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rn~w
A
ZEAIJ\.ND
A the
:;nlbmittet'l in
Ac;:ricultural
in the
University of
(} •
• RothVle 11
Gollage
1
CONTENTS
OHAl.')TER
1.
.
2.
REVIEW OF LITERATURE .
.,
...
PAGE
.,
. .
...
. . . ., . . . .
1
4
1•
Skin Structure and Follicle Anatomy.
4
2.
,.
The Folliole Outer-Root Sheath • • •
9
4.
Anomalous or tlDoggy" Wool
Folliole Capsules
• • • • • • • • •
14
•••••
17
5. Staple Crimp-Fibre H.elationships • •
23
6.
Relationship Between Fibre-Type,
Follicle-Type and Fl•• ce-Type
7.
•••
28
Pattern of Arrangement of Orthoand Para-Cortex
•
• • • • • • • • •
·.. ..
MATERIALS AND METHODS
.,
32
37
1.
Experimental Flock. • • • • • • ••
37
2•
Sampling. • • • • • • • • • • • ••
39
".
Laboratory Teohniques
RESULTS
• • • • • • •
•••• • • • • • • • • • • • • •
Group Data
1.
Mean Age
2.
Relationship Between Crimp Soore
and Tip
4.
Score
•
• • • • • • •
• • •
55
Incidence of Capsules With Age • • •
57
"
• • • • • •
tI
• •
in Relation
to Crimp Score and Tip Score • • ••
60
- ii PAGE
CHAPTER
4.
5.
Relationship Between Wool Scores
and Age • • • • • • • • •
6.
RelationShip
licle
~d
.
. ...
65
Type of Fol-
Betv~en
:'rcsence or Absence of
•• • • • • • • • • • ••
68
7.
Secondary:Primary Folliole Ratios •
69
8.
Follioles With More Than One Cap-
Capsules
aula
9.
69
Distribution of Ortho- and ParaOortex
10.
. . . . . . . . . . .. . . • • .
• • • • • • • • • •
pedigree Information
DISCUSSION
•.
•• •
• • . • • .
•
• • •
70
•
• • •
71
...
..... ..
ACKNOWLEDG&VtENTS
....
....
SUMMARY • •
..
REFERENOES
APPENDIOES
..
72
87
89
90
•
•
•
•
-
000 -
*
•
•
.. ....
96
1
is
, 1
a
some
.
an e
ace
(
no
(
.
,
... 2 -
more
less than that
t
the remainder
i. ~J no t:1 au1rt;
only
'been
by the
tion
attia.checl to
senGO of 0
of
.lll
fl
the
follicles
a:ncl
aso with
'1960)
s in
a
\'!laB
thJ2 crimp form
f ou.llcl
, 1
.,
(1
)
s att;ached to
ancl
'i
1
had also
i:n the New
(1
13.b..
OI'
oapsules were
Australia:n.
011
of
mind,
)"
- :3 te:t:'~·
folliclE?t
sence of
'-'-'n~"'~ fi ca:n:t1y
H, th.ere
of
A
D.
one
in the
of a
seri.ol1s
on
OlUil
invol1rsd
8.
survey
s fault;
Z()aland.
"'" 4 -
2
1
cell
"
),
co;rnemn"
)
011
-.5other
)
of
or
onEil"
A
IT
1\
1.1n.-
is
1
10080
wool
in
t:issue but;
bl
- 6 -
(
)
~
(1
)
f'ol.1icl(:; of
IHo(;l
('I
(1
(19'
r:()')
.
/:1-
Detailed.
shave
(19;58)
(1
(1
)
(1
i'oll:tcl(H3
of' aace
de va 1 opme ni; (
;
follicles
.
- 7 the tWCl typos
is no
produce"
s in the -presence or
glancls
OU8
follioles in.
malll-
!!lals ..
11cle::'4 in some
{)x
some follicles, '
1959) ;
, 1
and.
(
,
bandiooot
,
(
as in
Ar:roctor
wi 'l,b. some
and
OJ:
) 'll
); or with no
\
1
l)e
s
f
chinchilla (
,19(5)"
the elephant se
(1
as, as in
with
IS
(
as
(
, 1959).
and
wool follicle as an.
fi1)l~e
dermal
in
OIL
"
in
1"
homogeneous
2 ..
A. continuous thick
vitreous
of circular
tlssu.e
s ..
o.i
3
ssue
e wall
1 aye 1: of' 1
s ..
zones:
conns
1•
the
Oll.
.
s of the
they are
1
t
S
on
bulb
end
ell is
connective
the 'bulb ..
bulb in
more or
on
thi
".
fro:m
rooti
i
the
corneum
ous
- 9
of
the swaat;
in
.follicles
on, close to
wool
1i
on" .
are
t;ho
::tn a
c
ons ace
can
on
its
013111310
in
a
which
is
the
8
d.ivided. into three
(
OTIS
1.
