;"]SSI-N}] OREGOI: STATEGAiE GAI CO] coIIssIOH OREGCi:S?ATE SCALES SP.OU? IIGO}' OFSTEET,}EAD STEEUAL TROUT SCALES Rlii,IIiIG FRC]' TlE Th RELL PSSI"LTS RESULTS FRO? CATCH Ci]I.IRCIAL BAT TILIIJ.COT FROT TUE TILLAUCOK BAY CCIi ERCIAL CATCH T1:E FR.OI' bY Rcport by A Report Surnngr F. F. H. H. Sumner Ssalor BlcLogiet Senior Biclog1t to the tha to Oregon Stata State Gana Grie Comnieslcn Coiiasicn l]eedham R" Par:1 Dr. Paul H. Needharn lrn Fleherlee of Fi5herie Director Dlractcr of 1946 ]".Zn1946 July Ju)-y 12, *L-3.lntroductlon Introduction scales During D u r l n g t hthe e w iwinter n t e r o f l of g 4 l -1941-42, 4 ? r t } r g l r r the l t o r cwriter o l ] . a g tcollected odgcaler was so Hcvever, ltit was so dlfflcult d±fficult IIoNFvsr' steolhead" from anglsr-oaught angler-caught steelhead. from popular varioua popular by vlsltlng visiting various catoh by tbgir catch wlth their anglars with to flnd anglers to find vas that itlt was checkad), that npz"e checked), (63 steelhead stoslhead were and rsEorts areas resorts (63 ar€a8 and Clty Bay City tTio Bay at two st€alhead at steelhead glll-nottsd fron gill-netted data from decided to get data doeldcd to 50 Jaauary 30 from January c]:ocbed from wars checked fLeh were The ::etiad netted fish The plaats. packlpg plants packing 194?. 28, 1942, to Fabnrary 28, tc February tldeths tideln the re taken taksa in fleh rere the fish The groat gxeat raaJorlty majority of of the Iha from cans from The The remainder remainder came Rlvsr' section of the the l'l11son rilson River. esction of to the set-nets s a t - n e t s i in n t hthe e l o alower r e r b a y'cay. . l h e f l sThe h l e r efish d e l twere v a r o ddelivered totha 'wats' wate of the packing the packer p a a k e r bbr : , ' ttruck r u c k o or r b oboat a t a : : dand c u mdumped p o d o u t hon eflo o r ofloor fthepacklag from the fish before took shed e h e d oon n a pal .pier. o r o f h e The s ' r l . twriter ertookd . a t a fdata romthst.lshbefore markets were Yourk they packed. t , h a ywere werep a c k e d ' S t Steelhoad e a l h e e d s esent n t t o to l . i eNew wYou rkme'rkete$Jgrg Tl:ose Those daettrcd destined (net cleaned). e }eercd)' ror:::d (not .n t]:e the round boxed with !.ce, ice, 1n boxad, elth Flslx Feh ceLlfornla tagged wlth with aa Ca1forna a::.d taggsd clea*€d and rars cleaned for California caitfor*ia were for bozed lnin lcs' ice. beiag botod before being (la:no ComrLselon and Came Connissicn tag tag before and Dats Data gcaLs Of the scale 0f tho sxaninsd' rnrs examined. steelhead were Eight hundre* steelhead Elght hundred co growbh and so ln growth regeneration in percent) showed ehosad regenoratlon ?4. (9 percent) sainple, 74 sanpled, {9 dubloug" ware dubiou. epoctrnenswere 1?specimens to sex, eex, 17 As to ussd' As be used. could not be oould not spaclnena used of specimens nu:rber of the number brought the date brought Other gaps gapslnin tha the data 0thsr 702' to 702. down to tabi.eg down in appndod tables the appended ln the nBssultg from ths from the wrlter' "Results tha writer, by the report by prevlorrs report In Ia aa previous steelhead trout scales with recomreading r e a d l n g oof f g ucutthroat t t b r c a t a n dend atee].hgadtroutacaless'l.threoou!ar:mrary of work" (Juns (June 16, 1942)' a asuimnary 16' 1942), mendationsfor for future future 'ryorkt mndatlons is presented data d a t a ffrom r o m l 0100 o o f t of h o sthe O C e800 o w ]commercial € r c i a l e t e s l hstesihead eadlgpresaated anl 8B' 6 and in Tables Tables 6 ln ths 1n the fiEh in of fish number of by number The perosutages by Pr.o perceatages o,?