Antonie van Leeuwenhoek 1632-1723 Developed microscopes for observing living organisms 1674 discovered protists 1677 discovered spermatozoa 1682 discovered striated muscle fibers http://www.cartage.org.lb/en/themes/Sciences/Physics/Optics/OpticalInst ruments/Microscope/GlassSphere/usph_01.gif http://www.molecularexpressions.com/primer/images/introduction/leeuwenhoek.jpg http://students.ou.edu/J/Renee.E.Jones-1/Van%20leeuwenhoek%20Scope.jpg http://cell.sio2.be/introduction/images/microleeuw.jpg Schematic Human Sperm Cell http://academic.evergreen.edu/v/vivianoc/homunculus.gif Homunculus Human (Homo sapiens) Sperm Photograph Mammalian Syngamy: Plasmogamy Calcium release spreading across an egg cell during syngamy Amphioxus (lancelet) adult, zygote, and early cleavage not to scale! Amphioxus (lancelet) cleavage Amphioxus (lancelet) gastrulation and adult not to scale! Morphogenetic movements of cells during development Amphioxus (lancelet) nerulation Origin of some mammalian tissues or organs Primary Tissue Layer Tissue or Organ Ectoderm Mesoderm Endoderm Epidermis Muscle Lungs Hair Bone Liver Nails Blood Pancreas Notochord Digestive lining Kidneys Bladder Eye lens Nerve system Nasal oral mucosae Gonads Frog (Rana pipiens) embryology Frog (Rana pipiens) zygote Frog (Rana pipiens) embryo: early cleavage Frog (Rana pipiens) embryo: later cleavage Chicken (Gallus gallus) zygote and early cleavage blastodisc yolk Chicken (Gallus gallus) gastrulation Chicken (Gallus gallus) embryo after 4 days of incubation Human (Homo sapiens) embryo Human (Homo sapiens) embryo Human (Homo sapiens) embryo Human (Homo sapiens) embryo Comparison of vertebrate embryos of six species at 3 stages Ontogeny recapitulates Phylogeny Typical sigmoid (S-shaped) growth curve: logistic curve Weight plateau phase lag phase Age Weight growth in a mouse (Mus musculus) Puberty 16 Weight in grams 14 12 10 8 Weaning 6 http://www.c-wilkie.pwp.blueyonder.co.uk/jokes/images/mouse_sex.gif 4 2 5 http://www.sasionline.org/antsfiles/articles/images/Dino/mouse.jpg 10 15 20 25 30 35 40 45 Days after birth Weight growth in an insect (Malanoplus) with 5 molts adult Sudden increase when body lacks exoskeleton (soft shell crab) Weight (mg) 150 100 Nymphal instars: 50 5 4 1 zygote 0 2 3 20 40 Age in days 60 How does weight increase suddenly when not feeding? Multiples of weight at birth Differential growth in weight of the Homo sapiens body and its parts 20 Body Why does the curve inflect here? 15 Does growth continue indefinitely? Heart 10 Brain 5 0 5 10 15 Age in years 20 Average decline in a Homo sapiens from age 30 to 75 years Characteristic Percent Decline Weight of brain 44 Number of nerve cells in spinal cord 37 Velocity of nerve impulse 10 Number of taste buds 64 Blood supply to brain 20 Output of heart at rest 30 Speed of return to normal pH of blood after displacement 83 Number of filtering subunits in kidney 44 Filtration rate of kidney 31 Capacity of lungs 44 Maximum O2 uptake during exercise 60 bipinnaria http://www.cbu.edu/~aross/embryo/s-gastrula-2.jpg brachiolaria http://alpha2.bigelow.org/mitzi/images/library/subtidal%20ani mals/Sub-An-B_039.jpg planktonic larvae http://instruct1.cit.cornell.edu/courses/biog105/labs/deuts/media/echinowatervasc.jpg http://micro.magnet.fsu.edu/optics/olympusmicd/gallerie s/brightfield/images/starfishbrachiolarialarva.jpg http://www.snv.jussieu.fr/bmedia/oursinMDC/coursours3/bipinn2leg.gif Three larval stages and a sea star adult (Asterias vulgaris) gastrula benthic adult Monarch butterfly complete metamorphosis: zygote (“egg”) http://www.monarchwatch.org/update/2003/egg_glue1.jpg Danaus plexippus Monarch butterfly complete metamorphosis: larva Danaus plexippus Monarch butterfly complete metamorphosis: larva Danaus plexippus Monarch butterfly complete metamorphosis: larva Danaus plexippus Monarch butterfly complete metamorphosis: pupa Danaus plexippus Monarch butterfly complete metamorphosis: adult Danaus plexippus Monarch butterfly complete metamorphosis: adult Danaus plexippus Incomplete (gradual) metamorphosis in grasshoppers Brachystola magna http://www.ent.iastate.edu/images/orthoptera/lubber.jpg zygote Life span in years (log scale) Relative DNA repair rate among mammals 100 (Homo sapiens) Human (Loxodonta africana) Elephant (Bos taurus) Cattle 10 Hamster (Mesocricetus auratus) Rat (Rattus norvegicus) 1 Mouse (Mus musculus) Shrew (Suncus murinus) Relative rate of DNA repair