----------- " ; ,. .' ICES 1995 1 C.M. 1995/J:18 Baltic Fish Committee POLISII HYDROACOUSTIC ASSESSMENT SURVEY 01' IIERRING. SPRAT AND COD STOCKS IN ICES SUBDIVISIONS 25 AND 26 01' TIIE BALTIC CONDUCTED IN OCTOBER 1994 by Ryszard Grzebielec. Andrzej Paciorkowski Miroslaw Wyszynski. Wlodzimierz Grygiel Sea Fisheries Institute Department of ßiology and Conservation of Fish Resources ul. Koll~taja 1. 81-332 Gdynia. Poland . Abstract Within an international program of the ßaltic clupeoid fish assessment coordinated by the ICES Planning Group. the Polistl research vessel "Baitica" carried out a hydroacousticsurvey. devoted to llerring and sprat in the ICES Subdivisions 25 and 26. The survey covered the area of 14774 NH 2 . Echosignals were integrated over 1204 one mile distances. Twenty nine control hauls with pelagic herring trawl were conducted. 9263 specimens of herring. 4612 of sprat and 73 of cod were measured. Otoliths from 1459 herring. 709 sprat and 73 cod for ageing were co 11 ected. The estimated biomass density expressed in ton ries per 1 NH 2 eQualled 30.1 tonnes for herring. 44.2 tonnes for sprat and 3.7 tonnes for cod. The results compared to data from October 1990 survey conducted from on board r.v. "Profesor Siedlecki". (Orlowski 1993) indicate a slight increase of the herring biomass density (by 1.9 tonnes). a significant increase of the sprat biomass density (by 26.7 tonnes) and also a large increase of' the cod biomass density (by 2.3 tonnes). Distribution of the herring and sprat biomass has also exhibited some changes. Herring was less numerous in the Slupsk Furrow but was present in greater numbers in the Bornholm Decp. October sprat aggregations shifted northeastward. Cod cauBht with pelagic trawl appeared only in trace Quantities in thc Gdansk Deep area. The estimated herring and sprat biomass was recalculated into numbers at age using age-length key and mean weights for each ICES rectangle. Introduction During 3 - 20 of October 1994 r.v. "Baitica" carried out its maiden hydroacoustic cruise. The survey was apart of an international program of the Baltic clupeoid fish assessment prepared by the ICES Planning Group for Hydroacoustic Surveys in the Baltic (Anon. 1994). The main task of the cruise was assessment of herring and sprat stocks in terms of biomass and numbers in the ICES Subdivisions 25 and' 26. The r.v. "BaItica" survey continued the series of r.v. "Profesor Siedlecki" hydroacoustic surveys conducted during 1982-1990. Equipment and survey method Echosounding and echo integration were conducted during day and night continuously. using hydroacoustic system consisted of echosounder EK400. echo integrator QD and a PC. Echointegrator values referring to time periods equivalent to 1 NM distances covered by the vessel along the acoustic transect were recorded both as echodeflection and volume backscattering strength by recommended depth layers. The data together with mean depth of integrated layers were transferred from QD to PC and recorded on the hard disco The vessel speed during echointegration was maintained within 8-10 knots depending on the sea state and wind speed. The hydroacoustic system was calibrated using a standard copper sphere. The calibration was carried out next to the Swedish Island of Högön in the vicinity of Västervik harbour with assistance of dr Hakansson from Institute of Marine Research in Lysekil. Control fishing Control hauls were conducted only during daytime using midwater herring trawl WP 53/64 provided with 