_^ follows that for the larger various Y $ 1 o u 8 aage g € g lgroups o u p g € gessentially soatlallyfollorsthatforthglergar the number n D t r b € f o of f f l gfish h t r etreated a t e d o f l nof t h ain p r othe s a apresent t r a p o r t , areport, l . t h o u g balthough tJ:c adaguata' eo adequate. lnea:rlt so figures ara by no means f,lgurea are Stsolbead of Steelhead Sizes Slzgs of leagthr the average avsraga lengths glvas the rcport gives prevloua report of the previous Table fabb 99 of snal1 was such tuoh aa small thero was Slnoc there stsalhoad. Since ootrerolal steeThead. 8S oossnercial of 800 percentage p c r o e a t a g o of o f ffish l g b ddubious u b l o u s a sas t oto a asex, x , t h c the e a f learlier l c r f l g u rfigures caarr ppeated repeated hers. here. aze is fsnalec ir tha females of the length of fork length ths average fork The average lncbcr)o 28'? inchea). inches (a11' (all, 28.2 28'1 lacbag raalosp 28.1 the males, and of the 28.8 lacher, end ?8.! inches, 8'8 anaraglng 8.6 rolghcd' averaging wero weighed, Onehundrad hundredsomaty-elght seventy-eight flgh fish were Onp tha of the lcagth of shorter length lha ahcrtcr ??.1 !'nohes)n inches). The pouada (lengtU, pounds (length, 27.1 lnd!'nry indi2, msy Februsry 2, 3O tc February Jauua4.r 30 from January flah, weighed fish, taken tksn from rrlglrd prorc noad,adto prove ls needed but more data is ucrs data bu! dlstlnctreEs, cabs reia1 distinctness, oatg raglal polat. tbe point. the relationships of 702 conTable T a b l a l1g lgivea v e a t hthe e a gage-length e-l'gngthrolatlons}ripeofTO2eon- ireasured was smallest nrcial n r r a l a l ssteelhead. t s o l h e a d n T hThe gena l l e e t n a tnetted t e d f l s hfish noasuredgaa caught gsa-run steelhead gEsllgst sea-run etselhead caught ths smallest while the 1on5o shl1a 21.5 21n5 inches lnclres long, The 26-27.9-inch sport lin n t tthe reep o r t f l afishery h e r y r r awas g 1 1 11.25 . 2 6 t n cinches h e s 1 o long. ng.Ihaz6.z7.9-lncb sport fishgroup g t o u p lis g t hthe a l a largest r g o s t l n bin o t hboth t h a o the o g $ soomiercal r c l a l a n d eand portflgh- tho of the perocnt of 38 peroent aad 38 fors;r and tha former of the prcaat of 59 percent erl.es, bclng 39 erles, being the largest number of fish to latter. l a t t e r . T h This ! ' r g r ogroup u p a l salso o f u mfurnishes lshosthelargestn:mberofflsbto cLa!sos' year, classes. or year, aanulusr or the V annulus, fV and aud T II1, IV the III, Tablo ia Table precontod in ls presented groups is age groups of ace The average longtJrs of avarage lengths The thero that there notsd that be noted It will rr111be 2. It dlffer' grorth differ uttle is very little growth 1s very There is a regresence the c n c ' bbetween otTlenth c I - II-Il 1 a : rand d I 1 1 -Ill-Il I l g r o ugroups. ps.Thergllaregrca- as sion in ths the II-III Il-Ill as sloa 1n algo group' and also I-III tha I-Ill compared with the comparadwlth lin a t hthe e I I IIll-Ill - I I I a s o oas m ?compared a r a d r l i h with t i : e 1 1the - I I l Il-Ill g r o u p ' Igroup. ftflsh group1 If 3 fish the ellmlaatcd, thr wera eliminated, 111-111 group group were the 1II-I11 ln the under 26inches in r:ndcr 26.1nchgs