Winter Tree ID (Twigs) What is a twig? The book • Species organized into simple, meaningful categories*. • High-resolution photos of twigs, bark, fruits, etc. • “Similar species” list for each species. • *Some categories condensed/combined for this presentation. Learning twigs in 2 (or sometimes 3) easy steps! 1. What is the leaf scar arrangement? • Whorled • Opposite • Alternate 2. What type of bud scales does it have? • Imbricate • Valvate • Caplike • Naked 3. Other unique characteristics • Hair on twig or bud • Stout vs. thin • Zig-zag vs. straight • Buds clustered at the end of the twig Leaf scar arrangements Whorled (3 scars per node) Opposite (2 scars per node) Alternate (1 scar per node) Lay out your twigs in order, from 1 to 57 Leaf scars opposite or whorled (species 1-9) 1. Acer rubrum 2. Acer negundo 3. Acer saccharum 4. Cornus florida 5. Viburnum rufidulum 6. Fraxinus americana 7. Fraxinus pennsylvanica 9. Catalpa bignonioides #1 Acer rubrum (p. 14) • Usually reddish • V-shaped leaf scars that touch • Terminals a bit larger than laterals #2 Acer negundo (p. 10) • Green and lustrous • V-shaped leaf scars that touch • Deeply imbedded laterals • White pubescence on terminal (especially) and laterals #3 Acer saccharum (p. 18) • Brownish with distinct lightcolored lenticels • V-shaped leaf scars that touch • Sharply pointed terminals, variegated brown and white #4 Cornus florida (p. 26) • Dull and greenish • Large terminal, pointed and emerging from split twig • Round flower bud may be present #5 Viburnum rufidulum (p. 24) • Light brown to ashy-grey colored • Rusty red pubescence on buds • Terminals substantially larger than laterals #6 Fraxinus americana (p. 30) • Light grey to olive green colored • Occasional lenticels on recent growth • Buds deeply imbedded in “U” shaped leaf scar. #7 Fraxinus pennsylvanica (p. 32) • Light grey to olive green colored • Occasional lenticels on recent growth • Brownish bud sits on top of leaf scar #9 Catalpa bignonioides (p. 22) • Grayish green in color • Concave leaf scars are (usually) whorled • Skunky smell when freshly cut Leaf scars alternate, bud scales naked 11. Asimina triloba 13. Carya cordiformis 14. Rhamnus caroliniana #11 Asimina triloba (p. 94) • Brown in color • Dark brown naked bud, with velvety pubescence • Terminals much larger than laterals • Green pepper smell #13 Carya cordiformis (p. 102) • Yellowish brown in color • Terminal bud is larger than laterals and is covered in velvety pubescence • Looks naked but technically is not #14 Rhamnus (Frangula) caroliniana (p. 104) • Brownish twig • Naked bud with grey/tan pubescence • Terminals a bit larger than laterals • Musty odor when scratched Leaf scars alternate, single cap-like budscale 15. Platanus occidentalis 16. Salix nigra #15 Platanus occidentalis (p. 88) • Green to grey zig-zag twig • Single cap-like bud scale starts out green but turns red • Leaf scar surrounds bud. Bundle scars visible #16 Salix nigra (p. 90) • Slender, yellowish-brown in color • Single caplike budscale, beaked and appressed to the twig Leaf scars alternate, valvate (2 scaled) buds 17. Liriodendron tulipifera 18. Alnus serrulata #17 Liriodendron tulipifera (p. 108) • Shiny brown twig • Valvate buds, look like praying hands or a duck’s bill • Round leaf scar with stipular scar completely surrounding the twig #18 Alnus serrulata (p. 106) • Dark brown twig, possibly with some pubescence • Valvate bud scales, stalked laterals. • Triangular/semi-circular scar. • Male and female catkins often present Leaf scars alternate, twigs armed and/or buds submerged 20. Robinia pseudoacacia 21. Albizia julibrissin #20 Robinia pseudoacacia (p. 114) • Brownish angular twig • Submerged laterals • Stipular spines may be long, short or absent. #21 Albizia julibrissin (p. 126) • Zig-zag, greenish black with corky lenticels • Smells like green peas • Tri-lobed leaf scar, with tiny, partially submerged bud Leaf scars alternate, stout with large leaf scar 24. Carya illinoinensis 25. Carya tomentosa 26. Carya glabra 27. Rhus glabra 28. Rhus copallina 29. Melia azederach #24 Carya illinoinensis (p. 132) • Slight pubescence on brown twig • Buds dark brown, pointed, and somewhat beaked • Heart-shaped leaf scar #25 Carya tomentosa (p. 136) • Stout brown twig • Large terminal bud with blonde scales • Tomentum on recent growth • Elongated or tri-shaped scar #26 Carya glabra (p. 130) • Dark brown, almost black twig • Glabrous and thinner than C. tomentosa • Dark brown to black scales on terminal bud #27 Rhus glabra (p. 124) • Scattered corky red lenticels, twig generally