Schizophrenia and the Affective Disorders Chapter 16 See Table 16.1 Schizophrenia Symptoms POSITIVE SYMPTOMS Hallucinations Auditory/olfactory COGNITIVE SYMPTOMS Attention Low psychomotor speed Delusions Persecution/grandeur/control Deficits in learning and memory Thought disorders Disorganized, irrational Poor problem solving Negative and Cognitive symptoms are closely related, may involve dysfunction in similar brain areas Negative and Cognitive symptoms are not specific to schizophrenia Poor abstract thinking NEGATIVE SYMPTOMS Anhedonia Flat affect Motor Deficits Little Speech (poverty of speech) Social withdrawal DOPAMINE & POSITIVE SYMPTOMS PROJECTIONS Major CNS dopaminergic systems include: Nigrostriatal System (role in movement) POSITIVE SYMPTOMS Mesolimbic Dopamine VTA-amygdala in reinforcement/reward) Drugs that agonize DA Mesolimbic System (role Mesocortical System (role in short-term memory, planning, and problem solving) release (cocaine, etc.) can also cause positive symptoms of schizophrenia Brain Imaging ► Measure DA Rs in brains of recently diagnosed schizophrenics not previously exposed to neuroleptics Radioactive ligand of the D2 R was injected iv & its concentration measured in the corpus striatum Two Results ► Brains of schizophrenics have an abnormally high # of D2 Rs ► Higher level of occupancy of those Rs by DA ► Suggest both pre & postsynaptic abnormalities Too much release & too many receptors (modest) Side-Effects of Long-Term AntiSchizophrenic Drugs Similarly to long-term use of Parkinson’s drug treatments, there can be side-effects with long-term use of anti-psychotic drugs. Tardive Dyskinesia Tardus – slow Dyskinesia- faulty movement Late-developing TD: unable to stop moving Supersensitivity: possible that DA receptors become hypersensitive if they are blocked for long periods of time D2 receptors in caudate and putamen Problems with the D2 Theory Schizophrenia associated with brain damage Little damage to dopamine circuitry Damage not explained by dopamine theory It takes several weeks of neuroleptic therapy to alleviate schizophrenic symptoms Conventional neuroleptics (D2 blockers) mainly effective for positive symptoms Negative and cognitive symptoms might be caused by brain damage May be best to think of schizophrenia as multiple disorders with multiple causes Schizophrenia as a Neurological Disorder Predisposing factors (genetic, environmental, or both) give rise to: 1. Abnormalities in both DA transmission and PFC 2. Abnormalities in DA transmission that cause abnormalities in PFC 3. Abnormalities in PFC cause abnormalities in DA transmission Schizophrenia: Brain Abnormalities Evidence of brain damage Negative and cognitive symptoms Loss of brain tissue Lateral ventricles more than twice as large in schizophrenic patients than control subjects Possible Causes of Brain Abnormalities Epidemiological Studies Season of birth Viral epidemics Population density Prenatal malnutrition Maternal stress Possible Causes of Brain Abnormalities Season of birth (seasonality) Late winter and early spring (northern hemisphere) Reverse in southern hemisphere Possible link: viruses Effect seen in cities, not in countryside See Figure 16.5 Possible Causes of Brain Abnormalities Vitamin D deficiency Dealberto (2007) Northern Europe: 3-fold increase in schizophrenia in immigrants (equatorial regions) Thiamine deficiency Two-fold increase in incidence of schizophrenia in offspring of women pregnant during severe food shortage in WWII (Germany and Netherlands) Maternal/paternal substance abuse – smoking Complications during childbirth Mother: Diabetes of mother, bleeding, preclampsia (high blood pressure, protein in urine) Other: oxygen or blood flow deprivation Evidence for Abnormal Brain Development Home movies from families with schizophrenic child Compared to normal siblings, schizophrenic child displayed more negative affect and more abnormal movements 265 Danish children (11-13 years) were videotaped eating lunch Children who later developed schizophrenia displayed less sociability and deficient psychomotor functioning Hypothesis – although schizophrenia is not seen in childhood, the early brain development of children who become schizophrenic is not normal Age of Onset See Figure 16.8 Symptoms rarely begin before late adolescence or early adulthood Progression: Negative symptoms cognitive symptoms positive symptoms Abnormal Brain Development Brain damage is sudden (during young adulthood) Thompson et al. (2001) Adolescence with early onset schizophrenia MRI Normals: Loss of 0.5-1.0% Schizophrenic patients: 2-3% Loss: See Figure 16.9 Parietal to temporal lobe Somatosensory and motor Prefrontal cortex