s\) as sa:r€ as average would be be tho the same averagc rould _ for the 11-11lrs' II-IIi'e. for tho numLarger Largormum- such lrrrgularltlss' cut auoh irregularities. to smooth snooth out sadgd to data are ars needed bera oof data bers Fresh BrackishTJatgr rater aad Braoklsh Freah and Gfssrth Growth steelA A nmajor a J o r d l fdifficulty f l e u l t y i n r ein a dreading l n g t h o g o the a } C sscales o f a d u l t of g t sadult gltha ganoral' the Ir' general, In grovth' strean growth. the stream in interpreting lnterpretlag; the head le i la baad that for ocean o c € a n ggrowth r o r r r b hpattern p a t t e r n l is s m l c hch n cmore r € c o n consistent a l s t o n t t h a athan thatfor with o growth g r c ' s b h i in n f rfresh e s h w awater. t a r . A p p aApparently, r a a t l l r n a d u l adult t e t e a l eteelhead headwlthoe thaS the average, avarago' than on the fish, on growlng floh, stream anauLus annulusare are fastar faster growing strcara single are a f s tthose h o g 6 rwhich d i i c h e spend p e a d n more . o r e t l r tIme e i n t h ein s t the r e a nstream, , a l d t h e eand l n g l the e as nich ar€a area on on the the ecala ecale as as xrarcb snclosas as stream annulus oftea often encloses atrea.m ar:nulus growlug alower growing slower older' from older, eeales from oe scales annuli" on maytwo two oror three three annuli nay amiuli, the anmili may be f l g h o O : On r e c ascales l ' e e ' c rwith i - t h s c2r Bor a t3 r estream aclsnnull'theanaulluaybs fish. of but few cIrcuit, and ocr- wide seen, ora fnarrow, w l d s a and a d e aeasily sllysa€ arc:'n fcwrofbutfa'wclrcul1'aadoor- dletlngr:lsh' ta diatingush. reapondingly difficult to raspoadlngly ClfficuLt In I u B85, S r o ror l ' 212 p e r percent, o o n i r o f i ; hof e s pthe e c l specimens m o n s s t u d l o studied d t h e r e l s there is wj'thout grcnrblr wtthout watar growth ea1'Lwater or salt brackleh or gro,yyth rerglr:g stream merging into into braoktsh straam growth represents late eunurar down-. an s n qannulus. n n u ] . u g . T hThis l s p a pattern t t o r a p r oprobably bablyrapreseabslatesu!$grdwg- t}i g t r e a m nmigration. l g r a t l o n . T r aTrap p e t u dstudiea l a s E q . : lWill ] . a l d laid n c 1 gin r l fclarifying ytagtk.ls stream growth is svldsntly evidently 1aingrowtl: 1g tidewster, Brackish wetar, water, or or tidewater, Bracklsh polnt. point. speekons by by olrcull otrouli scala specimens tho scale cf the s0 percont, dicated on 420, or or 60 percent, of on 4?0, dlcated and ooarssr and are coarser vl1ich are pattern) ithich (outslde the etrs$l-grcwNh pattern) tf.e stream-growth (outside (tn circuit (in strsan olrou:.r snooth 1lks spaced, though like stream though smooth lrldely spacod" more ror.e widely Probabpattera)o oeean-groath pattern). ths ocean-growth ln the xarr-y clreuli contrast etrouli In tc the tha wavy oontrast to the I ish spent scme time In brackish lly y aa } alarger r 6 e r p ' oproportion p c ' t l . c n c f t } l of sfisi:spentgclptlrnslabracklsh of arsa of large large area s sufficiently er:fflclently '!vater. Tillamook providee a ay provides TiLlanock la;J water. stay of most of the young stealbrackish b r a c } i g h wwater a t e r t o to e x pexplain l a i n t n e sthe tarrofrrroetofthoyouaggtsE]'. oogann opga ocean. the open into the v.s:rtrrring into bgfore venturing ngdlug: before thr,tmedium head ln in tht hcad "tlal3q5!. he yot;ng *that rnoet of dstoglll-sjf besn detormIned has hag been 3f:3€.