United States Department of Agriculture, Forest Service Pacific Southwest Research Station Sierra Nevada Research Center Medieval Forests, Volcanism, and Response to Climate: Clues to ecological effects of future climate changes The Research: Whitewing Mountain, 3030 m, is a broad peak extending above current treeline that lies east of the main Sierra Nevada crest and adjacent to the Inyo Craters volcanic chain. The last eruptive sequence of Glass Creek Vent occurred ~500-800 years ago and covered the flanks and summit of Whitewing Mountain with more than 4 m of tephra. Strewn on top of the tephra over the otherwise barren summit plateau are hundreds of well-preserved dead tree stems, many of which are large, straight, and exhibit evidence of rapid growth from their tree-rings. The surprising growth rate, form, and occurrence of trees in this ‘ghost forest’ at this elevation suggest that they grew under climates very different from the present. Regular orientation of the tree stems and absence of stump bases suggest that the trees died in the Glass Creek Vent eruption. A similar situation exists on adjacent San Joaquin Mountain. We are using these natural events to unravel the story of forest origin, the climate under which the forests grew, and the exact year of the volcanic event. Objectives: • Identify species and determine dates of trees that grew in the ghost forest on the summit of Whitewing Mountain. • Accurately date the last eruption of Glass Creek vent. • Model the climatic conditions under which the forest grew and interpret the significance and implications for forest response to future climate change. Our Mission: Sierra Nevada Ecosystems are complex and our knowledge of them is incomplete. As a result, the long term outcome of any given land and resource management strategy is uncertain. We will provide assistance to land managers and policy makers by addressing this management dilemma through targeted research, emphasizing an integrated, ecoregional approach to examine particular physical, ecological, and socioeconomic issues, across a range of appropriate spatial and temporal scales specific to each issue. This unit will represent the collective research expertise and interests of scientists located in Fresno, Davis and Albany as well as other scientists within the Pacific Southwest Research Station. With a full spectrum of research, from long term, fundamental research to short-term, tactical applications, this Center is intended to support conservation, restoration, and sustainable utilization of the lands within the Sierra Nevada ecoregion. Application of Research Results: Using ecological niche theory and contemporary distributions of the species, we modeled paleoclimate during the Medieval period of the ghost forest to be significantly warmer (+3.2 degrees C annual minimum temperature) and slightly drier (-24 mm annual precipitation) than present. These values resemble projections for California in the next 70-100 years under conditions of global warming, and give insight into the potential responses of forests in subalpine zones. Based on conditions at Whitewing Mountain, forest managers and conservationists may expect much larger movement of tree species in response to climate change than has been anticipated to date. Non-equilibrium changes, with species moving individualistically, rather than in equilibrium with their current ranges, may be expected. Regional extirpations, such as happened to sugar pine in the eastern Sierra following climatic change at the end of the Medieval period, should also be a potential outcome that managers evaluate as they plan climate-informed policy. Location: Whitewing Mountain, 10,012’ (3030 m), eastern Sierra Nevada, between Mammoth Lakes, and June Lake, CA, Mono County, and similar environmental locations nearby (San Joaquin Mountain). Publications: PDFs available at: http://www.fs.fed.us/psw/programs/snrc/staff/millar • Millar, C.I., J.C. King, R.D. Westfall, H.A. Alden, and D.L. Delany. 2006. Late Holocene forest dynamics, volcanism, and climate change at Whitewing Mountain and San Joaquin Ridge, Mono County, Sierra Nevada, CA, USA. Quaternary Research 66 (2006): 273-287. • Poster: Millar, CI; Westfall, RD; Delany, DL; King, JC; and Alden, HA. 2004. HighElevation Response of Conifers to Climate Change in the Sierra Nevada and Western Great Basin, USA: Treeline Elevation is Not the Primary Effect. • Poster: Millar, CI; Westfall, RD; Delany, DL; King, JC; and Alden, HA. 2004. Climate as an Ecosystem Architect; Responses of High-Elevation Conifers to Past Climate Variability. http://www.fs.fed.us/psw/programs/snrc USDA Forest Service Pacific Southwest Research Station Sierra Nevada Research Center Albany Location: 800 Buchanan Street Albany, CA 94710 For further information contact: Dr. Connie Millar cmillar@fs.fed.us Mailing Address: P.O. Box 245 Berkeley, CA 94701 Dr. Peter Stine, Program Manager Phone: 510-559-6300 Fax: 510-559-6440 Davis Location: 2121 Second Street Suite A101 Davis, CA 95616 Phone: 530-759-1700 Fax: 530-747-0241 Fresno Location: 2081 E. Sierra Avenue Fresno, CA 93710 Phone: 559-323-3200 Fax: 559-297-3355