(
t;hin
the
bulb
CfHl
on
- 10 -
In
a level
it
extends
on "the con.cave
s of this
not
(liffer from
In
.,
tion 8,nd :tor-In
~iI,
bu"lge is con:tinEhl
deflected
, or more
t;han (,xn
on, -l:Jhe
siele ..
In
even
Itlembra:l1.e
OU.
into
form. one or
con-V'ex
concave
tllf,;
Ino
re
ased
with n
the "thickened
in u"eile
of the
into thtl
c
aonditiollS.
an e.ctally
Ii
aU.tol:"
decreases
jects
slig;ll tly or i
ental sVle
kl
of t;he calls
ac(~nt
.
to
€l
It
on (b) are
sheatl'l
1/
- 11 -
of
ow ttl,s
( c)
C OlJUllEH10fc) S
t3.!ld
B
until
region (0)
tb,$
"
found
In
root
regio:n
to the
bulb
S OOOUI:'
( )
1
equator
t;he mi to
B
is
atb in
more
on.
primary
is Closely
pili
on
in m::CUl (
1
..
outer-root
) 1
ons for
and,
It
the
as
:fOJ:'Jt!
on
Fl:oweve:r' Urma (1
s
l~lCl(in.g
the
wag sub-
an
,
)
"I
folliclfH3 (
of' the .Bandicoot; (
'.Pili mnsGle.
svmlling
oue 1.n follicles with
!nore
is also more
"
to Wildnu;ln ('I
the swelling
in
thH fibre
i~, d, rllHre
bulbs ane," th,fl.t
follicle~)
the" a.traction,
tion of tihe bull) ..
sence in
follicles W:LtJl
wi th ect;ally
:Ln<iim.ltes i tn
rlef1.oct;i()I1 (Aubel', 1
outer-root
concluded
flection ..
of
no evidence of a
the
suggeste<l
inner-roo·t
on"
ai
)
,
- 13
cells
ou't~r-root
of
..
(Cabb~3'i 1
()vittsnce
to
a,
by many
be
fI:hey a:c'e
Montagna, 1
a firm conna
on
( Br'W"'lJ:l ,
o1J
ons su.gge
pull
ath,
cells of
possibly
(1
(
outer-root
( 1964·)
hair
in
)
cell~l
motts()
in
aUl
lnitotic
,
par.t
of
a
o:f
sloughing
1311189:011,
just
iihe
-bhe
d.uc·(j "
a .stud.;]
(1962)
a.
b.ow
zone
level
brerucs
cells
sheath
sCJ.'"'ibed a
llarrOl"i
eous
oute
lov" the
duct.
this zone,
thE)
t;lw
sheath
tQ
s, finally
m,ass
at;
so
zone,
(
, 1
1
re-
7
or t tho
) was
so were
fully
fl.
more or
1
(19
- 15 numbers
a
the folliole
s which heii
oapsu10B
of
de[~enera1;ion.
to
from
():I'lly
elip;ht abno
The capsula
(1950)
by
to
two al terx1.ati va
sru.ption of
anti
,
all
fUld,
nOI'Ii1al
It CI;3,y:,
sule s If
1 irst
1
fo11io1e[1
s of
ona or m.ore
t;:tLlI3
IIcaps1l1ee1l
se oapsules
follicle"
may, in
, :have
with
follio1e region was
corneurll,
such
slocated
~process
s of t;1u.';
borne
of
O:rl.
(1
by
follicles
size
c
ort/sin
)
mouse
~a.fter
all
c:;rslJs of a
CarCi!logenic
agef}:lis"
T}I'OCeSl'3 of
by
11
is
11
eapaule s"
(Ryd.er nncl
the
, 1
- 16 , 1
, 1961)"
to an
on
"
by
s.
101' the
II
G
of folliole
of tbe
, 1
on wi tll
a
follicle
E\r(~
become
1ele
OUl'::l
bei;wTeen those outf?;rowth cap
or
o
'J:his
of
, 196413.).
s that;
(19~}O)
of
are
II Oli'2.'P
all.l e s \I
AulH:l:t" (19
with
)
a
or
(
II
(1
'1
)
to
t;tlo
lindi
II
one
) and
\
)
.
if
fibres.
llH}Veme nt
folli
s with,
as
Hct as
are
or
t
1')60).
'<tlJool
1
(Ghap-
are
like
clip
ar
- 18 -
the incidence
s"
s-
as
t
of
(1926)
OJ:'
(1
arSJ1Ce"
1
, 1
19 , 1
(P
<
)
,
)
(P
wool
( Ct ly-.nl1e ,
<1%)
, '"1960).
to
suint
and
1 •
Cloned.
