MADE BY:- Himanshu Bhatt * Contain Morphology The Root Modification Of Root Functions Of Root Rootless Plant Plants that are all Roots * * * Plant morphology (morphe-form+logos-study) is a branch of Botany which deals with the study of forms and features of different plant organs like roots, stems, leaves, flowers, seeds, fruits etc., including there development * * The root is that green cylindrical part of a plant which is given rise by radicle , the first to come out as a descending organ during germination of a seed * * Root Cap * Meristematic zone * Zone of Elongation * Root Hair * Zone of Mature Cells * * Root cap is a thimble- shaped or cap like protective structure present at the tip of root apex covering the root meristem. * It is a parenchymatous and multicellular . * The root cap secrets mucilage that lubricates the passage of root through the soil. * The soft and tender root would be unable to penetrate the hard soil if the root cap is absent . * * It is a small region(about 1mm in length) positioned subterminally behind the root cap. * Meristematics zone or zone of cell division consists of isodimetric meristematics cells arranged closely or compactly and therefor, represents the growing point or zone of the root. * The Meristematic zone continuously produces new cells that are added outwards to the root cap and inward to the basal region of the root resulting in the longitudinal growth of the root * * Lying just behind the meristematic zone, the zone of elongation is about 4-8mm in length and is made up of newly produced cells by growing point. * The cell of this zone lose the power of division but elongate rapidly thus increasing the length of the root. * The cell of this zone possess the power of absorption of water and mineral salts from the soil. * * Root hair zone is also called zone maturation or differentiation as the cells of this zone undergo maturation and differentiation into different type of primary tissues of the root, e.g., epiblema, cortex, endodermis, pericycle, phloem, xylem, etc.,. * * In aquatic plants root hair are usually absent. * Mycorrhizal roots lack root hair. * The number of root hairs are over 35,000 per square cm of root surface and many billions per plant. * * Laying behind the root hair zone, the zone of mature cells constitutes the bulk of the root. * The outermost layer of it consists of thick walled or impermeable cell do not undergoing any further change. * The important function of this zone is to help anchoring the plant firmly in the soil and to give rise to lateral roots from the interior(pericycle). * * Tap Root System * Adventitious Root System * * Given birth by radicle of the embryo. * Always underground. * Consist of single primary root. * Primary root produces distinct secondary roots, tertiary roots, and rootlets in acropetal succession. * Primary root is persistent. * It ay be surface feeder or deep feeder. * * Given birth by any part of the plants other then radicle of the embryo. * May be underground or aerial. * Consist of roots forming a cluster. * Primary root absent and there is no any type of distinction. * Primary root though develops but is shortlived. * If underground it is usually surface feeder. * The tap root modify for the storage of food called storage or fleshy roots. The primary root is swollen due to stored food in it and the secondary root remain thin.eg., carrot(Daucus carota), shalzum(Brassica rapa), chukander(Beta vulgaris) * STORAGE OR FLESHY TAP ROOTS RESPIRATORY ROOTS * * The secondary , tertiary or some time the primary root in leguminaceous plants bear small nodule-like swellings. * These swellings are called root nodules or tubercles that contain nitrogen-fixing bacteria (namely, Rhizobium leguminosarum). * The bacteria pick up the free nitrogen occurring in the soil atmosphere and convert it into organic compounds of nitrogen. * This relationship is beneficial for both the bacteria and leguminous plant and therefor, is called mutualistic symbiosis( or reciprocal symbiosis). * * Normally the adventitious root are fibrous and surface feeder( eg., cereals, grasses) but in some cases they get modified to perform special functions like storage food, mechanical support and vital activities .The modification of adventitious roots are as follow……………… * Fleshy adventitious roots:- like tap roots sometimes the adventitious roots swollen and become thick due to storage of food. Depending upon there shape and place of swelling s, these roots are classified in to following categories:1. Tuberous(single root tubers) roots eg., sweet potato(ipomoea batata) 2. Fasciculated roots eg., Asparagus, Dahlia * 1. Prop roots Prop roots arise from large horizontal branches of the tree. They hang down in vertical position. After entering into the soil, these root give piller like appearance and do not terminate into fibrous root system. Eg., Banyan tree(ficus