c ,:~:\i;:~~:~i,::, ~;'~i':J~~~r;{,'~:,,;,,NOTES FOR THE GAME FARMING 'fi§i"'~'\)~\r~l:j; ""'Co-,-:,;, INDUSTRY ~ 1991 J.t:. Haigh MRt:VS University of Saskatchewan ~. l ' ~~~~:;'i"~,[.. ."'. 'o'f' '.,;.~.!.. Moose (;~'":::'.;; '.;,. "".""',r,,"~ , ;.~" Introduction '~~('~~~:i:.. Moose (Alces alces ) are common to the boreal forests of the Soviet Union, Scandinavia, and North America. In North America the species tends to follow the general rule promulgated by Bergman that the further north a species is found, the larger it becomes. There are four subspeciesof the genus Alces recognised in North America. The largest is A.a. gigas which is found in Alaska, the western Yukon and perhaps the very northern tip of British Columbia. The range of gigas merges with that of its smaller relative A.a. andersoni which is found throughout most of western Canada in the boreal forest areas,the Northwest Territories and isolated pockets such as Moose Mountain Provincial Park in Saskatchewan. It ranges as far east as Northwestern Ontario, and occupies appropriate habitat in states to the south such as Washington, Idaho and Minnesota. In most of Ontario, Quebec, and the maritimes A.a. americana is found. The smallest sub-speciesis the shiras moose, A.a. shirasi which is found in the very southeasterncorner of BC and ranges further south than any of the others, being found in southern Utah. Even its habitat may be quite different than that normally associated with the forests and marshy grounds of the north. Much of the range in Utah is mountainous, but is covered in dry oak scrub for a period of the year. from 226-244 days, and 240-246 days is"q'uoted for Canadianmoose. A pair of yearling females that had no possible accessto a male, and were <;loselymonitored by Bob & Lyn Stewart, calved 216 and 218 days after being penned with a bull. This wide range is interesting, but there is no easy explanation for the differences. In the wild yearling female moose do not usually breed, unless they are very well grown and have abundantforage available. There do not appearto be any formal studies showing the critical weight at which females will come into estrus, but in many other deer species it is about 65-70% of mature weight. Domestication In 1865 Galton listed 6 requirements for successful domestication of species. They are that the animals:- should be hardy; should have an inborn liking for man; should be comfortloving; should be found useful...; should breed freely; and should be easy to tend. Moose qualify on all but the fIrst, but there is a modern interpretation that makes them less suitable than Weights of adult moose fluctuate throughout the year, as in other cervidae. The maximum weights may exceed 700 kg (1540 lbs) in Alaska whereas a bull moose weighing 550 kg (1210 lbs) has been recorded in mid-winter in Saskatchewan. The latter is likely to be at least 10% less than the maximum prior to the rut. Gestationis generallyquotedasbeing Publicationsupportedby the Canada/Saskatchewan ERDA agreement GF-M-I-OI J'""" either wapiti of fallow deer. This is the requirement that animals suitable for domestication form herds and have a dominance heirachy that makes them easy to handle. Moose are an example of a species which has failed to become widely established as a farm animal. It does appear that they can be readily domesticated and historical records in both Russia and Canada that indicate that they have been used as draft animals for at least 300 years. There have been moose farms in Russia for many years. There is a description from 1861 telling of the use of moose for drawing com from the fields, and the riding of moose was once common practice, until it was banned becausepolice on horses could not catch up with villains on moose! The first experimental farms were started in Russia after the 2nd world war, and by 1969 there had been a 4th generation moose that was 18 years old. The most successful managementsystems involved no fences, and the moose were allowed to range freely in search of food, returning in winter for supplemental feed. Bottle raised moose, captured at less than 3 days of age, are easy to tame, and may be milked with ease when they calve. Wild-caught adult moose do not adapt readily to captivity, and often die of stress related diseases. Perhapsbecausecalves were often found and were quite tamable, moose have always attracted attention. Samuel Hearne, a Hudson Bay Company employee in the late 1700s, considered moose to be: the easiestto tameand domesticateof the deerkind. I have repeatedlyseen themat Churchill as tameas sheep,and evenmore so;for theywouldfollow their keeperanydistancefrom homeand at his call return with him. Table 1 Ingredient GF-M-I-O2 Aspen* concentrate Corn, Ground yellow Wheat Sunflower hulls -whole Sawdust -"Pibrelite" Oats, ground Oats, whole 30.0 Barley, ground Beet pulp Cane molasses, dry Can molasses, wet Soybean meal (7.4% N) 12.5 7.5 "Pelaid" 1.3 25.0 15.0 6.3 Pellet binder Dicalcium phosphate Sunflower# concentrate 28.2 26.6 17.2 5.6 5.6 5.6 7.2 0.8 1.4 Biophos NaCI 0.5 Salt (cobalt iodised) Vitamin A,D,E 0.3 "Mycoban" Fig 2. Youngmoose calvesrea4ily adaptto handrearing. Hand-raised bull mooseare dangerous and unpredictable during the time that their antlers are hard, and must be handled wih great care (fromref# 4) Compositionand analysisof two pelletedmooserations Trace minerals & flavour trace trace Analysis Dry matter Crudeprotein Cell wall constituents Acid detergentfibre Lignin Ash Grossenergy(Kcal/g) Calcium Phoshorus Magnesium Zinc (ppm) Copper(ppm) Manganese(ppm) Selenium(ppm) 84.9 12.7 57.0 24.0 5.2 1.1 4.26 1.1 0.58 0.17 114.9 13.5 106.8 0.2 0.8 1.0 * Schwartz et al1985 # Stewart pefS comm) Figs 3 & 4 On the left is a print takenfrom a photo publishedin a newspaper(possiblythe WinnipegFreePress)takenin 1905of a mooseteamin harness.In racesagainsthorsessuchteamswere seldomif