'1
)
(
~
sence
It
on tho
"
or J1::ngli
arlIn];
(
an
on
1
as
1
W!;lB
on the
t
.,
1
, 1
)"
p
wave:3 or even crim p
(
)
.
in
)
se
- 21 -
s (1
m.o:re
?OO
bulge
of
,,,. n " mil-<
S
(ill
the .fibl.'le is
seen in
ctecl
1
one to
Hyland, 1960) ~
whioh
fX'QIll
to
thE1 cause
) concluded
man
fibre "
the
:alO!'€}
s t;o
U
effort
at on
:1
ell'!.
to
c
a of
is no
1
in
1
)"
have a crimpo(i form
fibres
the case
folliole,
s d.ue to
the
(
; Maule, 1
; MaY'stou, 1
i
Ryder,
, 1960;
, 1
(lef:tci
The;} a.if
or
n
'U
d,
wool
tensile
s
mor})hometry (
Auber (1950)
shovm that wool
(1
(1
)
)
o
wool
of'
,
form on. tihe
on
:f
)
involving;
~j
s of
vllithin the
)
ll'ol a1
on
tue:t:i.on 1:;;}
Oll ..
.fibre
forceEi
follicl~~
S ('1,1..1.0
to
0
on of the
pos~::ible
0[;1,1..1.S0
(
and
(lLstortion
form a:nd a
, 1
, 1
presence
or
ItVIlli d.i 'by
It; is
more
a fibro
dLfficult
staple is
't;;obe in
the
in the
all of
It is nena
it is
sively or
of
to di
f1 bra f
OI'1n ~
:fibre
2 ..
~Phe
staple,
"
"
fibre
the re
( GoldsWOI;thy
atortion tllat
.
Vloods (1
neohe
It
or
- 25 -
..
are
(1
(1
t;c assess
,
s
)
is
8.
(1930)
is
(1954 )
(1
fibres
a re
.
1 <I
l!
II
~
?
two can
of the!!la :t s
in
, or.
lnino:ci ty
on as one
o~ity
the staple.
wools as
13
., co€/,rse
'tend. to
Goldsworthy
Gold.sworthy
the
within
dimensionf1 were thought neces
banded
butt
to measure
on
length d1
were more
leng'th.
s
- 27 in
be
1
in
I:;
section
Sf)
rae an,
could
in
s of
, the meDll
c
per
d
wool.
causod
, 1
rail.'
th.at
is more
or
4 ".
.i.wJ
(1
)
fJion,
- 28 ..
ar
length.,
Merino
and mean fibre
']:ha!39
st;B.ple
7
1
.
~
so
of
cl"irnped
is
0.1'
nor
s of mean length ..
th.at
bet;ween
is a
suoh as tihe
t
13
s
t;he Merino,
dif'ferent
fOllicle
ty-r,hlBS
number
alBo
s variation is reI
, 1
different types
(ii
in
birthcoat
sheepo
/i3
fleece
The
improved
of
1)eiJwt3en fOllicle
i IZ3tics
sand
int;eracti ons
01
that
fibre
avol ved through
type.
~:md
lOIlI;?;Wool breeds, wh,ieb
ar to
the
double
s from
unifol'fm
llomney t
t~nd
groups (
1;'ri thin
the
~l
:medium
a113,11d
ced.
shee]? in which "i;:;he p}.':i.mary :fo11:1.e10
tin(yl;ly eoaroer
t~rO'Ll]? i11
the
~rl;a:pl(;J
f:i."t)ref~
th8.Jl
formed, a
dis·~'
title seoondo,ry foll:L·,
1961) "
tho ,mecliulll
flG(H~0(1
i1Ilcesto:I' eould
i.:ricrease
stirtot;io:n. boi;v{('HJH long\7m<)1 and double
lOllgW(lOl "
'I:he reduction in
S:L<:H1
and grov.rth
J:'at~~
of fibres from
grow 1:w.lo-ha:L:t'B an<l t,heae are see:n. In:'ojecting above the I'ost;
of t;h0 birtilwoati.
fibres
11:he reduction in size 811(1 growth l"ato of
ch~';1.racteri~~jtic
of' the double
cO(:i,i:;ed bro€}dEl
simil&.r in both ·the double coatHcl fJheep and
breed.s although the modul18:bion of t;hese
H0m11,ey. a pril1u,t:Lve longwool
br(H~d,
thE~
fi'br'~s
@
111he
longvvool
may be
show tl1c},t tIle growtl1 of
30 in thH
fi1n'es is f
on.
the
sons
i,E;l
in.
'l'his
)
be
in oom,pe
the
.
l.fJ
evolutiorl of
fe
the
S::l
(
to
, 1956)..
