Cnidarians “The guys with the stinging All About Cnidarians • Evolution of multicellular animals with tissues that perform special functions • Cnidarians: Stinging animals • Mostly marine • 10,000 known species • 2 body forms: medusa and polyp Anatomy A. Polymorphism- Cnidarians have more than one body form: 1. Polyp 2. Medusa • The life history of some cnidarians includes both polyp and medusa stages. Others spend their entire lives as either polyp or medusa medusa • Free floating form which is transported by water currents, mouth with surrounding tentacles are positioned downward Medusa • Umbrella shape • Tentacles around mouth • Motile, Free-swimming polyp • Sessile, attached form with mouth and tentacles positioned upward Polyp Form • Tube with tentacles around the mouth • Sessile Coral polyp Polyp (sea anemone) Polyp (Hydra) Cnidarians Cnidarians actually exhibit radial symmetry in which similar body parts can be grouped around a central axis. • Here we also observe a large jump on the evolutionary scale: tissues that perform specific functions. • Animals with radial symmetry look the same from all sides and have no head, front, or back. • They do, however, have an oral surface, where the mouth is, and an aboral surface on the opposite side • See figure 7.6 page 119 Radial Symmetry Radial Compass jellyfish tissues • Two layers present • Epidermis covers body surface (external) • Gastrodermis lines internal body cavity and is specialized for digestion • There is also a narrow, gelatinous middle layer, mesoglea,that usually doesn’t contain cells. Forms “bell” of medusae Two Tissue Layers 1. EPIDERMIS – OUTER LAYER 2. GASTRODERMIS – STOMACH CAVITY MESOGLEA – JELLY MATERIAL BETWEEN TISSUE LAYERS ABUNDANT IN JELLYFISH TO HELP THEM FLOAT Cnidocytes • Stinging cells • Nematocyst capsules located on tentacles • These are used for protection and feeding Cnidocytes-Stinging Cells • Within the cnidocyte is the nematocyst (which is like a capsule) • The capsule has the stinging structure which is a hollow thread (Filament) with barbs • Most contain a toxin • Very small, but discharged in large numbers • Usually the sting is only strong enough to kill zooplankton or tiny fish • But there are exceptions! • Discharged nematocysts, stinging cells used for defense and prey capture, have been the plague of more than fish. • Stings from certain jellyfish have resulted in death in a matter of hours, especially for infants. Discharging nematocyst • http://www.youtube.com/watch?v=6zJiB c_N1Zk Jellyfish stings • http://www.youtube.com/watch?v=Tp38DUjUnM • 1:01 • http://www.youtube.com/watch?v=SP6T B8kiZB0 Digestive and nervous systems • Digestive system is incomplete (sac-like with mouth only) • Centrally located mouth surrounded by tentacles • Tentacles capture and handle food • Mouth opens into gut where food is digested • Nerve net throughout body coordinates movements • Some jellyfish also have sensory cells and contractile cells One Opening System FOOD IN Food enters through the mouth and is digested in the cells. Wastes are excreted through the mouth. Sperm and eggs are released through the mouth. WASTES, GAMETES OUT Response • • • • No nervous system No brain Nerve net around mouth Nerves cover the body but do not tell the difference between body parts. Respiration • Via diffusion • Body is two cell layers thick Habitat Aquatic: • Most are Marine • A few are freshwater Feeding 1. Carnivores (predators) 2. Process of feeding Lion’s mane eats a. Tentacles sting prey with another jelly! nematocysts b. Tentacles grab prey c. Prey pulled into mouth 3. Prey moved into gastro-vascular cavity (GVC)* 4. GVC makes enzymes, breaks down food, extra-cellular digestion 5. Undigested food moves back out of mouth Process of feeding 3. Prey stuffed into gastrovascular cavity (GVC)* 4. GVC makes enzymes, extracellular digestion 5. Undigested food back out mouth *incomplete digestive tract (no anus) Locomotion A. Medusa- motile, free-swimming B. Polyps- sessile, attached to hard substrate Exceptions: 1. Hydra tumbles on tentacles 2. Sea anemones glide on pedal disc Reproduction 1. Asexual budding 2. Sexual a. Medusae release sperm & eggs b. Larvae are free-swimming Sexual