11 mm mesh bar length in the codend. Vertical opening of the trawl equalled 17-36 m (mean 26 m). Trawling speed was maintained at 3.5 knots. Standard trawling time of 30 minutes was followed. Two hauls per unit of survey area (ICES rectangle) was planned as a minimum required sampling density. Area stratification and estimation of fish abundance in number In accordance with the ICES Planning Group rccommendations all survey results were recorded by ICES statistical rectangles. The parallel transect design was used. with equidistant spacing of approximately 15 NM. similar to that followed during the r.v. 2 "Profesor Siedleeki" eruises sinee 1952. Fish abundanee in numbers was ealculated based on a total biomass estimate and me an individual weight of fish. The total biomass was estimated using Orlowski's method (1993). Target strength of 1 kg of herring and sprat was calculated applying the following formula: where: - mean length in a reetangle (ern) w - mean weight in a reetangle (grams) The results of the caleulations has been collated in the standard format reeommended by the ICES Planning Group (Anon. 1994) and forwarded to the Group Chairman. • • Results The survey track along which the echo was integrated and positions of the control hauls are presented in Fig. 1. The area of reetangles included in the echointegration amounted to 14774 NM 2 . Length of the vessel traek eovered with eehointegration totalled 1365 NM but only 1204 NM were accepted for final analysis as elementary sampling distance units (ESDU). In total 29 hauls were carried out of which 3 were rejected as not representative (Table 1). The index of rectangles coverage with control hauls equalled 1.7 (less than planned index of 2). The index of area coverage with representative acoustic transects amounted to 12.3 NH 2 per 1 ESDU (in 1990 9.26 NM 2 ). The greatest biomass of 653.522 tonnes constituted sprat (Tab. 2. Fig. 2 and 3 - created using SURFER Golden Software Inc.), one third of which was concentrated in reetangles 4064 and 4065. Herring biomass was estimated at 444.284 tonnes. Near half of its biomass was eonfined to 3 rectangles in the northern part of the survey area. Cod biomass was estimated at 54.579 tonnes; almost all of it was found to be distributed in the Bornholm Deep. The total length 6f sprat caught (Fig.4) varied within the range from 6.5 cm to 16.5 cm. The Subdivision 25 apart from rectangles 3760 and 4062 was dominated by sprat of 12.5 - 15.0 cm in length. In the two latter rectangles sprat of 7 - 9 cm were predominant while within the remaining rectangles sprat belonging to those length-classes were not numerous. The Subdivision 26 was dominated by sprat of 11 - 14 cm. Examination of the sprat age distribution (Fig. 5) also indicates differences between Subdivisions 25 and 26. In the 3 .. Subdivision 25 mainly sprat from age groups 2 - 4 were taken except for rectangles 3760 and 4062 where sprat of the age group o were prevailing. In the Subdivision 26 younger (age groups 1 3) sprat were caught. Only in one rectangle (3764) a greater share of juvenile sprat (407. constituted age group 0) was found. The total length of herring taken (Fig. 6 and 7) ranged from 7.5_ cm to 29.0 cm. Fish ·from 16.5 cm to 22.5 cm in length were predominant. A considerable number of juvenile herring from length classes 8 - 12 cm was found in rectangles 3760 and 3764. Results of age readings (Fig. 8 and 9) indicated that they were herring from 1994 year-class (age group 0). Catches in the