A s a re
sed from 5 - 6: 1 in
1
in
to
follioles in
the
at
60-
nun2
on the
)
re
wool are tha,t
stiCH; of
fibre
are
sand f::Lln:'e s
s are
vnlich is often seen
to
(Hyder
t
other brae
1
on
are
(D:r:'y, 1
a:nd, oocur
ted
th.e
II
'
A
is
Short, 1
(
"1
schinckel. 1
of a coarse
(low 13/1:')
low
Bhort, 1960)..
f 0111
on
is more marked (
rJ:his is due to
s on the #type
at on
fleece
a,me
breeds, i3uoh
£U3
an increased vari
le:ng~th
cause a
a,
of
and ti1
sta~p'le
a
to influence ~3
non uniformity
for smile tirafil.
on
t1118
(1
configuration
SJ:J
a
h.11m:l. (1:1. t;y ?
swellinf~
other ..
dyeing
in
furthermore,
between the two
parti on of. i;he
s (194-1) outlined
(1
reaction
clifference
th~)
a tvlO s:tclEH.l
s in
folliole\>
Haria
(1953)
stence
bilateral
now
mR'p1y
that the
- 33 ....
into
s is
of
(Horio
, '1
to as
c
(soft or
the
)
,less
soribed
as
t\r~ro
torm
fibre
110a11y.
WOllrlCl
of'
s (1954 ) f O1J.net
the
th.e
this
by
crosso OIl.-
(1955)
of
1 ..
wool
In
in
clemarcation is
the
Alwve
steel:
be
ss
of
j1.lsti,fiable on the
s
s-
[Uld.
are
so
1
(
diV:L
sowoll
)
two
.
- 35 woOls, or
cells
result
occu.rrence
i
to tlle view
the
is
of
on of" the
is a
on in
layer
dave
"
seve
(1
.Ahmad
di 9tr:1. btl ti OIl
in C08.rser
II
\I
.Jt~ffries
(1960) ..
IS
(1
(1964)
on
the f01l10l(J bulb
bY'
distribution
(1
)
, in
IS
with the
of
that the
view,
reconcile
/3tri bution
on
to
in
(a)
the
on an
was a
soil
ewes
of'
"1
<-
'-)
aI'S
1
3
II
"1
4
II
1
5
fI
'1
6
If
waaAugust 1
2
"1
-17iih 1
"
et~1,
low 'Wool
''t
hi
home
,
ewes)
ws.s
were
1961
s was of
1
srune as
ora
at wltich
e:n.oes were
ation due
all
(b)
run
five eQ'le
s ,1:1.et with
feed,
the
"~Tinter,,
lSi
a.s
Autumn saved
Down raml3
'I'he
on a
was one rllonth
em.
COlItmenced.
s
months
is
exoellence
COIM10n to
f'aul ts
the d:1.
(a)
to
e:x.peri1l1ent, an
ze
axu,\lyse 20 sample s
Vias decided on ..
(1
the remail1.:lng
for
c~o
were
sheep
were aeleeted
sampled
ba08.11Se
sldn sample s were
ewe.
1
1
1
1
1
2'7
- 40 -
left
on
8k1:n
ruml! area. was
!ol11olf~s
rump
al"'e fil"'st
, 1960) Ii
and
OIlS
1\(;1
o1;rt; ain an
of'
................. Y's
,
sat
as
senCt';) of
(0)
s were "tmke:n
a an<:l
skir~,
of
; 1939) ..
1CIfl (
ath
fixed
1
when
f'rom
(Gu:t'r,
sheep"
fluid
for
of
back
- 41 -
Plat e I .
Crimp Score Standards.
- 42 -
Plate II .
Tip Score !itandarda .
s.
(
(i)
wool
WGx'e
crimp
ware e
on a 1 - 4shed.
of'
these
score grades.
il·
;}
2
1
ss as
(Chapman
for
s obvious
is
with
"
c
on
- 44 as
the
it
(
(Chapman
)
.1'or entry
case in
sampled,
score and
score should
throughout
age
alter
does
in one or
criteria.
words
"
scores will therefore
to
cally
to
some stage
to soum environ-
contribution
:tnfluence"
nT",,,T':'
the
s would.
of
relation-
two
=1- ~~ola te (1
al
the sa.JJle
were run
the enVirOXlJllental e
or! t;eria
will also
on t;h.e
constant
age
(ili)
sooras were estimated
the
9S
i:ng the
of
"
of
Il~tanda:r.'ds
within
was
by the
and
- 45 -
(lv)
crimp scores.
-
0
MU
-1.16
1
? ,,0
Q
7.£30
Sheep x Obll-3erVel'
226
58.2
0.26
Within Ob::3erver
2'::l?
40.0
0.18
fj::O'I',IUJ
680
;69.0
CICt
s1!
Hheep
226
Observer
**
,,>0
')(-'3
b,}
~G.
_
HI
6 .. 44"'*
1.1-3 .. 3*
*'
1 • '-I.,1I. ""
P<O.01;
Wr.1.~ll a
fllllel....e
highly sigrdfice.nt difference 'between
ob~3erVeI'fJ
'II)
(p <O.01~1~ H.:nd
/';
'by on6
€I.
obsI3rvcn:~
si€:!;:nificWlt d,lfferenee between observations
('I'able IV)"
ti()l1. of' t11e soores..
