Stand Density NASP IMDS THE BIG THREE: • Absolute stand density • Quadratic Mean Diameter • Basal Area Graphical Guides Size-Density Relationship The predictable relationship between mean size of trees and their associated density in crowded (i.e., stem exclusion) populations Density Age Boundary Line Examples (AKA: “-3/2 Power Law”) Log-log Scale Growth Trajectories Wiley 1976 Stand Density Index Stand Density Index (SDI) – Woodard paper Reineke (1933) SDI Species-specific upper limit Density when the stand has a standardized QMD Calculated as: Imperial QMD TPA 10 1 . 605 SDI Metric QMD TPH 25 . 4 1 . 605 Stand Density Index Values Suggested Maximum SDI by species and source. English units are number of 10-inch trees per acre. Metric units are number of 25.4cm trees per hectare. Species White fir Red fir Mixed conifer for CA Douglas-fir for WA-OR Douglas-fir for CA Eucalyptus Redwood Ponderosa Pine Loblolly Pine Longleaf Pine Slash Pine Shortleaf Pine Upland Oak Ponderosa Pine Lodgepole pine Douglas-fir Western Hemlock Maximum SDI Maximum SDI (English) (metric) 830 2050 1000 2470 750 1850 595 1470 600 1480 490 1210 1000 2470 800 1980 450 1110 400 990 400 990 400 990 230 570 830 2050 690 1700 587 1450 790 1950 Source Reineke, 1933 Reineke, 1933 Reineke, 1933 Reineke, 1933 Reineke, 1933 Reineke, 1933 Reineke, 1933 Reineke, 1933 Reineke, 1933 Reineke, 1933 Reineke, 1933 Reineke, 1933 Schnur, 1937 Long, 1985 Long, 1985 Long, 1985 Long, 1985 Long, J. N. 1985. A pratical approach to density management. Forest Chronicle 61:23-37. Reineke, L. H. 1933. Perfecting a stand density index for even-aged forests. Journal of Agricultural Research 46(7):627-638. Schnur, G. L. 1937. Yield, stand, and volume tables for even-aged upland oak forests. USDA Technical Bulletin No. 560, 87p. Stocking Charts & Guides Size vs. Density Log-Log scale SDI on the “third axis” SDImax = 1000 Taylor Woods Level-of-Growing-Stock Study Flagstaff, AZ Relative Density Index Relative Density Index (RDI) Basically the percent of maximum SDI Communication across species (species have maximums) Intuitive feel Magic thresholds for stand dynamics 0.15 - 0.30 for crown closure 0.40 for scheduling of growing space thinnings 0.55 for entering “the zone of imminent competition” 0.80 for certain mortality agents Density Guides Density Management Diagrams Packaging the concepts together Simple management tool Old GSL “boat” diagrams Additional information on the diagrams INTERPRETING CHANGES 6” (800 TPA, 5.1”) 4” 340 TPA 653 TPA Density Guides Density Effects on Growth Density Effects on Growth Taylor Woods: Four decades of TREE DIAMETER growth; 1962 - 2002 4 y = -1.3146Ln(x) + 7.106 R2 = 0.97 y = -1.415Ln(x) + 7.6447 R2 = 0.9793 DBH growth (inches/decade) 3 2 y = -1.4821Ln(x) + 8.2557 R2 = 0.9873 GSL40 1962-1972 GSL60 1962-1972 GSL80 1962-1972 GSL100 1962-1972 y = -1.1867Ln(x) + 6.8112 R2 = 0.9767 GSL120 1962-1972 GSL150 1962-1972 1962-1972 1 1972-1982 1982-1992 1992-2002 Log. (1962-1972) Log. (1972-1982) Log. (1982-1992) Log. (1992-2002) 0 0 20 40 60 80 100 basal area (sf/ac) at decade's beginning 120 140 160 Density Effects on Growth Taylor Woods: Four decades of TREE HEIGHT growth; 