106 01\' Q..,IE bNITED STATES AND CANADA Society of American Foresters F, H. Fyre, ditor 1980 Nom-H PAan;E'OREST COVER 'l'YPES WESTERN Foru:sr Covrn 'IYI'n "$ -! I::; ·0 :E Vl -· . : Ig Pacific Douglas-Fir .ro':l ·,-., Vl 229 Definition and composition. -Pacific Douglas-fir is one of the most well known, extensive and, in vol­ ume of wood, highly productive forest types in the world. Stands that comprise the type contain at least 80 percent Douglas-fir. Other species that may occur in light mixture include western hemlock, grand fir, Pacific silver fir, white fir, noble fir, Cal­ ifornia red fir, western redcedar, bigleaf maple, Pa­ cific madrone, red alder, tanoak, Oregon white oak, giant chinkapin, vine maple, redwood, Port Orford-cedar, ponderosa pine, we5tern white pine, incense-cedar, sugar pine, and Sitka spruce. Geographic distribution. -The type is present in southern British Columbia, but it is widespread west of the summit of the Cascade Range in Wash­ ington and Oregon south to approximately 43°. north latitude. Farther south, a drier, less extensive portion of the type becomes increasingly intermixed with other types. In discontinuous stands, it contin­ ues south throuJ;!h the Klamath Mountains of Ore­ gon and northern California through the Coast Range to the vicinity of Mendocino County, Cali. fornia. Although pure stands o1Douglas-fir occur in the southern Cascades and the Sierra Nevada, they are small, disjunct, and probably should be consid­ ered local phases of other types containing this species. Elevation of the type is generally from sea level to about 500 m (1,500 ft.} in the northern part of its range and to about 2,000 m (6,000 ft.) in t_he south. ern. Annual precipitation ranges from about 600 to 3,000 mm (23 to 120 in.) (Franklin and Dyrness 1973). Ecological relationships. -As its wide latitudinal and elevational_ ranges show, the type is capable of a apting to diverse site conditions. Where the type is excluded, the reason may be excessive heat (Sileo 1960, Hallin 1968), drought (Isaac 1938, Tappeiner 1966), cold (Cleary and Waring 1969), too much ; moisture, or high water tables (Minore 1968). The : type generally occurs on moderately deep and welldrained soils derived from highly variable parent t materials (Franklin and Dyrness 1973). , Natural Douglas-fir stands typically regenerate after wildfire in partial.shade· cast by fire-killed trees. Seed is provided by scattered surviving trees : or islands of trees. The young stands that result ' often are so dense as to crowd out competing vege- i tation. As stands develop to pole size or larger, mor­ tality from snow or 1ce breakage, small fires, root pathogens, bark beetles, windthrow,· and suppres­ sion periodically relieves overcrowding. Mature stands can remain healthy for decades. Finally, when mortality h caused openings in the canopy, the climax species, typically western hemlock, west­ ern redcedar, and Pacific silver fir, become estab­ lished. Eventually they take-over the site unless wildfires renew the cycle. · · I ro :E In the absence of fire or other ·catastrophes, the type can be maintained only through the actions of man in creating mineral seedbeds necessary for 3 Vi' a.. ro seedling establishment and in opening dense cano. ::t> a.. oor g pies to permit development of a new stand (Isaac 1943, Williamson 1973). W here canopies exceed Vl ?5 § ;! about 70 percent, the type is replaced by others con· Vi' taiping more shade tolerant species. ;:::1:' -< Combining a moderate climate and ample pre O"Q ;g/ d long-live a and soils fertile deep, with on cipitati 5ru :l species creates an extremely productive forest. Stem-wood volume alone commonly exceeds 5· ri5 ' 1,3003/ha (about 150,000 bd. ft., Scribner rule) in ;;. q · ' . 5· g [ -o 1 8 £/ . ! :::; Vi' ::::: n' ', ,:::· j g.i · old-growth stands. Fire is a destructive agent in Do glas-fir, but it prepares a seedbed for regeneration. However, other agents, such as the Douglas-fir bark beetle (Keen 1952) and the black bear (Poelker and Harty.rell 1973). cause mnch stand damaue vet leave the torest floor undisturbed and do not provide the nee­ . essary mineral soil seedbed. Instead, they preserve a ing surface better suited to competing conifers and woody shrubs. U large open areas are created and no Douglas-fir seed source is present, stands of red alder and bigleaf maple may capture the area (Newton et al. 1968) Variants and associated vegetation.- Douglas-fir occasionally forms uneven-aged but essentially pure -stands and apparently exists as a climax in drier areas occupied by the type. This climax situation can be caused by topographic rain shadows, soil, and climate characteristics that lead to droughty, but not xeric, soil conditions. Such conditions occur from the east sides of Vancouver Island (JCrajina 1959) southward through the east sides of the Olym­ pic Mountains and Coast Ranges, and through the intermediate mountain slopes of southwestern Ore­ gon. to the southern limits of the type range (Dau­ bertmire 1978). Sometimes on these sites Douglas-fir \s mixed with hardwoods, such as Pacific madrone, . Oregon white oak or tanoak, and with conifers, principally ponderosa pine. This mixing leads to a merging with Douglas-fir-tanoak- Pacific rna­ drone and Pacific ponderosa pine- Douglas-fir types The . former type may replace Pacific Douglas-fir after wildfire or disturbance by man. Douglas-fir climax stands often are associated with oceanspray and salal. Grand fir, western hem­ lock, vine maple, Oregongrape, and swordfem are common associates in more mesic, seral Douglas-fir stands (Franklin and Dyrness 1973). . RicHARD L. WILLIAMSON USDA Forest Service Pacific Northwest Forest and Range Experiment.Station