One observer scored consisten'bly higrler
th.'3.n th(; other al-bhough the trend. in scoring i s
sim.ilu~c.
fella important consideration is the degree of ag::C'eemen:b
a significa.nt difference t a repeatl:"!.bility estimate of 0 . 662
was obtained from the formula t
1966) ,
~[lhis
repeatahili ty estimate COt'lJH,),reS favourably with.
other estimates of rSI?eatabili ty
ti va sooring
~
G
observutio:ns by' subjec-
Niorley (1951) gave a figure
t "" 0 .6fJ6 fol"
(1
)
0.614· (rruI\6)
a
( I'talus)"
II
0 .. 47(3 (ewe
(ewes)
cover"
was
by
1'3 ..
s.
i:
347 .. 5
13 .. 1
1
,
1.54
13
.A
0"
51.,,0
0
479,9
V) ..
t "" 0 .. 701 w'as
II
11
.A~J
was
score
)
(i)
(a)
s
were
7.
'"
59 ..
*
1
$
total
from
of
five
total
100~
The
folliole
the
d. by
('1949)
each
, 3,'0 all
Is.
s were removed trom
with
al)soluta
a. triangle
s to
6
Ii
hours)
Gurr'
ss of 12,r
011
a
flOX'es
(0)
, 95
(
,
70 pEn:'
.
"
, 50 per
the
- 48 -
50
"
1
5
(95 per
)
10 ..
.
01 ..
Glear
ens were
(0)
used was an
(
was as
t
1954 )
..
OWS:-
1 ..
24·
all
come off
on~
2 ..
01.
(
lIed
Aoid
4 ..
OIl -
20
II
)
~
cent - ?2rnl
)
- 8ml)
wa:tel' ..
\
7 ..
5
Blua l::\olution
)
) - 19
)
)
2.
- 1
10.
(
)
11 "
12
- 0
5
Eooin
(1
GlefJ..r
x;ylo1~
"
~
)
(
rolliolH
(1949) "
i\.
V!~lS
OUS
an
and.
6/0.18 objeotive (
IS!.
)"
the screen was
piece
on was
caleulat€Hl from the
catj.on
:-
on x
-;;~
on x 1 .. 2 screen
was x10a.
on
area was
s area
011"
each follicle was
ons were then
point to
on
the
ViT,lS
on a
f3UCC(~
on
po:pul
As
oap
with
s were
which.
were
enee
a
ation of
level
s
- 51 comple
',F'rOlli
could.
1"
It
11
one
more tlU,111 one capsule
capsules
(b)
s of
8
was a.chievect by
20
the 1
level,
on
type
map
4
cent
(11
and 14 1
for
d.t
on ..
were
eli
IE!.
modi
by
(1956) "
or axis or
(1)
:ts
if
(
)
axis.
(1
)
not on
(
.:. :.: .
(2)
) Diffuse
. . . ....
....' .'.'. ...'. ..,
, ... "'.
•
'.
I,'
•
~.
~
~
'
di
(
) Ii
chwmel
occurrence
pe ri phery -
) it
the
<,q·O
pe:r:
area...
o
aircular occurrence of
by a
at
12
.
s was
on in £i 'bra s
al:' (')ld ewe s was
on.
SOllie
(d)
on
G/AO.
a
(2) x 6 objee-
wera
t
1
follioles
0
of
by
(e)
folliole
senee or
vIera
IE!
was 112 ..
4
soores
comes
d
1961
1
1
1
1
1 ..
-J$
+('
2.
&;:.
± C8
2
2"
2." '"
2.
:iO.
2 ..
+0
.-
7,,0
± 0,/17
2 .. 19
± 010
:;SIlO
+ 0 .. 67
2 ..
",.
~
! 0 .. 79
B ± ()
-
+ 0 .. 94
+
16.5
10,,9
0
.
-8.9
17 ,,30 ±10~6
,,12
6 ..
0
- 55 -
1J~his
l'elfli;ioru3hip
inoluded all
2~~?
were cl:l.lc.ulated
WL1.S
examined in eaoh ag;e group, an.o.
sheep sampled@
(2~a'ble
VII).
Oorrelation coei'i'ioientr::l
Ino,ividual sheep
f300res arE)
shown in ./'l.PIiendix II ..
Oorrelation coeffioients betvJeen crimp score
Ilnd tip score _ _ _ _ _ _ _
__,,_._,_'<!-._________
.
If
._m_¢"'=~_'
~I~ ... _
1961
-0 .. 12
196~?
-0.17
1963
0 .. 12
O .. l~O
~196l~.
"II
1965
analysis of variance..
'l1hese resul us show that there
Wt~Ul
no signifioanoe correlation between crimp score ancl 'tip
score for
EnVElr3
born
in.