benghalensis) * 2. Stilt roots Stilt roots develop from the basal nodes or the stem near the soil. They develop obliquely from the stem. After entering into the soil, they terminate into the fibrous root system. Eg., Pandanus, Zea mays etc.,. * BANYAN TREE 1.Primary functions Anchorage Absorption of water and minerals Preventions of soil erosion 2. Secondary functions Storage of food Providing mechanical support Help in floating Help to climb * * Many hydrophytic plants e.g., Utricularia, Wolffia, Myriophylleum etc., Do not develop any roots as they need little absorption of water & minerals salts that is fulfilled by the general surface of the plants itself. * Aldrovanda is also rootless angiosperm. * CONTENT Stem Buds Forms of stem Modification of stem Function of stem * * Stem can be define as that ascending part of the main axis(shoot axis) of plant which develop form the plumule of the embryo and is usually negatively geotropic and positively phototropic. * * Stem grows in length by means of terminal buds. * It is clearly differentiated into nodes and internodes. * When young, the stem is green and photosynthesizes. * When mature, the stem bears flowers and fruits. * It may bears hair on external surface which are commonly multicelluar. * * A buds represents a condensed immature or embryonic shoot possessing a growing point enveloped by closely place immature leaves. However, a buds can be defined as a compact young shoot consisting shoot stem overlapped by crowded immature leaves surrounding the growing point. * 1.On the basis of there nature vegetative Floral Mixed 2. On the basis there position Terminal Lateral Adventitious * * Areal stem Reduced stem:- Here the stem is very much reduced to a small disc which lies above the base of the root and is non photosynthetic. E.g., radish(raphanus sativus ) etc. Weak stem:- Here the areal stem are delicate thin, weak and thus are unable to erect. e.g., lableb , pasiflora etc. * * Underground stem:- Underground subterranean stem are those that lies below the surface of the soil. They are not aerial, non green, store food and contribute in perennation. E.g., Banana( musa paradisiaca) . * 1. Prostrate or sub-aerial week stem Trailers(=stragglers)., the main stem and its branches of some sub-aerial weak stem trails or spread horizontally along the ground without producing roots at intervals. Creepers :- are those prostrate or sub-aerial weak stem whose main stem and its branches trails or spread horizontally along the ground producing root at interval. * * Upright weak stem modification Twiners :- These stem are long flexible and twin around and upright – support like rope by a special type of growth movement (eg. Convolvulus arvensis ). The twiners can be considered as of sinistrorse type and of dextrose-type. Cilimbers:- contrary to the twiners that twin on the support by their own growth movement, the climber require the help of certain clasping of clinging structure(eg., Tendrils, Spines, Thorns, Hooks). eg., Smilax, Piper, Bignonia etc. * * Underground stem modification Sobole is an underground- runner. It does not swell as it does not store any food material and remains thin. Eg., Agropyron, Imberata(Ulau grass). Sucker It is underground modify thin stem. It may be said as runner like and underground runner . It is apex of the sucker that grows into an aerial shoot. * Rhizome Rhizome is well developed and well branched underground modify stem. Food material is stored in entire stem and branches. Rhizome is placed generally obliquely or horizontally in soil. Roots are produced all along rhizome. Eg,. Ginger. * * Primary functions stem bears and supports foliage, flower and fruits. Leave manufacture the food which is translocated through the phloem of the stem to root, fruit and storage organ. * Modification of stem for: a)storage (b) support (c)protection (d)spread and vegetative propagation * Secondary functions Stem store food in their parenchyma cells and underground modified form. Eg., Potato(solanum tuberosum), Ginger(zingiber officinale), Banana(musa paradisiaca). The underground stems like those of dryopteris, mentha etc. functions as orgons of perennation to overcome unfavourable conditions. * CONTENT Leaf Kind of leaves Types of leaves Phyllotaxy Leaf modification Function of leaf * * Leaf is a green lateral flattened outgrowth borne on the node of a steam or its branches and is characteristically photosynthetic. Leaves originated from the shoot meristem as leaf primordia and gradually enlarge . They are the most important vegetative organ of the plant as they synthesize food in them. * * Cotyleadonary leaves:- these are the embryonic leaves(cotyledons) emerging at the time of seed germination. Two cotyledonary leaves are present in dicots while only one in monocots. * * These leaves contain flower or inflorescence in there axil and are called bract leaves. Usually