reproduction • Medusa is normally the sexual stage with epidermal gonads. Eggs and sperm released from medusa • Zygote develops into swimming larva called a planula • Planula settles on bottom to form colony • Eventually, new medusa are formed Planula Life Cycle of a Typical Cnidarian • Alternation of Generations • One form is the polyp (a cylinder with the closed end attached to the substrate and the open end with the mouth and tentacles directed upward) • The other form is the medusa, freeswimming, with the mouth underneath (like a jellyfish) Life Cycle of a Typical Cnidarian Hermaphrodite sexual asexual Alternation of Generations Cnidarian life cycle • http://www.youtube.com/watch?v=ct9Ky Lmnu0I • http://www.youtube.com/watch?v=K9cF bJ8qYWg Phylum Cnidaria The Classes of Cnidaria are: • Anthozoa- Sea Anemones and Corals • Hydrozoa- Hydra • Scyphozoa- “true” jellyfish • Cubozoa- “box-shaped” jellyfish • Ctenophora- non stinging Cnidarians: Comb Jellies (covering separately) Class Anthozoa • • • • • • • All marine “flower animal” Corals, anemones, sea fans/whips, sea pansy Colonial polyps that normally lack a medusa stage Sting organisms around them Can move by crawling along the substrate Corals secrete calcium carbonate “shells” Most coral species possess symbiotic algae within body tissues called zooxanthellae Anthozoa- sea fan Anthozoa- anemone Sea anemone Giant Sea Anemone Anthozoa- coral Sun Coral Brain Coral Anemone Video • http://www.youtube.com/watch?v=NFO 4DwEkkr0 Homework • Research the relationship between clown fish and anemones • http://video.nationalgeographic.com/vid eo/animals/invertebrates-animals/otherinvertebrates/clownfish_amonganemon es/ Class Scyphozoa • True jellyfish. All marine • Free swimming large medusa forms with polyp only in reproductive life • They move by rhythmic contractions of the bell, but cannot fight against prevailing water currents • Many with powerful stings Scyphozoa National geographic jellyfish • Dance of the jellyfish • http://video.nationalgeographic.com/vid eo/animals/invertebrates-animals/otherinvertebrates/dance-jellyfish-eorg/ • 1 minute National geographic • http://video.nationalgeographic.com/vid eo/animals/invertebrates-animals/otherinvertebrates/jellyfish/ Super Jellies • http://www.youtube.com/watch?v=NbpB 5F9CcLc • 3:31 Jellyfish video • http://www.youtube.com/watch?v=B3fa XiTZdjo • 3:43 • http://www.youtube.com/watch?v=pimIb TqJLZc • 11:08 Homework • The case of the killer Cnidarians page 122 • Research most deadly jellyfish • Irukandji • Carukia barnesi Killer Jellyfish • http://www.youtube.com/watch?v=GK_ Cl_54Qh8 • http://www.youtube.com/watch?v=Ml7b O021DMk • http://www.youtube.com/watch?v=rQ2W YbJSvTk worksheet • “Killer Cnidarians” (Hoyle) Life cycle of Scyphozoa Largest known jellyfish • Lion’s Mane Jelly • bell of a lion's mane jellyfish can be over 8 feet across • tentacles can reach over 100 feet, and they have many of them - the lion's mane jellyfish has eight groups of tentacles, and there are 70-150 tentacles in each group Lion’s Mane Jellyfish • Encountering a lion's mane jellyfish probably won't be lethal, but it won't be fun, either. A lion's mane jellyfish sting usually results in pain and redness in the area of the sting. The sticky tentacles of a lion's mane jellyfish can sting even when the jellyfish is dead, so give lion's mane jellyfish on the beach a wide berth. In 2010, a lion's mane jellyfish washed ashore in Rye, NH, where it stung 50-100 unsuspecting bathers Class Hydrozoa • Mostly polyp forms with reproductive medusa • Freshwater and marine • Physalia • Portuguese man of war • hydra Hydrozoa Hydra • Exist only as polyps Portuguese man of war National geographic portuguese man of war • http://animals.nationalgeographic.com/a nimals/invertebrates/portuguese-manof-war/ National geographic port. Man of war • http://www.youtube.com/watch?v=RBd CpcapB0s Ecological Role A. Predators and prey B. Neurotoxins in medical research C. Coral – jewelry, building, reefs (surfing!) D. Coral reefs - habitat for many different species, great biodiversity, protect coastline E. Symbiosis with other organisms Cnidarians Documentary • http://www.youtube.com/watch?v=AjMWOeM3uA • 9:17