remaining rectangles were dominated by 3, 4 and 5 years old herring. An exception was age distribution in the rectangle 3863 where also age group 1 was Quite abundant. The estimated biomass of herring and sprat has been converted into numbers by successive age groups (Tab. 3 and 5). For each rectangle age-length keys and mean weights by age-groups were calculated (Tab. 4 and 5). That kind of presentation of the Baltic fish biomass hydroacoustic assessment survey results has been prepared in the SFI (Gdynia) for the first time. In 1994~ the percentage share of herring, sprat and cod in catches changed compared to 1990 (Fig. 10). A nearly stable distribution of the herring share in control hauls observed in 1990, changed towards a considerable increase of the herring proportion within rectangles of the Subdivision 25. In case of sprat an opposite trend took place i.e. their share has increased within rectangles of the Subdivision 26. On the other hand, cod taken in the pelagic layers of the Subdivision 25 occurred in a greater share than during 1990 survey. Following the change of the percentage share of herring. sprat and cod in catches from control hauls conducted in 1994, the pattern of biomass distribution in the Polish EEZ has also changed (Fig. 11). Compared to 1990. the sprat biomass in the Subdivisions 26 and 25 increased considerably. Large quantities of herring appeared in rectangles 4060. 4061 and 4062 outside of the Polish zone. However, within the zone the.herring biomass dropped by about 6.500 tonnes. mainly within the central part of the zone. A considerable increase of the cod biomass was no ted in the western part of the EEZ. An insignificant Quantity of cod biomass occured in the Gdansk Deep (rectangle 3863). In the Polish EEZ changes were also observed in the biomass distribution with regard to north-south (N-S) and west-east (W-E) direction (Fig. 12). The biomass distribution in the diFection N-S was analysed by summing up the estimated fish biomass, originated from rectangles having identical two initial numbers in the rectangle code. The distribution in the W-E direction was estimated taking into account two last numbers in 4 the code. Compared to 1990. the presence of herring was much more distinctly marked in the western part of the surveyed area (e.g. rectangles 3700. 3800. 3900. 4000 - were conventionally depicted by --50) and to a lesser extent in the eastern part (rectangles --53. --04). With regard to N-S distribution a considerableincrease of the herring biomass in the southern part (--38) was noted. Sprat biomass shifted northward (rectangle 39--) and eastward (--02. --03). Cod shifted to a large ext~nt to the southern part of N-S direction and to the western part of the W-E direction. Acknowledgments Authors extend their thanks to dr N. Hakansson from the Institute of Marine Research in Lysekil for consultations as weil for his participation in the calibration of the r/v "Baitica" acoustic system. and to dr A. Orlowski from SFI in Gdynia for his advice on the biomass calculation method. S~edish • • Summary During 3 - 20 of October 1994 r.v. "Daltica" carried out its maiden hydroacoustic survey. The investigations were part of an international pro gram of the Baltic clupeoid fish assessment prepared by the ICES Planning Group for Hydroacoustic Surveys. The main task of the cruise was assessment of herring and sprat stocks in terms of biomass and numbers in the ICES Subdivisions 25 and 25 (the southern Baltic). The r.v."Baltica" survey continued the series of r.v. "Profesor Siedlecki" hydroacoustic cruises conducted during 1982-1990. The survey covered the area of 14774 NM 2 . Echosignals were integrated over 1204 one mile distances (ESDU). Twenty nine control hauls with midwater trawl were conducted. 