glabrous • Leaf scar wraps almost completely around the bud. #28 Rhus copallina (p. 122) • Corky red lenticels and soft pubescence all over twig • Bud sits on top of a U-shaped leaf scar #29 Melia azedarach (p. 150) • Dark green or almost black twig with small lenticels • 3 lobed leaf scar with small brownish bud on top Leaf scars alternate, buds with numerous scales (imbricate), buds clustered at the end of twig 30. Quercus alba 31. Quercus stellata 32. Quercus nigra 33. Quercus coccinea 34. Quercus velutina 36. Quercus phellos 37. Quercus falcata 38. Quercus rubra #30 Quercus alba (p. 38) • Grayish or grayish-red twig, hairless, lustrous. • Reddish brown glabrous bud scales, usually with a rounded point. #31 Quercus stellata (p. 62) • Yellowish grey twig with pubescence. Usually stout. • Orangeish-red buds, with pubescence #32 Quercus nigra (p. 54) • Slender, with very divergent buds • Reddish brown buds with a whitish, pubescent tip (a little frosty) #33 Quercus coccinea (p. 40) • Reddish brown twig, a little darker and stouter than Q. nigra • Buds reddish brown, with frosted pubsescent tips #34 Quercus velutina (p. 64) • Very angular twig, generally hairless • Bud scales very hairy and tan in color. • Buds very pointed #36 Quercus phellos (p. 58) • Slender hairless twig • Small, slender brownish divergent buds. Also hairless #37 Quercus falcata (p. 42) • Hairy twig, otherwise similar to Q. velutina • Reddish brown buds with some hair #38 Quercus rubra (p. 60) • Very angular, bronzed to greenish twig. No hair • Glabrous, shiny reddish brown buds Those pesky oaks… Hair on the twig YES NO NO Quercus rubra YES Hair on the bud Quercus alba Quercus phellos Quercus falcata Quercus velutina Quercus stellata Quercus coccinea Quercus nigra Leaf scars alternate, buds with numerous scales (imbricate), twigs generally small w/small leaf scar 39. Oxydendrum arboreum 40. Prunus serotina 41. Carpinus caroliniana 42. Diospyros virginiana 43. Cercis canadensis 44. Betula nigra 45. Celtis laevigata 46. Amelanchier arborea 48. Nyssa sylvatica 49. Liquidambar styraciflua 50. Ostrya virginiana 51. Ulmus alata 53. Sassafras albidum 54. Tilia americana 55. Morus rubra 57. Fagus grandifolia #39 Oxydendrum arboreum (p. 162) • Two-toned twig, usually greener on one side, redder on the other • Small leaf scars, and small submerged buds. #40 Prunus serotina (p. 178) • Glaucous on one side, glabrous on the other • Variegated reddishbrown/green appressed buds • Bad smell when scratched #41 Carpinus caroliniana (p. 154) • Slender twig • Lateral buds appressed to the twig • Buds variegated white/brown, look like snakeskin #42 Diospyros virginiana (p. 160) • Light brown twig, shieldshaped leaf scar • Dark brown/black triangular buds • Crescent-shaped bundle scar sometimes present in the leaf scar #43 Cercis canadensis (p. 166) • Reddish to purplish zig-zag twig with white lenticels • Tiny lateral buds submerged in the leaf scar • Large globose flower buds sometimes present #44 Betula nigra (p. 70) • Slender twigs, usually with obvious white lenticels that feel like sandpaper • Sometimes pubescence on the twig • Greenish-brown buds w/white pubescence. #45 Celtis laevigata (p. 182) • Usually dark colored, zig-zag, with small white lenticels • Triangular brown bud sitting on a little “porch” • “terminal” bud is at a 90degree angle #46 Amelanchier arborea (p. 78) • Reddish brown twig • Pointed, elongated buds, ranging from green to pink to (sometimes) red #48 Nyssa sylvatica (p. 72) • Light brown to grey twig • Large terminal bud with small (or missing) laterals. • Purplish bud scales, usually with white pubescence • “sloth face” in the leaf scar #49 Liquidambar styraciflua (p. 120) • Light brown to grey twig • Large shiny terminal, with smaller laterals • Buds may range in color from green to brown • Sometimes has a corky wing #50 Ostrya virginiana (p. 158) • Slender, reddish-brown, slightly zig-zag • Sometimes fine pubescence on the twig and bud scale • Buds variegated green and reddish-brown #51 Ulmus alata (p. 184) • Slender and brown, may or may not have wings • Small, divergent brown buds • Crescent or half-circle shaped leaf scar #53 Sassafras albidum (p. 172) • Light green twig • Small appressed laterals • Club-like terminal bud • Spicy smell #54 Tilia americana (p. 86) • Reddish to green zig-zag twigs • Well-pronounced buds with 23 bud scales • Similar to M. rubra (55) but fewer scales #55 Morus rubra (p .76) • Grey twig • 5 to 6 greenish-brown bud scales #57 Fagus grandifolia (p. 74) • Yellowish-brown zig-zag twig • Elongated, very divergent buds