1962 - 2002 10 height growth (feet/decade) 8 y = -2.3839Ln(x) + 16.391 R2 = 0.6455 y = -2.1886Ln(x) + 14.675 R2 = 0.6736 6 GSL40 1962-1972 GSL60 1962-1972 GSL80 1962-1972 4 y = -1.4543Ln(x) + 11.654 R2 = 0.3773 GSL100 1962-1972 y = -1.8418Ln(x) + 13.568 R2 = 0.6672 GSL120 1962-1972 GSL150 1962-1972 1962-1972 1972-1982 1982-1992 1992-2002 2 Log. (1962-1972) Log. (1972-1982) Log. (1982-1992) Log. (1992-2002) 0 0 20 40 60 80 100 basal area (sf/ac) at decade's beginning 120 140 160 Density Effects on Growth PER ACRE Taylor Woods: Four decades of CUBIC FOOT VOLUME growth; 1962 - 2002 1200 GSL40 1962-1972 GSL60 1962-1972 GSL80 1962-1972 y = 444.4Ln(x) - 1246.2 R2 = 0.8503 GSL100 1962-1972 GSL120 1962-1972 1000 GSL150 1962-1972 1962-1972 MCF Volume growth (cf/ac/decade) 1972-1982 1982-1992 1992-2002 800 Log. (1962-1972) Log. (1972-1982) Log. (1982-1992) Log. (1992-2002) 600 y = 278.49Ln(x) - 516.98 R2 = 0.7585 y = 240.24Ln(x) - 387.18 R2 = 0.8335 400 y = 209.37Ln(x) - 241.19 R2 = 0.8082 200 0 0 20 40 60 80 100 basal area (sf/ac) at decade's beginning 120 140 160 Density Effects on Growth Taylor Woods: Three decades of BOARD-FOOT VOLUME growth; 1972 - 2002 PER ACRE 5000 GSL40 1972-1982 GSL60 1972-1982 y = -0.2786x2 + 71.106x - 626.9 R2 = 0.8283 GSL80 1972-1982 Scribner Board-Foot Volume growth (bf/ac/decade) GSL100 1972-1982 GSL120 1972-1982 4000 GSL150 1972-1982 1972-1982 1982-1992 y = -0.3955x2 + 56.14x + 847.11 R2 = 0.6366 1992-2002 Poly. (1972-1982) 3000 Poly. (1982-1992) Poly. (1992-2002) 2000 y = -0.2407x2 + 33.39x + 1134.4 R2 = 0.31 1000 0 0 20 40 60 80 100 basal area (sf/ac) at decade's beginning 120 140 160 DENSITY EFFECTS ON GROWTH Tree vigor and mortality rates Plasticity of the trees – think about this at “Hoskins” LOGS study (chapter 7) Live Crown Ratio and “canopy depth” Crown Radius (branch length/size) and crown closure Height:Diameter and “stability” (e.g., in wind and snow) Wood utilization products; size and quality Understory richness, diversity and abundance Net Primary Productivity distribution Ecosystem “stuff” OTHER DENSITY MEASURES CURTIS’ “RELATIVE DENSITY” RD = BA √QMD RELATIVE or “AVERAGE SPACING”: (can be relative to heights, as well) TREE-AREA RATIO: area occupied by tree crowns per unit land area; TA can be estimated from D or D2 CROWN COMPETITION FACTOR; TA relative to open grown trees Spacing impacts Example calculations - Douglas-fir (SDImax = 600) QMD TPA BA SDI RDI CRD 6 50 9.8 22 0.04 4 6 200 39.3 88 0.15 16 crown closure 6 400 78.5 176 0.29 32 well-stocked 10 50 27.3 50 0.08 9 10 200 109.1 200 0.33 34 well-stocked 10 400 218.2 400 0.67 69 moved into "the zone" 15 50 61.4 96 0.16 16 crown closure 15 200 245.4 383 0.64 63 moved into "the zone" 15 400 491 767 1.28 127 24 50 157.1 204 0.34 32 24 200 628 815 1.36 128 impossible, would have self thinned 24 400 1257 1630 2.72 257 completely crazy idea Description low-density young plantation low density, but great tree growth impossible, would have self thinned well-stocked UNEVEN-AGED structures? SDI calculated by diameter class from a stand table BUT don’t stretch too far…why? 2 6 10 14 18 22 TPA BAPA 50 1 10 2 0 0 60 64 80 141 20 53 220 261 SDI 4 4 0 103 205 71 388