9203 specimens of herring. 4512 of sprat and 73 of cod were measured. Otoliths from 1459 herring. 709 sprat and 73 cod were collected for ageing. An insignificant increase of the herring biomass density from the level of 28.2 t/NM 2 in 1990 to the level of 30.1 t/NM 2 in 1994 was noted; moreover a considerable increase of the sprat biomass density from 17.5 t/NM 2 to 44.2 t/NM 2 as weil as also marked au~mentation of the cod biomass from 1.5 to 3.7 t/NM 2 • respectively. was observed. Certain changes in the distribution of herring and sprat biomass compared· to 1990 in the Polish EEZ has also occured. Relatively smaller quantities of herring were found in the Slupsk Furrow and greater in the southern part of the Bornholm 5 .. Deep. The October 1994 aggregations of sprat shifted northeastward. Pelagic biomass of cod moved to the western part of the W-E direction and to the southern part of the N-S direction. Catches from control hauls conducted in the Subdivision 25 were dominated with sprat of age groups 2 - 4 while in the Subdivision 26 predominated fish by one year younger. The juvenile sprat occurred in trace Quantities in almost all rectangles of both Subdivisions but the southern part of the Bornholm Deep. Gulf of Gdansk and the southern part of Middle Bank where they prevailed. In the above mentioned areas juvenile herring was also present whereas in the remaining rectangles 3. 4 and 5 years old herring dominated the control catches. The estimated biomass of herring and sprat has been converted into numbers by succesive age groups. For each rectangle age-length keys and mean weights by age-groups were calculated. That kind of presentation of the Baltic fish biomass hydroacoustic assessment survey results has been prepared in the SFI for the first time. References . ANON .• 1994. Report of the Planning Group for Hydroacoustic Surveys in the Baltic. ICES C.M.1994/J:4 Orlowski A.• 1993. Distribution of fish stocks vs environmental factors in the polishfishery zone; Determined by the acoustic method (cruise of R.V. Profesor Siedlecki. Oct. 2-24.1990) Bulletin of the Sea Fisheries Institute. 1(128)/1993 •• 6 Table 1. Calch data r. v. "BaItica" , 3-20.10.1994, hydroacouslic survey No. ofhaul Date Time start UTC I 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 94-10-05 94-10-05 94-10-06 94-10-09 94-10-09 94-10-09 94-10-11 94-10-11 94-10-11 94-10-11 94-10-12 94-10-12 94-10-13 94-10-13 94-10-13 94-10-15 94-10-15 94-10-15 94-10-15 94-10-16 94-10-16 94-10-16 94-10-17 94-10-18 94-10-19 94-10-19 94-10-19 94-10-19 94-10-19 07:50 16:00 07:30 09:00 11:00 17:00 07:30 12:05 15:40 17:55 13:00 15:55 12: 15 15:30 17:50 07:25 09:55 12:05 15:50 09: 15 14:55 16:45 09:35 15:55 07:10 09:50 13: 10 15: 15 07:05 Latitude start DD 55 55 55 55 55 55 54 54 54 54 55 55 55 55 55 55 55 55 55 55 55 55 55 54 54 55 54 54 54 MM 22.9 21.1 32.7 35.6 26 3.2 31.8 53.6 55.7 57.9 25.4 15 44.1 25.8 20.5 35.4 36.5 25.7 16.2 59.8 49.9 52.6 40.8 29.8 55 2.5 45.3 35.7 29.4 Longitudc start MM DD 15 15 15 15 15 15 15 15 16 16 16 16 17 17 17 18 18 18 18 19 19 19 20 18 18 19 19 19 19 20 27.3 52.8 56.1 54.9 35.7 50.4 37.7 11.6 2.1 49.1 49.5 46.5 50.9 40.1 27.8 55.2 56.8 46.8 13.5 28.9 37.1 6.1 54.8 49.7 11.6 22.3 23.8 33 Duration