1961 '*
196~:or
1963..
r:t:he corr~~la­
tion ba"blNeen oriln];> seOl'e ancl tip score we.s posi ti va
xligltly
si~~nifioant
(1' <. 0.01) for ewe s born
19fA, a:n.d.
positive and significa:nt (p (0 .. 05) for ewes born in 1965·.
-
saore"
1
on
0.,
30.0
.. 5
1
0 .. 48
15"
o
01162
16 .. 10
on
sidual
Total
1
4a
0.51
49
3'+.90
1
48
7
7",6
9
**
6 .....1
!It
0
4'7.,;4
1
9
;8.6
43 .. 5
4·9
0.71 N.B.
0"
4<3
Residual
0.51
4.9
0 .. 8
...,''''7 s with
1961
'1
1
1
16 .. 5
10.9
7.0
12.12
3.0
6 .. ;7
1965
22.65
17,,30
0
0
ssed
in
1 ..
follioles vii tb oapsules with
"
(a)
"
0
10
10
1
13
6
5
13
2
20
1
10
(3
1
~O
1
20
0
0
20
1961
1
1
19
- 58 ...
FIGURE
1
20
tf)
W
-.J
:.:>
(/)
0...1
«
U
I
3 10
If)
w
-.J
U
-.J
-.J
0
LL
0
0-
0
1965
1964
A
1963
GROUP
1962
1961
s of
crula.ted
va.luGs were convt;lrted
s
s (1948).
Analysis
)"
of fo111
.. 0
641tO.1
9;$60
***
<0 .. 001"
re
A
s was
the
mel'lt variance
Cubic
Wl;i.U3
of·
parti"hioned into
, the.
, "i.uadratio and
and tllE!
(
oOllt;ri buti O)'lS
1'0
total
the
f)
l
~
"
)
(
(
1
)
)
.,.
score
sion
II
of Y on
,.1 "" partia,l
.
1
"" ° Y1
b
'1
)('1
overall
the
- 10 ..
- 111
is
s in Y
ancl is o.er1 ved by the
",
:B'ur
...
b1
III
1<'
I?
*'
...",
y1
Ow
0 ..
*
< 0.01
(a)
on it;!
the
(
S in
ssion: ...
,... 63 -
standard
1
(b)
are shown
The re
the
)
**
77
was
to be
i9
on
(/)
W
...J
U
...J
...J
0
LL
/-0
1
2
M AN TIP SCORE
4
cr:tmp
'1:;0
a
in
follicles
on
with
equivalent of 10.
s·with
2
(
scores of
score
1
1
1
1964
1
1
2.07
2,
2 .. 3'+
2
is
(
)
FIGUR
3
3
w
o~
____
1965
~
____
1964
~
____
1963
AG
GROUP
AG
GROUP
3
w
cr:
0
()
2
If)
a..
-1--
1
Z
«
w
0
1965
~
____
1962
~
1961
very
the
9 ..
O.. 4·?7
105"
Total
**
'11/~,,97
,F'(O
)
scores of
mean
are shown
1
2 .. 30
1
2.19
1963
2 ..
19€J~
2.18
.,
2.32
tween
was no indication that a
by
a.nd
(
-
1
1
1
1
69 -
of
were
xx
in
1
1
1
4
13
,+
16 .. 7
4.3
15",
1
l~ .. 6
'16,7
1
4e2
17
os
stant
more tha.n one
)
s with more than one
.Most
)
liclel:3
Of
.
follicles with
he.va a
2,134·
1961
1
1
? ,.
,-
1
2,.
75
1
1
1
1
15
2LI,~6
104·
15
10
'1
,0
9
;;
2
11
4-
1961
",0
1
..
22 0
~)
I,':,~
- 71 (1)
commOl1,
1no
s 1
, :;> (
oocurrences
OOcurrence
(
)
(
se
)
is
YE'r'!'Y
the:!
between the '1
group when
be-
(iv)
(
(19
) ..
- 72 -
5
,
effects
s
a·difficult
IIllas
the Ronmey
of
that
s
•
Bhort (1
by
obvious
a
..
the
less
:ts
en in
it is
Short (1964a)
in the Merino
this was
to
to
of
could,
the
has
, 1
the
was
crimp
the
breed
was
" 1(60)"
and a
and
is
)"
eo:m:nwn
Because
sent;
11 be
of
e:(.).ces
of
under cornmon
it is
in
coat
of'
til
length
.form.
s of'
with
by
a
ems
i;;o
ee.ch
these
causes~
score
this
score
V:13.S
score
tihe se c V.....,'' 'IC''.....
1'011:i.016
sJ.milar
is
sl10uld be
by
on
acora was
'1
, 1
1
110
seem that one
re
c
to
OIl.
wi
able to aost:une
a
ss
'.Nle
1101as
s stndY'
ttve.
to
in a
:t ash:t on from
a
)
on
"
- '16 -
s.
