FOTOMORFOGÉNESIS • Desarrollo foto y escotomorfogénico • Fotorreceptores – Fitocromos – Criptocromos – Fototropinas Transducción de señales por los fitocromos Ritmo circadiano COP9-signalosoma Plant development normally shows plasticity within certain bounds, that is, without falling into extreme phenotypes. The occurrence of multiple receptors and downstream signalling pathways, negative regulators, and positive and negative interactions provides the homeostasis that prevents exaggerated phenotypes under extreme environments. This system leaves room for change because the actual phenotype obtained within the range of normal developmental plasticity can depend on just a few receptors, negative regulators that establish the need for further environmental cues, and interactions that modulate the relationship between signal input and output. Convergencia de la información de los fotones De-etiolación de una plántula Plantulas creciendo en distintas condiciones de luz y análisis de la expresión génica correspondiente mediante microarrays. CONCEPTO DE FOTOMORFOGENESIS Y ESCOTOMORFOGENESIS fotorreceptores de plantas Dominios en el fitocromo Luz blanca Luz Roja Luz Infrarroja Luz Azul Mutantes de fitocromo en Arabidopsis the network that connects developmental genes to environmental cues. Phytochrome photoperception and signal transduction in seedling photomorphogenesis. Light-induced nuclear translocation of phyA. PIF3 PKS1 PKS1 interacts with phytochrome PKS1 expression Light-regulated PKS1 phosphorylation by phyA in vitro. Seedlings overexpressing PKS1 have reduced sensitivity to white light. Direct targeting of light signals to primary photoresponse genes. Photons convey information. Cryptochrome and flowering time Phot1 Phot1-2 EL CONTROL DEL DESARROLLO FOTOMORFOGÉNICO -Sistema de control por degradación --EL COP9 signalosoma -COP1 -Componentes del sistema de identificación y degradación -Papel del criptocromo cry1 -Control del hy5 La red que conecta los genes del desarrollo con las fluctuaciones ambientales Hy5 The light regulation of HY5 is dependent on the nuclear abundance of COP1 and its ability to interact with COP1.