Speed ffiJn. kn 00:50 00:30 00:30 00:30 00:30 00:30 01:00 00:30 00:30 00:30 00:45 00:30 00:35 00:30 00:30 00:30 00:30 00:35 01:00 00:35 00:30 00:30 00:30 00:30 00:30 00:30 01:00 00:35 00:30 3.5 3.5 3.5 3.6 3.5 3.5 3.5 3.5 3.5 3.5 3.5 3.5 3.5 3.9 3.5 3.5 3.7 3.6 3.5 3.5 3.5 3.4 3.7 3,4 35 3.9 3.5 3.6 Water depth m 91 93 80 78 88 78 52 76 54 Headrop depth m 43 46 40 50 12 12 34 72 48 78 61 73 83 100 89 94 88 97 62 62 74 62 97 103 101 78 48 19 34 22 32 30 68 60 64 30 54 30 20 42 25 57 55 41 45 18 Nel opening m 0 17 26 26 28 28 24 26 0 0 26 26 26 25 25 24 25 25 22 36 24 25 28 24 28 28 26 25 26 Validity invalid invalid invalid Total catch kglh 0 50.7 101.6 155.3 505.7 57.9 6.9 55.1 0 5.9 37.2 60.6 11 875.6 3":.7 201.2 192 670.6 43.2 325.4 207.6 615 231.8 79.5 54.7 151.6 334.5 661.7 14.9 __ ~ __Table 2. Fish density and biomass estimates by lCES squares r.v. "Baltica", 3-20.10.1994, hydroacouslic survey • leES rectangle Area [NW'2] 3760 3764 3860 3863 3864 3960 3961 3962 3963 3964 4060 4061 4062 4063 4064 4065 685.0 162.7 1043.8 706.4 946.1 985.0 1032.2 1032.2 1032.2 1032.2 1019.3 1019.3 1019.3 1019.3 1019.3 1019.3 14773.6 <I > <Svc> [mm] [dB] -46.58 62.40 -46.14 69.23 -48.47 40.39 78.15 -45.61 59.37 -46.80 23.75 -50.78 39.81 -48.54 -47.69 48.36 -48.02 44.84 74.38 -45.82 56.44 -47.02 -47.50 50.54 -54.05 11.18 46.60 . -47.85 107.75 -44.21 -44.44 102.24 Biomass area density [t/NM"2] Herring Sprat Cod Biomass [tl Herring Sprat Cod 6.25 1.89 41.36 40.42 52.86 18.47 23.77 17.77 3.85 9.50 59.08 65.59 19.93 21.37 72.24 0.31 25.89 71.25 8.70 51.37 36.36 17.37 36.13 51.56 54.36 86.69 18.16 16.21 0.95 41.33 81.98 127.06 25.02 0.00 12.19 1.03 0.30 2.55 1.51 0.00 0.00 0.00 13.16 6.09 0.00 0.00 0.00 0.00 4279.4 17736.3 307.0 11592.7 43167.0 9076.5 28555.4 36285.1 50011.2 34402.3 18190.4 17112.5 24538.4 37292.5 18341.2 53220.3 3978.1 56114.3 9802.7 89485.7 60221.5 18513.4 66851.5 16518.0 20318.7 964.1 21780.2 42132.2 73630.8 83560.5 310.9 129515.3 17141.2 0.0 12727.9 726.1 288.0 2511.1 1559.4 0.0 0.0 0.0 13417.4 6207.7 0.0 0.0 0.0 0.0 30.07 44.24 3.69 8 444284 653522 54579 Table 3. Estimated number (in milIions) of herring r.v. "BaItica". 3-20.10.1994. hydroacoustic survey Strata SD 25 26 Total AgeO Age 1 3760 3860 3960 3961 3962 4060 4061 4062 420 927 353 584 512 1180 1559 456 391 25 7 77 48 3764 3863 3864 . 3963 3964 4063 4064 4065 27 1334 1225 116 238 585 1998 7 21 152 33 0 2 1 11 201 7 0 0 Age3 Age2 Age4 Age6 Age5 Age7 Age8 6 167 58 19 4 103 48 29 0 106 29 42 35 107 143 73 0 259 89 192 100 228 483 91 6 221 109 171 176 284 458 111 6 133 49 44 114 293 109 63 0 4 2 29 25 68 86 35 6 12 7 9 6 66 22 21 5 386 126 9 12 40 24 3 0 180 162 14 28 85 0 331 207 34 66 223 453 2 2 109 260 42 76 150 753 0 0 108 290 14 27 32 370 0 0 23 127 2 14 26 134 1 0 0 20 2 1 6 16 112 0 16 17 72 0 Agel0+ Age9 6 0 0 1 3 15 3 12 0 6 4 72 0 0 0 0 1 1 2 3 101 0 01 0 0 1 0 01 31 0 9 0 0 0 7 4 01 01 01 01 01 01 11 01 01 I Table 4. Estimated mean weight (in gramms) of herring r.v. "Baitica", 3·20.10.1994, hydroacoustic survey SD Strata AgeO Total Age 1 Age2 Age4 Age3 Age5 Age6 Age7 Age8 Age9 Age 10+ 25 3760 3860 3960 3961 3962 4060 4061 4062 10.2 46.6 51.5 42.0 35.8 51.0 42.9 44.6 7.6 20.9 19.3 6.3 5.6 5.1 6.2 5.0 5.5 39.0 39.6 38.2 20.4 39.0 34.4 24.9 0.0 49.6 49.1 41.7 33.2 39.1 40.5 31.5 0.0 49.1 53.7 45.7 38.9 48.6 45.6 37.3 70.0 48.6 54.7 46.7 37.2 50.8 47.7 41.3 40.0 46.6 56.5 53.4 40.2 56.4 57.1 50.8 0.0 112.5 80.6 57.5 45.1 61.6 64.1 56.8 75.0 66.7 56.8 76.7 66.6 61.6 82.1 77.6 40.0 0.0 0.0 83.8 65.2 