal convoluted tubule, descending and ascending limbs that form loop of Henle. The tubule finally joins collecting duct to empty urine into renal pelvis. URINE PRODUCTION Three steps in the production of urine in the kidneys 1) glomerular filtration, 2) Tubular reabsorption and 3) tubular secretion. 1.Glomerular filtration The first step Occurs at the glomerulus-a ball of capillaries that makes close contact with the end of the nephron. Driven by the beating of the heart, blood plasma (the fluid component of blood) is forced out of the capillaries and into the Bowman’s capsule., the Larger molecules mainly proteins and blood cells cannot pass. Glucose, urea, creatinine, Na, K,Cl passses through. Clinical importance- Proteinuria, haematuria 2. Tubular reabsorption Large part of urine production involves removing filtrate for delivery back to the blood-reabsorption, is highly selective. Cells in the proximal and distal convoluted tubules actively absorb nutrients out of the filtrate- help of microvilli seen in the lumen of tubules. Reabsorption of water as well- osmosis. Sodium ion, glucose, amono acids and other nutrients are also reabsorbed into the blood from the filtrate. (Diabetes-glucosuria) 3. Tubular secretion At the same time, other cells in the proximal and distal tubules are engaged in secretion. This process involves active transport of specific molecules into the filtrate. An example of a commonly secreted substance is hydrogen ions, potassium ions, ammonium ions, creatinine and urea. Finally, the tubule empties its contents into one of the many collecting ducts. The urine now contains a concentrated solution of whatever the body currently considers being "wastes." Water continues to be reabsorbed in the collecting duct, so that little water is wasted in the process of excretion. Finally, all the urine collects together in the kidney and is passed out through the ureter, and from there to the bladder. When the bladder is filled or enough urine is produced it is expelled. MICTURITION Process of expulsion of urine from bladder. When the bladder is distended pressure builds up and it stimulates reflex center in the spinal cord, which relaxes the smooth muscle sphincter at the neck of the bladder and contraction of bladder expelling urine out. But…reflex emptying of bladder can be controlled by urethral muscle. Ureters Each ureter originates at the renal pelvis. It is a muscular tube that conveys urine from pelvis of kidney to the urinary bladder. The smooth muscle undergoes peristaltic waves of contraction that brings the urine into the bladder. Urinary bladder Is hollow muscular organ that vary in size and position with the quantity of urine it contain. When empty it is thick walled pear shaped and located on the floor of pelvis. when filled with urine wall become thin and displaced towards the abdominal cavity. Neck of bladder continuous with urethra caudally. Muscles at the neck of bladder is arranged in circular manner forming sphincter that regulate passage of urine into urethra. Urethra Is the continuation of neck of bladder. It is a tubular structure that drains out urine during micturition. In male it divided into pelvic and penile urethra while in female pelvic urethra opens into vagina. In male pelvic urethra is joined by vas deferens and ducts from accessory sex glands as well. CLINICAL TERMS Nephroectomy- Surgical removal of kidneys Nephritis-inflammation of kidneys. Urolithiasis- Presence of stones in the urinary system. Nephrolith- Stone in the kidneys. Diuretic- Any substance that increase urine output. Oligouria- Scanty amount of urine formation. Polyuria- Excess amount of urine formation. Dysuria- painful urination. Enuresis- involuntary urination/bed wetting. Pyuria- pus in urine Uninalysis-Microspic examination /analysis of urine. Cystitis-Inflammation of urinary bladder. Micturation –urination /voiding of urine. A reflex action stimulated by distension of bladder. Can be prevented by voluntary control of urethralis muscles sorrounding the pelvic urethra. CLINICAL TERMS Kidney worms-Dictophyma renale worm in kidneys of dogs and cats -may damage the kidneys. Kidney stones formation results from crystallization of chemical salts like ammonium urate or sodium urate etc. it may obstruct urethra especially in males. Nephritis may result either due to infection or presence of irritants Ingestion of some plants may cause urinary tract tumors Glucosuria (glucose in urine)- Uncontrolled diabetesexcess level of blood glucose-cannot be reabsorbed in the proximal tubules. Proteinuria (proteins in urine)- diseases that affects the glomeruli. Renal insufficiency- degree or relative loss of renal function, but the animal can survive its state. Renal Renal Failure-complete loss of renal tissue function and the animal can't survive or can't continue its existence.