*Lor'I-t taiie g o li-'re r tr:br: tributaries. i(i 1'::i! 3 F!ver ,i ate. from f rrn KilchI *s Trapping TrappJ ::E data It It *+* ocaa:l' to the ths ocean. thelr way to ateeThead migrate dosnEtream downstream ln in ilay iay oa on their stealhaad nlgrata only aa mlgraats spend Bp€nd only of the the migrants naJortty of the majority It is probable probable thab ths the It ls (4 percent) percoat) 28 (4 only 28 for only wat,:r, for few months at at ths the most most lntracklsh in'ackish water, fer nonthe a,rauius. blacklsh-vrater annulus. a brackish-water shc+r a 7o2 specimens spacksns show of 702 of of A comparison conparlaon of A thosa rrlthout anC those anaulus and without an anmulus wj.th such suoh an of fish fieh with average lengths lengths of average euperior is not not gonersLly generally superior ln tlco?tarer growth in that growth has shown t±dewater ls shown that groa*rhn to to freslmater freshwater growth. conJ:Ience, thebrackish-1rgter brackish-water annull annuli afs are conl{€nco, tha tsbles. gtresm annuli tha tables. n the gtderad as stream a:suli i-a sidered rpeoiof epeciThe numbers of 11:e nr:mbars ?' ?ab1s 2. of Table the bottom bottop of at the shcvna at e:r3 shown e&:3u:.us are ii*e?sater aimulus r.'i.th aa tidewater mans neas with Age and Sox groups arid and r:'sbers numbers prevlouely previously age groups 4 show show the Tables the age and 4 Tables 3 and previously. epalrned praviouaiy' parceat had had spawned 21percent fis-n, 21 all the the fish, 0f all spawned, by sax. sex. Of spawned, by percent onl'y 15 15 percent previously, only spaarned provloue11;, had spawned fe:oaies had tl'-efemales Vhile percent cf oftho T,hllo 26 ?6 prcent of readraaddifficulty of Tne dlfflcultv so. The done eo, harra done to have fsurd to of the meles were found tha males of "vera differoaee' of the for scme the difference. ee:e of aeco';::t for shecks me;r lug male spawning may account epar,nL*g checks !.ag:aale than rrora than hsd spawned porcent h spa:rned more 3? percent apa*moc, 27 had spawned, tb.at had Of fsnales that of the ths females tT o had spawned lpa'iraed two percont had s? percent spa"*msd, 37 hsd spawned, thst had maLaa that of the the males once, wtrlLe while of onos, 8€a?afdr ni.gratlng seaward. bafore migrating epa'naedbefore aay have spawned fsw may or more t1ues. times. AA few or mor€ 'rilsrs notsd' flowever, no no clear-eut clear-cut axa:apLee examples ware rioted, Ilormevar, Ratio Sx Sax Ratio usod' ?0t specimens epocimas used, the 70 frcn the d'aterninod frcn as determined The sex retlo, as sax ratIo, the mslssn to 1.0 L'0 males. is fsrlalas to 1.52 females Ls 1.32 R:is Thl ttt practlcaLly t1' figure le practically ftguro is (t.lOr1'g)' ( f g E e ) for f l a h (l.36l.0). f o r 800 8 0 0 fish a p o r t e d (1942) p r o w l o u s L y rreported aeas s t h a t previously s a @ , athat forales, probab probabto select csl€ct females, te nets tend to note tend that t.rre lndlcato that The figures indicate Ilre flgures pLumpnass' grea'tor plumpness. ly because of of ths: th&rr greater 1y because r t Sexual A * a at aturlty Se:cuel-1I aturlty Age -bile steelhead sa:iual on reaching r€schin3 sexual staei.hsaC on of the ai;a of tl;o age Table shovre t;e ?ab1e 5 shows -oo -5.. maturlty. maturity. psrt naa spant spent tha the better better part (5tr.g percent) peroont) had