KEEP THIS IN MIND • Microclimates determine local conditions A CLASSIFICATION Body Temp • Poikilotherms and Homeotherms 5 10 15 20 Environment Temperature ENVIRONMENTAL IMPACTS • Gains Losses SUN RESPIRATION WIND AIR GROUND GROUND REMEMBER • Biochemical reaction rates are temperature dependent A LAW Reaction Rates • Van’t Hoff’s law: Reaction rates double every 10 C Temperature A THOUGHT • Enzyme reaction rates can affect many processes including development, physiology, behavior, etc. BARK BEETLE RESIDENCE TIME TEMP. Male Female 15 20 25 30 32 35 85 32 12 9 8 8 82 32 11 9 8 9 ATTACK RATE TEMPERATURE EFFECTS ON PARASITIC WASPS TEMPERATURE TEMPERATURE EFFECTS 4 3.5 3 2.5 2 1.5 1 0.5 0 0 10 20 T em p er a t u r e 30 40 DEVELOPMENTAL TIME STORED PRODUCTS BEETLES TEMPERATURE TEMPERATURE AND DEVELOPMENT RATE HOW DO POIKILOTHERMS DEAL WITH EXTREMES? • COLD: • Super cooling is cooling of a liquid below its freezing point without it becoming solid. • Freezing point depression is cooling below freezing point of a pure liquid due to a solute e.g. NaCl Ex: PAINTED TURTLES • Hatchling painted turtles typically remain inside their shallow (8–14 cm) nest throughout their first winter where temperatures often drop to -10C • Their skin resists ice nucleus penetration Ex: GALL MOTHS • Freezing point depression through synthesis of antifreeze (glycerols) HEAT SHOCK PROTEINS ANOTHER OPTION • Maybe you should just leave IMPACT OF GLOBAL WARMING Bergant et al. 2005 The future? “In a future with increased mean temperatures … heat waves would become more intense, longer las:ng, and more frequent .” WHY DOES GLOBAL WARMING MATTER? • Both mean and variance can both affect ecological processes A CONCEPT • For poikilotherms, time is temperature dependent but for homeotherms it is not A QUESTION • Q: How do insectivorous birds determine when the best time to breed? • A: They use calendars (photoperiod) A PROBLEM • For insects, timing depends upon time and temperature AN IMPACT • Mismatch timing can be important Year! Year! A BIGGER PROBLEM • For long-distance migrants there is no way to anticipate changes in timing in a given year Swamp Forest WHAT ABOUT HEAT WAVES? • They will cause continuous stress • Many animals and plants cannot escape these stresses MICRO HABITATS CAN VARY DRAMATICALLY • Position on a rock can be critical for a mussel on a rock