76.8 121.0 83.1 0.0 0.0 0.0 80.0 105.0 112.8 98.3 107.8 0.0 0.0 0.0 0.0 68.8 0.0 0.0 97.5 26 3764 3863 3864 3963 3964 4063 4064 4065 11.3 21.4 40.8 34.2 41.2 37.2 36.9 47.5 5.9 5.7 6.4 0.0 10.3 10.0 0.0 0.0 27.5 12.7 23.3 34.3 35.8 25.1 20.7 36.7 0.0 20.5 35.0 29.6 36.3 30.5 29.3 0.0 0.0 24.3 44.2 32.9 39.7 34.2 34.8 40.0 32.5 31.9 44.0 31.0 37.2 34.6 31.3 0.0 0.0 27.3 46.5 43.0 44.3 55.8 40.6 0.0 0.0 73.4 44.1 64.3 53.8 55.2 42.3 95.0 0.0 64.9 77.7 80.5 76.5 72.0 57.3 0.0 0.0 66.2 76.1 83.8 68.2 101.6 55.4 0.0 0.0 68.8 0.0 0.0 0.0 75.8 111.7 0.0 0.0 0.0 0.0 0.0 115.0 90.0 0.0 0.0 Table 5. Estimated Dumber (in millions) of sprat T.V. "Baitica", 3-20.10.1994, hydroacoustic survey SD Strata AgeO Total Age 1 Age2 Age3 Age4 Age5 Age6 25 3760 3860 3960 3961 3962 4060 4061 4062 4295 663 1211 2394 3698 1051 988 213 4088 204 315 156 0 68 48 180 69 25 40 7 126 28 0 0 90 145 285 318 828 188 112 0 47 159 344 807 1513 306 288 7 0 92 138 594 754 251 296 7 0 27 521 280 215 203 161 10 0 2 21 211 122 5 78 7 26 37641 38631 3864 39631 39641 4063 40641 40651 1803 3136 3433 4185 7228 3459 7124 11533 7901 72 137 100 361 62 781 0 435 834 1243 126 380 1251 1284 1670 3448 1785 3733 4809 162 646 697 16821 1944 854 1026 11071 16 210 45 360 405 367 93 265 131 441 171 2341 2961 111 21 1381 0 34 10 13, 221 142 21 0 I 1077 138 2137 5213 Age8+ Age7 0 11 15 19 0 0 4 3 71 0 0 01 01 01 0 01 1 0 0 2 0 0 0 0 0 0 44 0 0 0 01 0 0 Table 6. Estimated mean weight (in gramms) of sprat T.V. "Baltica", 3-20.10.1994, hydroacoustic survey Strata SD AgeO Total Age2 Age 1 Age3 Age4 Age5 Age6 Age7 Age8+ 25 3760 3860 3960 3961 3962 4060 4061 4062 4.1 13.7 14.1 15.6 14.4 17.6 16.7 4.5 3.6 4.2 4.0 4.5 0.0 4.0 4.5 2.5 11.7 13.9 13.3 10.0 12.6 12.2 0.0 0.0 16.3 16.8 16.6 14.5 13.2 15.6 15.6 0.0 15.0 18.1 17.9 15.4 15.0 17.7 16.2 12.51 0.0 19.1 19.1 17.2 15.9 18.4 18.1 16.7 0.0 20.2 19.9 19.6 18.2 19.7 19.5 17.2 0.0 21.7 19.8 15.8 16.4 21.8 16.6 15.0 0.0 20.8 21.0 20.0 0.0 0.0 20.7 15.0 0.0 0.0 21.5 0.0 0.0 0.0 0.0 0.0 26 3764 3863 3864 3963 3964 4063 4064 4065 6.4 11.6 10.0 13.4 12.4 12.2 11.7 11.2 2.9 4.4 3.2 3.4 4.5 2.8 4.2 0.0 8.1 9.5 9.1 10.9 9.9 9.9 10.5 9.9 9.6 11.4 10.4 12.7 11.9 11.6 11.9 11.2 10.21 13.71 11.5 13.81 13.6 13.1 12.71 14.9 11.4 13.3 18.1 16.1 14.3 13.9 16.5 20.5 16.0 18.1 14.0 17.4 16.5 15.2 . 17.61 15.0 0.0 17.1 22.0 20.0 20.0 13.7 17.6 0.0 11.7 0.0 0.0 0.0 0.0 0.0 0.0 0.0 0.0 17.41 0.01 0.01 0.01 0.01 0.0 0.0 ..... N i. Fig. 1. Location of acoustic sample"s and control hauls T.V. 59 "Baltica", 3-20.10.1994, hydroacoustic survey 62 61 60 63 20 56.00 . L-.---,.c..kL-+------+-----+-....:.....------t----:::---r::-=.:::;;,--=-~. ~~'"IF:::----I . . . ' .: .. .:'" ..... . .. ...- - . .•.~. ..' .....' .•.. .., . 1~ ~:::".: . 2 w.-: .. - -.-. : ..- . )_ 23\ .. \ .. ..•.. 'i ........-- 2 ~. •.. 55_00 ' :: 104-.00 .:' ...... .... M.oo 65 64 15.00 .... ::- -...- ........ ~ .... ::::::;-...•...... . - . 21.00 Fig. 2. Distribution ofherring biomass (t/NM2) T.V. "Baltica", 3-20.10. I 99-t, hydroacoustic survey 60 59 62 61 63 65 "- 56.0 55.0 54.0 14.0 64 15.0 16.0 17.0 18.0 . Fig. 3. Distribution of sprat biomass (tINM 2 ) r.v. "Baltica", 3-20.10.1994, hydroacoustic survey 59 60 62 61 63 64 65 56.0 0 ~ ..... U'1 Q) C"J 55.0 CI) C"J 54.0 14.0 15.0 16.0 17.0 18.0 19.0 20.0 SubdIvIsion 25 Subdhlsion 26 3764 3760 i~Lh 18 30 Lfllt.-D 6.5 7.5 8.5 ~L. ;g .1..--.•111. _ 3860 3863 SOL ~L.__1.1••._ I •. 40 30 fg 30 •• 1 _ &5 ~5 8.5 a5 • •1_1_•. 10511.51~51351~51~51a5 5°L 65 7.5 8.5 95 10511512.5135145155165 3864 ~gL 40 30 ..llL__ ~~ •.