<A
>" ..
some
is
is
,
~l
as
as
"
[;:1
II
a
s
s.
1$ no
c~Qoe
is
1.s
scorc"
6coriil
to
)
(1
)
was
This does
ex-
s
(1965)
..
the
aa
but little
"
s
assume that
asas with
in
cause
fomn
follicle
sa that ten,a
s
in
seem to
the
s
SOine
control
an
to
follicles
level ..
on
, occurs.
- 79 l11ay
is
seerns
doubt
is an
•
on
L1UlI1,hel's of
deteri
crimp
thfl.'\J the
is
the
re
a
s in the
of
a
No
a
f'ollicles
other
on.
produce more
s
in
waf~
that <both
production
in
of arrangement of
The
fibres
OI."OUllCE;tQ.
by the follicles
(1957),
IS
s.
(1961)
Gl~ynnt
in
is
(1
to
1
is
:ffi.easnre
f
s
- B-1
on in
sy
(195?) was
of
"
(1
t
cells
(1
8
(
(
) which
more
aula
(.me
(1956)
)
(1960)
s
!i
some
attf:~ched
the
bulb
bulb to very
from
they were
in
(1
bulb ..
cle
bulb~"
8,S
, thus
on of the
s/]? ratios me
in this
are lower
the Homneyby
S
til
8/1'
(tf
(1
)"
5 •.4 with
>J
t ile
" mean f·'/p··
) quol;e /"J a B/p
on of' 16.0
'Romney
are three
s in
1 ..
son (1
breed."
) were
(1
)
found
of an
'II
.follicle
lmt
time
tb,E)
(1
2 ..
)
Aug LU3t
cycle,
s could
count
no
to
Origin
if!
(1
orIgin
lower
s aren of
is
no
on
on
:t s no :reaso:o.
ass
:i.e no
on
is
.; c!.
kf.,.!~
85 is
c
s
this
on :ts
on
on
was not
on
re
s al:'a l1.ece
f21ellCe O:f'
ly'
to
i
a
on
upon
(;3)
of
studies
on
be
occurs,
6
1"
A
Homney
of a
in
Southland, was
a
(
ace
v/1th
grown
type
on a 1 - 4
of
5.
was
..
Crimp soore was
.
score showed no
An
was found.
1':1ssoeiatio!l
was
scora~
sules
with equal frequency on
20 per cent of
:more
no
in. the
were
omn.ev~
one
..
The
, "fJroi'esaor
A.E.
illg thl':l course
technical
College,
BUt
dur-
S f3tU.iy ..
Seience
in prepElration
...."""""""''1'', Ili:acoll1
measuremellt
the
l::m i; 11 or
, Bouthlanda
v1l0uld like
st~"lCe
of
is gratefully
-
, VI .H.
';:IV
-
1
It
J ..
• "IO:11EL
"
.
.
J?roc ..
• B"
• VII.
1950 ..
4th,
, A.D .. ,
1
.
,
,
,
r1' .1)"
'>
A.
~
1
..
1
.
.
~r
" Anat.
..
..
.
H..
1
~
.
"
t
• f? ..
s ..
.
Anim ..
• H.
1
1947Q
C
;r"
.
"
8:
.
"
1950
164 ..
~
:64 ..
"
1
Hdh
..
:207 ..
•
,
,
., J.
1960"
J ..
..
1
"tJ.
:1(Yl.
.
.
• J ..
Comm.,
11.13 •
•
1
s"
"
"
J ..
B.
'!l
"
.
- 91 -
,
1
Et.Jj;.
1
J
~
f... ust
.
It
c ..
.. 11 ~
•
1
.
:1
II
, lL.U'.
..
1
19640(1
" 1" •
(.
1
Aust .. J.
1
211"
1
1
11
K,,~J..
1
1
"
.
II
1
..
Oh.. IV iii 2n.d.
iii
(
Go" ,
6:
• R.O.
B:64:
1
"
1952 ..
.
11
"
1
..
"
1
..
.
1 ..
.. 131
:B1l11iarcL~
G·lynn
~T"
Lan5i5, VI. It ..
s .. 11:
1
11:"
"
..
..
1
Y"E"
.. J",
Vet ..
die
M.H. and IJyne ., A.G.
Con! .. Aust .. F:26 ..
1955.
Hardy,
.
Froc.
..
1
,
"
1
i>
.
. N.Z"
1967 ..
1
..
1
..
.. ,
It
fI
: rl
•
If
1
..
.,
.
Soc •
..
, rl!"
1
.
.
s .. ,J ..
.. Jv
11
~1 amEllS ~
1
iJ '" Il' •
1A: 12 ~
s, G"
12:
"
1961,.
.
t
A"
.
1850"
1· Ot::.Q
/./U"
Wool
1
..
