We are grateful to the editor of Conservation Biologyforarranging this Forum and for the willingness of suchdistinguished conservationists to respond to our essay.Our critique of sustainability has clearly touched a nerve.By way of rejoinder we would like to revisit themain pointof our essay, approaching it from a different direction, andthen turn briefly to the three responses.At its root, conservation is all about nature and the wayspeople interact with it. From a conservation perspective,some ways of using nature are acceptable and some arenot. How do we tell which is which? We can ask thisquestion in other ways: Where is the line between usingland and abusing it? Or, if conservation succeeded, whatwould success look like, in terms of nature itself?Currently, there is no consensus answer to these ques-tions, not among society generally, nor among conserva-tionists. Indeed, these questions are not asked very oftenand are not asked in ways that keep these questions sepa-rate from the many other questions that conservationistsfind important.As posed, our questions are designed expressly to focusattention on the core ends or aims of conservation, havingto do with the physical condition of nature. Everythingelse about conservation, in our view, is secondary to theachievement of these ends. Conservation is about livingright in relation to lands and waters. If we do not get thatrelation right, judged in terms of our effects on natureitself, conservation has failed. Conservation might also beabout other things—about living right with one another,for instance, or living right with future generations— butit is first and foremost about the direct human–naturelink. If we get that wrong, we can never declare success.A central reason why we do not like the concept ofsustainability is because we cannot extract from it a clearanswer to our questions about humans and nature.