•.• -llIIJl__~ 6.5 7.5 65 9.5 10.5 11.5 12.5 13.5 14.5 15.5165 3961 3963 11_ ._11.11 __ 01 ------_----- 6.5 7.5 6.5 9.5 10.5 11.5 12.5 13.5 14.5 15.5 165 ~o _ _1.111_1.__ 1, __ 6.5 7.5 85 95 10.5 115 12.5 13.5 145 155 165 3962 __ 3964 ___ .I.lh.. lo!g 1... -_ 6.5 7.5 8.5 9.5 10.5 11.5 12.5 135 14.5 15.5 16.5 ro~IL. •.1111.....,_ _ ~ _ 6.5 7.5 8.5 95 105 11.5 12.5 13.5 14.5 155 165 4060 4063 ~og1---------..-.....dlLAI'''._ ro~I. . .llu.. _ 1U • • 1 6.5 7.5 8.5 95 10.5 11.5 12.5 13.5 14.515.5 16.5 6.5 7.5 8.5 9.5 10.511.512513514.5 155 165 4061 4064 ___.dIJh__ ~g~L . du.. I.U. 90 111_ 0 _ 6.5 7.5 6.5 9.5 10.5 11.5 12.5 13.5 14.5 15.5 16.5 0 • . __ 20Ig 3960 ig _ 6.5 7.5 8.5 9.5 105 11.5 12.5 13.5 145 15.5 165 9.5 10.5 11.5 12.5 13.5 14.5 15.5 16.5 6.5 7.5 8.5 9.5 10.5 11.5 12.5 13.5 14.5 15.5 16.5 4062 ~lm. &5 ~5 8.5 a5 _ 6.5 7.5 8.5 9.5 10.511.512.513.514.515.516.5 4065 . u __ ~o l _~ I 1-.-.. 1Q511.51~51351~51~51&5 --_.-----------_._-------- --------------- ------ bI. __ 6.5 7.5 8.5 9.5 10.5 11.5 12.5 135 145 15.5 165 --------- length (an) length(an) --------------------------------- --Fig. 4. Length distribution of sprat by redangles 16 Subdivision 25 Subdivisioll 26 3760 3764 '~I~ o _2gl•• _ 1~1 2 3 4 5 6 7 o 8 1 234 40 2g o 1~L.. 60 40 2g _ ••• _ 1 234 5 6 7 o 8 1: 3960 1~l.. 2g .. 40 o 1 2 3 40 2g _ 4 5 6 7 8 1 u.__.. 2 3 4 5 6 7 8 Lu 1 2 456 7 8 567 8 ......... _ 3 3963 100 80 60 40 -.l.-..JI • • - o. o 2 3 4 5 6 20 7 o 8 'rogl_~_ . ....1.2 3 4 5 2 3 3964 6 7 -'.~ o 8 1 _. 234 _ 5 6 7 100L. ~ u.__.. _ 80 o .----o 234 ---- 567 o 1 234 5 6 7 8 8 4064 4061 . '~IO----..-. ....u.--..-----o 234 5 6 7 8 '~I _ oW....-------o 1 2 3 80 100~ 60 .. ~--.. o 2 3 4 4 5 6 7 8 4065 4062 40 ,.2g 8 4063 4060 '~I 4 40 2g u o o 1:L. 3962 • _ 3864 o 3961 '~I 8 3863 3860 1~~ 567 5 6 _ 7 '~Lu o o 8 age 1 2 L.J. - 3 - - 4 - - 5 - age Fig. 5. Age distribution of sprat by redangles 17 - - - - 678 3760 40LJ 3962 30 20 10 o .-J.IIJ.u.'___ _~_JI.'__................. t 7.5 10.' U U.S 1.5 1'.' •• 19.5 2l 115 ~_ _ US .z4 27 21.' 1ll 0 UUAl.. .-JI.... IIJI.- ..h 7.' 9 10.' U l~U n u I'.' .1 3860 io~l, . ~__ .... _ 7.' • t 10.5 11 13.5 1S 16.5 11 20t 22.S 15.5 11 18.' i'o~I, __~ . . 7.S , 10.' U U .6.' .1 1'.' t 10.' 11 13., U 16.' 11 lLS ioll. .••. ........ 14 25.5 21 Je.' 7.' , 10.' 12 40 ••I.Wlh..... _·..-.IJ.IJ.IJ.............. , 10.' U U U 16.5 .1 I'." 11 21.' 14 io~ 1~. . __~. . - -.l1,UJ u., 1'.' )T la.' 19.' ns 21 1-4 aB J" D.S lf.' .1 ." 11 21.' z.4 "t 27 11..' 4062 30 7.' I1 -"...... ..111111•••••• 13.' 3961 ~o~ 1. _ M 4061 .....1I. ..111111.1•••••. U.J 12.' -"...... ..111111111.... _ 7.' 3960 io~I_~ <-21 4060 ldlJl..h. .1 JIH ••. , US 27 2:1.5 Ic;ngth (an) 7.' I .I.IJIJU............... 10.' 11 13.' 15 16.' .1 .,., Jt 11., Ic;ngth (an) Fig. 6. Length distribution of herring by rectangles in Subdivision. 25 18 "__~ lI.-J'_...... ••11.1111......... _. 9 J4 U, J7 'U 3964 3764 ~lku. 7.5 , IO.J U u.. . . U.5 U 16.5 11 19.' _ 11 n.s 14 215 11 11.5 i:o~ ...........................JI.JIJ............_ - I; 1.' 9 lOS 11 13., .111111111111••••• U 16.' 11 19.' 11 21.5 24 1.5.5 14 ns 4063 3863 40 30 20 1l , -__.•.•. . __...1111.1111111•••• .. 0 ~ 1.' c a.JIUl.lLaA...................... , 10.' U n.s 15 14.' tS 1'.' 21 12.' _ 14 U5 27 UJlIJl 1111.. 1...... 1_ 'Oll , 7.' 11.5 , 10.' 12 11.' U 1'.' .1 19.' 21 n.s 4064 3864 40 30 20 10 o 1---_ _~__JU Jll.JIJhul.... III.JIJ ....... _ ......... 1.' , 10.S 1Z U.S U 16.' .1 19.' 21 12.' 24 2.5.5 27 2a.S ioll,_ . ..11111111111•••••• . . I.I.IJ 1.' , 10.' 