"
~J
.
..
•
• 10,
.J ..
• 51:
1959 ..
end.
" .1,. Bi
21 :
~T
..
..
..
:1
11
.. It •
.. of
"
p .. 20'7 ..
l~ ...
s.,
"
Ow
1
..
Jill
1
1
,
Aust" (Joun.
..
Text ..
"
s~
J. 24:
'" J
..
.
..
1
"
1946.
~ CI!231 "
'"
, e.o.
,
t
I( •
It"
1
•
.
J,.
19:407 ~
.
1
l/>
J~
.J~J.
.
J'1lI
"
"Notes for
N.Z"D~A"
Handel,
.
.
1
1
,
1
p. 1,11
1
II
<Ii
100:1 '"
.
,8.K~
N.Y~
.G.
'1
1
• f"
If
.
'1
if
.
'1
J3.Jr.
l~_
I!
Arb ..
.
1
.. t
II>
. Zool"
.. .I.H.
.L.
2 LH401B ..
'1
'1
~20'7.
.
.. s .
.
..
Sydney
(1964) ..
1
'" 2
.. 11 :19 ..
,
t
B.B..
'1
1
.B.
: 51? ..
Ann.
'"
J ..
"
k...L.J.'~ jif J.,\.A.:'~Q.,.L.
~.iL.z.vWJ':!
""",v~.J",.,.,Ji,,,'V~<;Oi
".A.Q~c;J.
ll:,lVI
-
IN"IOIO_
1S
13
2
11
1
15
2
13
6
13
, ___
"'.
""'0."" ""... """'"w '"
1
-
r
0
t.
1+
-
2
1
7
~
i-:;t
ry
9...-"
1
1
c::.
4-
I
1
6
2
1
'1
1
2
4
12
16
'1
161
uc'
8
5
10
T~
2
1
c::.
10
'1
-.
?
17 .. 0
13,,3
....
?"7 .. c:.
~
0.,5
1
o.
~
111,:/
15.
.
- 1
e
4-
1
<5
?
123
'1
13
2
0,
11
16
-1
5
1
1
1
,..,
~1
4-
c:
0
1
23
9
10
7
1
1()
!"\
'j
1
"'I
9
2
2
2
13
1 .. 3
2 .. 1
0
..... _"'"""''1'1'''...._ _ ' ' '
(2) -
s.
'1
7
5 -'7,'' \
9
'1
a,,;
'1
)
2
'1
17
3
1
10
1
10
2
4-
2)
2
/
\..
8
"
,~
., i'}
10~9
.,
...
-,..
;
?,
",-
\,...j
1 0./
1
6
2
1
1 .. 3
2
4"
?
::z,
'0
6
1
vQ
6
1
5
L~
6
6
A
...-
3
2
?,-
,;:;,
'V
r
1...-
-''''3
6 .. 9
1
'1
-
6
6
c;.u,,,,;:;
4
1
?
1:':'
16
1
1
12 .. 5
.",
1
1
'1
.r,:.
,-,,u 9
1
3
1 .. 5
15
0 ..
f'
.u
1
7 .. 0
(
R'l
•
n
'"'"')'"
412,
-'1
.:::
1
'1
?
1
r,
v
.."
:)
'1
10",(
2
;"'1
...
~ ,\
'1
'1
"1
"
~
2
1"1
l
2
,,6
,,8
2
5~7
1
L~
53
~
,. ,...,
'1
2
l'
~
·5
Lt
"
11
'1
1
(>
2
:3
l
- 1
\,
r,~
t:illi""l> .. wmr
~~
_~~,:1lb"';"
2
5
145
1
'1
8
~
1
'1
-
1'-1;<"\
1
...
_ _ _ _ _ _ _ _ _ _ _ _ _ _'_W
..
'~oIIi3
1
_""'"'I......
::tm*'i'mrO\ll.
I>l
"i~
III.
f
Bcores
-1
.,
:;
2
2
1
2
2
.2
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In.cidenoe of
scores - 1961
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(Oont'i).
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of eapaulea
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- 61
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- 61
- 61
-
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?
- 119 -
Appendix VI (Cont'd).
Sheep No.
Type - Numbers of Follioles of Eaoh
1
467
446
237
15
248
122
194
350
136
205
244-
408
124
69
ortho-pe.re.-cor'tex - pattern of
arrangement. 1965 ewes - all follicles normal.
2
18
13
11
17
17
2
19
18
19
8
2
10
4
5
6
6
23
6
56
8
41
13
62
22
12
3
23
22
16
21
13
1
17
14
23
24
3
1
-
8
5
3
5
7
36
23
47
4
18
6
42
2
23
32
1?
33
14
95
2
29
65
33
8
28
2
10
36
?
2
10
3
1
29
41
13
34
36
21
Fibre J/JissinEi
1
15
8
16
5
2
1
4
5
2