everbeaten. On the right, a morerecentPolishpicture of a moosepulling a sleigh. Archives contain many photographs of moose used for riding or pulling sleighs but they are usually yearling and subadult animals. (see figures on page 3) Moose continue to be difficult to maintain for long periods of time and captive populations are seldom self-sustaining. Moose may be behaviourally suited to life as domestic animals but are susceptible to disease, have exacting requirements for feed, and generally require a great deal of space. Studies of captive moose in North America have occurred at Ministic, near Edmonton, and on the Kenai peninsula in Alaska. At Ministic, production was the main focus of attention, while at the Kenai, other subjects, . particularly as they relate to the biology of freeranging moose, was the objective. Nonetheless, the scientists at the Kenai have developed a ration which has improved the longevity of moose in captivity and overcome some of the digestive problems seen. As moose, the largest of the cervidae, are concentrate selectors, seldom grazing unless forced to, they have ~ highly specialized dietary requirement that involves the ingestion of relatively high proportions of fibre. Rations involving either aspenconcentrate, or sunflower hulls, have beenused with some success.(see table 1) However moose are more of a solitary than a herd animal and their high productivity is balanced by high mortality rates. Although they can evenbe hand-milkedefforts in bothRussia and to been develop large scalemooseoperationsCanada havenot successful. . Handling There are few records of moose being handled in chute systems, but it is certainly possible with animals that are used to captivity. Minor modifications to handling systemsdeveloped for wapiti would probably work well. Trained animals can be held in minimal chutes, for simple procedures such as blood sample collection. Immobilizing drug doses are similar to those for wapiti. Problems When kept inside fences on game fanns moose will always try to browse in preference to any other style of feeding. This will eventually cause irreversible, damage to trees. Only if sufficient browse is made available to them year round will moose do well without special diets. Manage~ent staff at the Quebec City Zoo have been able to keep moose on display for long periods by providing browse cut by a contractor clearing under power lines on a year round basis. The browse is stored in a cooler and kept just above freezing for year-round use. Moose are susceptible to a variety of disease conditions. Probably the most important of the viral conditions to which they succumb is malignant catarrhal fever (GF-S-6). They GF-M-I-O3 ~ usually die within about a week of contracting the disease,and there is no treatment or vaccine. There are several parasitic conditions to which they are susceptible. Ticks, especially the winter tick Dermacentor albipictus may cause extensive or even virtually complete hair loss, and this parasite has often been incriminated in moose die-offs, especially in late winter when the coat is so important for insulation from the cold. (GF-S-8). (see fig 5) FigS Moderatetick infestationon a mooseatMinistic. In severecasesalmostall of the guardhairs may bebrokenoff. In suchcasesthe animalappearswhite, which is probablythe origin of the "ghostmoose" mythuncoveredby Dr. Bill Samuelandhis teamat the University of Alberta. Conclusion The brain wonn, or meningeal wonn Parelaphostrongylus tenuis is also an important parasite of moose, causing so-called "moose sickness" or circling disease.(GF-S-2). There are a number of gastrointestinal parasites to which these animals are susceptible, and they may develop diarrhoea that is difficult to control. In fact chronic scours, due to improper diet, worms, or coccidiosis, may be the most common problem seen in captive moose. Other than the important abomasal parasitesof the family Ostertagiinae the whip-worm, Trichuris spp., which parasitizes the caecum, may causeproblems. Personal experience has shown that ivermectin is not always reliable for treating moose infected with this worm. An alternative is the use of mebendazole in the feed for 3 days at a dose of 10 mg/kg. Mebendazole is available in Canada as Telmin, which either comes in powder or as a paste in a syringe. It can be administered either with the syringe, or inside a banana, as moose that have been accustomed to this fruit will eat it eagerly. A note of caution however, there is a preparation of Telmin for horses called Telmin B. This contains an organophosphateas well as the mebendazole. The one experience that I have had with it was almost a disaster, and the moose showed signs of poisoning for several days. Moose may be possible to keep in small numbers behind fences, as long as they are properly fed, and have access to some form of browse or high fibre diet. Unless some new information on management comes to light they are unlikely to prove to be a commercially viable livestock species for the production of venison, as the costs of maintenance, and the problems associated with management, are likely to preclude high stocking rates. Bibliography GaltonF (1865) The first stepstowardsthe domestication of animals. Trans.Ethnolog.Soc.Lond. N.S. 3: 122-138.Reprintedin Inquiries into HumanFaculty. MacMillan & Co. New York. Haigh,J.C. (1991). Managementrequirementsfor farmed deer. In Biology of deer. Brown,R.D. (ed) Springer Verlag. New York. In press. Schwartz,C.C., Regelin,W.L. and A.W. Franzmann. 1985. Suitabilityof a formulatedration for moose.J. Wildl. Manage.49: 131-141. Syroechkovsky,E.E., Rogacheva,E.V. and L.A. Renecker 1989. Moosehusbandry. In Wildlife Production Systems.(Hudson,R.J., Drew, K.R. and L.M. Baskin (cds). CambridgeUniversityPress.369-386. Photocredits:-Fig 1,2,5.J. Haigh. Fig 3. ?Winnipegfree press1905. Fig 4. A. Krzywinski GF-M-I-O4 @