12 13.' U 16.' 11 19.' 21 12.S 4065 3963 40 30 1lo1_ 7.S 20 10 o 1111tttJ1••••••• --JUJIJ , 10.S 12 11., U 16.5 .1 .,., 21 ns 24 US length (cm) .- .. 27 lI-S .....-JL-__-JL_ _......_ ~ 1.' , 10.S U 11.' U 16.' .1 19.' U length (cm) _---------- Fig.7. Length distaibution of herling by rectangles in Subdivision 26 19 ll.S J.. nJ 3962 3760 '~~ 0 'OOl 80 60 ~~ 1 2 3 456 7 8 9 10 o ..d.... 1 2 3 4 I 2~ _ o UW.. . .~-~~~~ •• 1 2 3 456 7 8 9 10 '~I o 8 9 10 _ 12345678910 4061 100 80 60 100 80 40 40 60 20 0 0 1 2 d. 3 4 5 6 7 8 9 10 2~ .........-......IJ..-~-....o I • o 4062 . __ 1 2 Ul,J-......-__-_~~ 3 4 5 _ 2345678910 1 3961 2~ 7 - • . 0..--..--. 3960 80 100 60 40 6 4060 3860 80 100 60 40 5 6 7 8 9 '~j 10 o ..... _-- _ 1 2 3 4 5 6 age 7 8 age Fig. 8. Age distribution of herring by rectangles in Subdi\'ision 25 20 9 10 - ------ ---- - - - - - - - 1 3764- ,ooL lEI 2~ 80 60 40 • 0 1 2 3 4 .dL... __ 2~ 5 3964- 6 7 8 9 0 10 1 2 3 4 I~I 100 80 60 40 20 0 1 2 3 4 5 6 7 8 9 0 10 100 80 60 60 40 40 20 1 2 3 4 5 20 0 0 2 3 4 5 6 7 8 9 10 0 1 2 3 4 3963 I~I 0 -.d. 1 2 3 4 5 9 10 6 7 8 9 10 4064 100 80 1 8 -.11 ___ 3864 0 7 4063 3863 0 6 5 ._-5 6 7 8 9 10 4065 100 80 60 40 20 0 6 7 8 9 \- 0 10 1 2 3 4 5 6 age 7 8 9 age Fig. 9. Age distribution of herring by rectangles in Subdhision 26 21 10 e - "Tl ciQ' ..... N~O\OOO 000000 0 o CZl :3 g-, 3760 0"0 "0 e; (l) ~ o· 0..0 ... :3 0"0 ... 0 ::r-tn ...... o ll) N N -, 01 ::rc Ei> S' ..... \0 \0 .+:0 3760~ ~ N~O\OOO ; - 000000 3760 JIi 3960 3961 3%1~ 3961 3962 4060 4060~ 4061 4061. 4060 4061 4062 ~62b- 4062 e: (') () 3764 3764 ~ 3764 3863 3863 3864 3863 3864 3963 3964 3963 3964 4063 4064 4063 3963 3964 4063 4064t;= 4065 j 4064 4065 -I I \0 \C \0 \C O~ ~ ... ... ~ ~.~ '"o ll) N~O\OOÖ 000000 3960 \0"'" \00 0 0 e. - (') 83860 -otO o... _ ;.> ::3 • 3860 0 ....,::3 ..... 0 ......, e ~ t:". ... () ;.>_ 3962 CI) CS" 4065 3962 , i 3860 3960 3864~ ::c 0 ::l 5' 0'Cl .--------------~- - - - --- .. Ilerring 250000 200000 150000 100000 50000 0 _.§iJ.mJ 0 'D ,..., t'- 0 o - NON ~ ~ ~ ~ ~ ~ M f""') M o _ 'D M ~ ~ ~ N 'D 0\ 0 'D ~ lt) N Cl v.l oBlllh.Il _a-~ ,..., "1" ,..., "1" ,..., 'D 'D 'D 'D 'D t-e ,..., 00 00 0\ 0 ,..., ,..., ,..., 0\ ,..., "1" "1" "1" 'D 0 "1" lt) ~ lt) 'D 0 "1" 'D N Cl v.l ~ !-< Sprat 250000 200000 150000 100000 50000 0 0 'D 0 'D 00 ,..., ,..., t'- ~ ,..., ~ ,..., 'D N 'D ,..., ~ ~ ~ "1" ,..., "1" "'" "I:t 'D 'D 'D ~ ~ t'- ,..., 00 00 ,..., ,..., ,..., ,..., lt) 8 v.l f"") 'D 'D 'D 0 ~ ~ "1" 'D 8 v.l ~ !-< Cod 25000 20000 15000 10000 5000 0 0 0 'D 00 ,.,t-e ,., ~ o - ~ ~ ,..., ,..., N ....~ 0 'D 0 "1" 'D N 'D ~ ~ "1" 'D lt) ,.,t'- 8 v.l ,..., 'D 00 ,..., "1" 'D 00 ,., M ~ M '7 ~~'D'D ,..., ,..., ~ ~ lt) 'D ~ 'D N Cl v.l ~ !-< - 1994, r.v. "Baitica" c:=:J - 1990, r.v. "Profesor Siedlecki" Fig. 11. Hydroacoustic estimates ofherring, sprat and cod biomass (t) in the Polish EEZ by rectangles in 1994 compared to those of 1990 23 Herring 250000 200000 150000 100000 50000 o 37" 38" 39" 40" "60 "61 "62 "63 "64 Total 37" 38" 39" 40" "60 "61 "62 "63 "64 Total 37" 38" 39" "60 "61 "62 "63 "64 Total Sprat 250000 200000 150000 100000 50000 o Cod 25000 20000 15000 10000 5000 o 40" Northward (two initial numbers ofthe rectangle code) Eastward (two last numbers of the rectangle code) ~ - 1994, r.v. "Baltica" biomass estimate c=J - 1990, r.v. "Profesor Siedlecki" biomass estimate Fig. 12. Changes in the distribution ofthe herring, sprat and co biomass in the Polish EEZ in 1990 and 1994 in N-S and W-E directions 24