they are small and green but exceptionally in some plants are large and bright coloured. * * Photosynthetic normal leaves borne on areal stem or its branches are called foliage leaves. The word leaf is normally used to refer to the foliage leaves. * * Leaf base(=hypopodium):- it is lower most part of the leaf borne onto node of the stem. Usually the leaf base is border to get sufficient base base for attachment onto the node. * * Petiole(mesopodium) stock of the leaf that connect the lamina with the stem or its branches is called petiole. The upper part of the petiole remains embedded in lamina in the form of midrib while its lower parts is attached with the stem in the form of leaf base. * * Lamina(=epipodium) lamina is the terminal, thin, flattened, green and conspicuous part of a leaf and is specialized to manufacture food. Each leaf possesses its own shape, to margin, one apex and two surface. * * Venation refers to the arrangement of veins and veinlets in the lamina . Veins and veinlets, in addition to the forming the structural frame work of lamina or blade. * * Reticulate venation :- the vaniation is reticulate when veinlets is are irregularly distributed and for a network or reticulum in the lamina. It is the characteristic feature of dycotyldons. * * Parallel venation:- it is also called sriate venation. In it, all the veins are run parallel to each other and do not any network. Parallel venation is the characteristics of the monocotyledons. * * Simple leaf The lamina of the simple leaf do not divided into distinct lobes or leaflets. An axillary bud occur In the axil of simple leaf. Simple leaf develop in acropetal succession on the stem. This may borne on the stem in one or more planes. * * The lamina is distinctly divided into two or more leaflets borne either at the tip of the petole or laterally on the rachis. * An axillary bud may be present in the axil of the whole leaf but never in the axil of leaflets. * Leaflets are always borne in the one plnae. * * Pinnate compound leaf:- In this type of compound leaf, the leaflets are borne laterally on an elongated axis which may be the midrib or lateral vein of a simple leaf. * * Palm of hand like outline * Leaflet originated from common point * * Phyllotaxy is refers to the arrangement of leaf on the stem or its branches. * Type of phyllotaxy Alternate:- Only one leaf is borne on each node and the leaf of adjacent nodes roughly lie toward the opposite side thus giving a spiral arrangement. Opposite:- In opposite phyllotaxy, two leaf are borne on each node lying always opposite to each other. * Whorled or verticillate :- In whorled phyllotaxy, the three or more then three leaf are borne on a single node is whrol or circle. * * Though the main function of leaves is the photosynthesis , a leaf or its parts is occasionally modified into specialized structures to perform some other functions. Which are as follow :- * * Tendrils are wire like sensitive structure that help the plants in climbing by coiling around a support. They are usually unbranched. * * In some plants, the leaves or the parts of leaves are transformed into spines that protect the plant from grazing animals and reduce the rate of transpiration particularly in xerophytic plants. * * Leaves or lamina of some insectivorous plants get modified into pitcher to trap insects e.g., Nepenthes, Dischidia etc., * * Primary function of leaves:Photosynthesis:-it is the most significant single function of a leaf by which the latter synthesizes food material for plant. Transpiration :- leaves are the main seat of transpiration which create force or tension in the water column to cause ascent of sap.i.e., translocation of water or minerals from soil to the top of the plants via Xylem channel. Exchange of gases:- leaves exchange gases from and to the atmosphere mainly through stomata present on them. * * The branch on which the flower are arised is called peduncle, it develop in axile of leaf or on the terminal branch of the plant .The term pidcel is use to the stalk of each individual flower, attached to the peduncle. The specfic mode of arrangement of a group of flower on the peduncle is called inflorescence. * * The inflorescences are classified on the basis of position in the shoot system into the following fourtypes: 1. Terminal inflorescence: A strictly terminal inflorescence, terminates the growth of the branch. 2. Axillary inflorescence: It is terminal on a short axillary branch, which is reduced to an inflorescence. * 3. Intercalary inflorescence: These are terminal clusters that have been left behind by continuing apical growth of the main axis, which form alternately fertile and sterile sections. Example: Callistemon (Bottle brush) 4. Cauliflory: The development of inflorescences on older branches. Example: Theobroma cacao (Cocoa plant), Arctocarpus (Jackfruit), Cannon ball tree. * * On the basis of sequence of development of flowers on the peduncle the inflorescences are of thefollowing kinds: 1. Racemose (Indefinite or indeterminate) 2. Cymose (definite or determinate) * * Different types of racemose inflorescence(a) simple raceme (b) spike (c) catkin (d) spadix(e) corymb (f) umbel (g) capitulum (h) panicle S.No Character Racemose Cymose 01. Peduncle It does not end in single flower instead grows continuously. 