Abnormal Brain Development See Figure 16.10 Hypofrontality and Negative and Cognitive Symptoms Decreased activity of the prefrontal cortex (dlPFC); believed to be responsible for the negative symptoms of schizophrenia. Above fig: Task required increased concentration and attention Possibly caused by decreased DA activity in prefrontal regions NMDA , Dopamine and Hypofrontality PCP and ketamine can cause positive, negative and cognitivelike symptoms NMDA receptor antagonists; decrease DA and metabolic activity in frontal cortex Role of D2 Receptors in Development of Schizophrenia Abnormalities in the striatal DA system may be the cause of schizophrenia Virus inserted into striatum that increased D2 recpetors Caused the development of behavioral deficits characteristic of schizophrenia Abnormal activity of dlPFC Treatment with Partial DA Receptor Agonists Atypical drugs are able to reduce ALL symptoms Increase DA activity in PFC Reduce DA activity in the NA Aripoprazole: Atypical Partial DA agonist High affinity for receptor but less than ligand Can act like an antagonist Nucleus accumbens Can act like an agonist Prefrontal cortex http://www.youtube.com/watch?v=USxHsSWCaJA AFFECTIVE DISORDERS Chapter 16 MAJOR AFFECTIVE DISORDERS • Bipolar Disorder – serious mood disorder characterized by cyclical periods of mania and depression. • • • • Men and women in equal numbers Mania: last days or weeks Depression: typically 3x as long as manic episodes Major Depressive Disorder – serious mood disorder that consists of unremitting depression or periods of depression that do not alternate with periods of mania. • Continuous • Episodes MAJOR AFFECTIVE DISORDERS DEPRESSION MANIA Little energy Move and talk slowly May pace aimlessly and restlessly Cry Anhedonia Loss of appetite for food and sex Disturbed sleep Depressed bodily functions Sense of euphoria not justified by circumstances Nonstop speech and motor activity Delusions Full of their own importance & become angry or defensive if contradicted Long periods without sleep Working on “unrealistic” projects Constipated Decreased saliva production Chen and Dilsaver (1996) 15.9% of people with MDD attempt to commit suicide 29.2% of people with BD attempt to commit suicide AFFECTIVE DISORDERS: HERITABILITY Evidence indicates a tendency for affective disorders to be heritable, although many genes may be involved. Monozygotic twins – 69% Dizygotic twins – 13% ANIMAL TESTS AND MODELS OF DEPRESSION Animal tests of depression Forced swim test Sucrose preference test (test of anhedonia) Animal Models of depression Learned helplessness Social defeat Copyright © Allyn & Bacon 2010 24 ANIMAL TESTS OF DEPRESSION Porsolt swim test • measures the effect of antidepressant drugs on the behaviour of rats/mice Animals are subjected to two trials during which they are forced to swim in an glass cylinder filled with water The first trial lasts 15 minutes. 24-hours later, a second trial is performed that lasts 5 minutes. The time that the test animal spends without moving in the second trial is measured. Immobility time is decreased by antidepressants. ANIMAL MODEL OF DEPRESSION Social defeat • • • Repeated exposure to an aggressive male mouse Decrease in subsequent social interaction Increased time immobile in the forced swim test Berton et al., 2006 BIOLOGICAL TREATMENTS MDD • • • • • • • • • MOAIs SSRIs & SNRIs ECT Transmagnetic stimulation (TMS) Deep brain stimulation (DBS) Vagus nerve stimulation Low-dose Ketamine Bright-light therapy (phototherapy) Sleep deprivation Bipolar Disorder • • Lithium Anticonvulsant drugs BIOLOGICAL TREATMENTS 1950s • • TB drugs found to elevate mood First antidepressants • MAO inhibitors Monoamine oxidase (MAO) inhibitors • • • IPRONIAZID (e.g.) MAO breaks down monoamines in the terminal buttons Increases release of DA, NE and 5-HT (not specific) MAO inhibitors: Cheese Effect • • Dangerous dietary interaction Excess tyramine hypertension BIOLOGICAL TREATMENTS Tricyclic antidepressants (TCAs) • • • E.g. Imipramine Inhibits the reuptake of norepinephrine and serotonin but also affects other neurotransmitters. Safer than MAOI’s BIOLOGICAL TREATMENTS ‘Selective’ Reuptake Inhibitors • • • SSRIs: serotonin • Prozac, paxil SNRIs: norepinephrine & 5-HT • Fewer side-effects than TCAs • No more effective than TCAs, but side effects are few and they are effective at treating other disorders ‘Selective’ is relative as they still have effects on both 5-HT and NE though range in affinity: • Affinity for 5-HT:NE • Milnacipran 1:1 • Venlafaxine 1:30 BIOLOGICAL TREATMENTS MDD • • • • • • • • • MOAIs SSRIs & SNRIs ECT Transmagnetic stimulation (TMS) Treatment-resistant Depression Deep brain