02. Position of Flower are arised laterally Flower actually arised terminally flower on in all sids on the in one side only. peduncle peduncle. 03. Arrangem- The flower are arranged ent of in an acropetal manner. flower The flower are arranged in a basipetal manner. 04. pollanation A pollyinator can not pollinate many flower in one visit though flower are grouped more frequently * A single visit by a pollinator can pllinate a large no. of flower. It ends up into a single flower. CONTAINT flower calyx corolla androecium gynoecium * * Flower is the most beautiful part of angiospermic plant and arise latterally or terminally and the axile of leaf which itself arises from the node. It is the reproductive unit in angiosperm. * * Actinomorphic flower:- when flower is radially symmetrical, it is termed as actinomorphic. * Zygomorphic flower:- when flower is bilaterally symmetrical , it is termed as zygomorphic flower. * Hermaphrodite or monoclinous flower:-a flower is called bisexual when it contain both male and female reproductive organs.e.g.;china rose mustard etc. * * Cyclic and acyclic flower:-when the floral part of flower arranged in a whorl, the flower is caled cyclic, and if floral part are arranged spirally and not in whorl, the flower is called acyclic. * Complete or incomplete flower:-when the flower of aplant have same size ,colour ,and arranged of various floral whorl/organs, then flower are called regular and if any flower of plant shows dissimilarity in any of it’s part of trait , then the flower is called incomplete. * * * It form the outermost covering of a dichlamydous flower. The unit of calyx are called sepals.each sepls is green ,coloured leaflike structure in most of the cases. * When the sepals of flower are free from each other , the condition is called polysepalous. * When the sepals of flower are fused with is other, the condition is called gamosepalous. * * It occurs next to calyx in a dichlamydous flower. It is a colorful structure other then green, thus help in insect pollination to the flower. The unit of corolla is called petals . * When all petals of flower are fused , the condition is called gamopetalous. * When all petals of flower are free from each other , the situation is called polypetalous. * * Aestivation: The mode of arrangement of sepals or petals in floral bud with respect to the other members of the same whorl is known as aestivation. * (i) Valvate: The sepals/peta ls close to each other without overlapping or may be in contact with each other. (ii) Twisted: Overlapping is regular, i.e., one margin of the sepal/petal overlap the next member and the other margin is overlapped by the previous. * (iii) Imbricate: Out of five sepals/petals one is completely internal being overlapped on both margins and one is completely external with the rest of the members arranged as in twisted aestivation. (iv) Quincuncial: Out of five sepals/petals two are completely internal, two external and one has one margin external and theother margin internal. * (v) Vexillary: Out of five sepals/petals the posterior one is the largest and external almost completely covering two lateral members which in turn overlap the two small anterior sepals/petals * * Types of aestivation in corolla : (a) Valvate (b) Twisted (c) Imbricate (d) Vexillary * It is a collective term given to group of stamens or male reproductive organs present in flower. Each stamen has two part a long filament and a terminally situated bag shaped anther. Most of the anther of angiospermic plant are dithecous, i.e., have two lobes but some have only one lobe, i.e., monothecous,e.g. China rose. * * Anthology is known as the study of flower. * The term staminode is given to non-functional stamens. * A stamen is a modified leaf (microsporophyll) * * The pollen grains are produced in pollen-sacs. A sterile stamen is called staminode. * Stamens of flower may be united with other members such as petals or among themselves. * When stamens are attached to the petals, they are epipetalous as in brinjal, or epiphyllous when attached to the perianth as in the flowers of lily. * * The stamens in a flower may either remain free (polyandrous) or may be united in varying degrees. * The stamens may be united into one bunch or one bundle (monoadelphous) as in china rose, or two bundles (diadelphous) as in pea, or into more than two bundles (polyadelphous) as in citrus. * There may be a variation in the length of filaments within a flower, as in Salvia and mustard.