stimulation (DBS) Vagus nerve stimulation Low-dose Ketamine Bright-light therapy (phototherapy) Sleep deprivation Bipolar Disorder • • Lithium Anticonvulsant drugs BIOLOGICAL TREATMENTS Electroconvulsive Therapy Early 1900s: von Meduna (drug induced) 1930s: Cerletti (electrical) • Electroconvulsive Therapy (ECT) – brief electrical shock, applied to the head, that results in an electrical seizure; used to treat depression. • Antidepressant effects are seen quickly (unlike antidepressant drugs) • May exert effects through activation of inhibitory neurotransmitter systems • • Increased GABA & Neuropeptide Y Potential cognitive impairments TREATMENTS • Transmagnetic stimulation (TMS) • Strong localized magnetic field into the brain by passing an electrical current through a coil of wired placed on the scalp • • Deep brain stimulation (DBS): • Subgenual ACC • • • • 1 month – 35% improvement, 10% remission 6 months – 60% improvement, 35% remission NA • • Response rate <30% Reduce symptoms in ~50% Vagus nerve stimulation Low-dose Ketamine 33 MONOAMINE THEORY OF DEPRESSION NORMAL DEPRESSION TREATED Underactivity of serotonin and norepinephrine synapses MONOAMINE THEORY OF DEPRESSION Monoamine antagonists can produce the symptoms of depression • Reserpine (lower blood pressure) – blocks the activity that transporters that fill synaptic vesicles in monoaminergic terminals with NTs • Blocks release of NA and 5-HT Monamine agonists treat depression • TCAs, SSRIs, SNRIs, MAOIs Copyright (c) Allyn & Bacon 2010 35 PROBLEMS WITH THE MONOAMINE THEORY OBSERVATIONS: • Depleting tryptophan from diet does not alter mood in healthy volunteers • • Does in others on antidepressant treatment And in healthy people with family history of depression • 2-3 weeks for antidepressants to become effective • Some people never respond to drug therapies BRAIN REGIONS IN AFFECTIVE DISORDERS Amygdala: involved in the expression of emotion • • • • • Negative emotion 50-75% blood flow and metabolism in amygdala increases during depressive episodes Abercrombie et al. (1998) • Activity in amygdala of depressed patients correlated with severity of depression Metabolic activity of amygdala increases in normal Ss when they look at faces with expressions of sadness Metabolic activity in amygdala increases when depressed Ss remember episodes in their lives that made them sad BRAIN REGIONS IN AFFECTIVE DISORDERS Medial Prefrontal Cortex: • • Subgenual ACC lower level of activation (right) in depression Increased during mania Figure 16.19 THE ROLE OF THE 5-HT TRANSPORTER 5-HT transporter (5-HTT) The promoter region that codes for the 5-HTT comes in two forms: Short Long Three possible combinations: S-S S-L L-L The Role of the 5-HT Transporter See Figure 16.20 Major depression Suicide ideation Caspi et al (2003) Science 847 people for 20 years Stressful events (> # of stressful event; > depression) Increase was greater for those with the short allele THE ROLE OF THE 5-HT TRANSPORTER Rausch et al. (2002) • People with 2 long alleles tend to respond better to drug treatment Lee et al. (2004) • Depressed people with 2 long alleles treated with antidepressants have better long-term outcome Rhodes et al (2007) • People with higher levels of 5-HT transporter in amygdala showed less activation of the amygdala when the people looked at emotional faces Several meta-analyses of studies investigating the role of 5-HTT promotor in depression No significant effects! ROLE OF NEUROGENESIS • • • • • Stressful experiences that produce depression (in animals) suppress hippocampal neurogenesis Administration of antidepressant treatments increases neurogenesis (in animals) Delay in action of antidepressant treatments = length of time it takes for newborn neurons to mature If neurogenesis is suppressed by low-level dose of radiation, antidepressant drugs lose their effectiveness (in animals) No way to measure adult human neurogenesis ROLE OF CIRCADIAN RHYTHMS Symptom of depression: disordered sleep • • • • Reduced Sleep Latency Reduced REM Latency Reduced Slow-Wave Sleep Increased Sleep Disruption 1 of the most effective antidepressant treatment – sleep deprivation (total or REM selective) • • • REM selective - therapeutic effect occurs slowly, long-term improvement after deprivation is discontinued Total sleep deprivation – effects are immediate, but depression returns the next day Intermittent total sleep deprivation can have beneficial results SEASONAL AFFECTIVE DISORDER Attacks of depression and lethargy typically occur every winter Cause: Attacks are triggered by a reduction in sunlight • • Higher incidence in Northern US than in Florida Light therapy is often effective in reducing symptoms