group (55.6 The largest group The largeet watst' ealt water. years inin salt about 22 yaara anC about water and frssh water la fresh of yeaqs in 2 years of 2 in the also spent years in about ?2 yoare spant about gnaLler groups hed aleo groups had The next smaller Tha nart ylars :rearly 22 years nature after aftEr nearly aaxually rnaturo becarm sexually parcent became In 88percent Ia all, all, 82 oo€lnr ocean. osea:x. the ocean. (II in the (II annuli) snouLl) in g!'van parcont given 62 percent of 62 The comparableflgure figure of fhe conparabS.e and percant, and 78 percent, to 78 corrected to ehould be corrected report should of the page 5 the 1942 report E of on pags fish made nadc the the fish 3ixty-seven of the percont of Slxt1r-s€YgB percent perceat. 81 percent. fish, 81 for sport fish, for sport atfeesn secondrsinter winter lain tha the stream, gaa.rard mlgratLcn seaward migrationafter after thair their seoond H;'brids Hybrids are considerably oonslder*b1y thrse are studied, three sarylos studled, Of the the 702 scale samlos ?0? scale 0f g!l{" Two are fves from f1rma3'ete fomaes, ltro are their size. a?Efqgs for smaller thaa than the the average for thglr srallor rches long3 the thlrd third loogc ths 26'5 !.cches 26.5 I*lI'01 and 1-11-0, i.nchea, and Ill-Il-C, 24.5 34.5 inches, III-II,Or It is believed long. lis g f from r o m a maamale, l e n l l l - I111-Il-I, l - 1 ' ? 5 ' S l n c25.5 } : e a 1inches on6"It1 sb€1lc?€d nothing that t h a t tthese h c a g f lfish s h n amay y h , a vhave o b e abeen n h y b rhybrids, l c a , a l t h o ualthough ghnothle5 e t r i k . atrikapart' ihea apart. t them to 161i apFaafanca &o sxtersal. appearaneg 1mg wasnotod notedlnin their their external iag was Sieelhoad Steelhoad Coserclal Comparaon S-ort and asd Connercal Compariaon efofS'rort sport The T h e cconclusions o n c l u g l o n e cdrawn l r r y s : rfrom f r c i n aac comparison o n p a r l e c n o fof spo r t a n dand c c mcornunchanged' report remain :'ercsin unchanged. 1943 report the 1942 in the xrcial st€eLbsed in merci.al steeThead Tl:g The A smaller smallar ycungtr fleh. take smaller fish, A a::d younger e:nal1er and tends to taka sport ftehl*g tends eport fishing spad previously. proportion p r o p o r t l o n o fof t h othe e p osport r t f t s h fish h a v shave apa*r*edpreviousS.y"Bolh Both aunbor dieproportionate number tek: aa disproportionate flshera,En take sport and corerclal coercial fishermen sport and largor €Ten larger sneven tEkse an anglor takes sport angler t}:e sport of females. Ii{F',tsver, Hover, the of fsmaleg. the In all the In all the nets. nete' than ihe (i.s6i.o) than proportion of females fersales (1's6r1'0) proportion of tho from the ouffors' sport fisl:ery fishery suffers, judged Judgad from the sport above comparisons ccrnparlsoas the ebova standpoint of conservation, ccnse:irEtlo:: n etaadPolnt of valuatlon' cante valuation, of dollars 8nd cente dollara and vlowpoi"nt of tha viewpoint Ho'ver, fron the l{ow8vsr, from ccrcial the is t h e ssport p o r t f l afishery herylsun d oundoubtedly u b t e d l y e u p asuperior r l c r t o t h e cto o r rthe r.ngrcla}i 3D- 1ocal by local on by carried on isls carried glll-mtting tha gill-tetting placa, the the first place, Sport anglers, anglare' Sport tcvolvad' ls travelllag Yory litt1e little travelling i involved. residents. Very rogldente. tleoir reaoh their to reach :resey to deal ofofmoney 8pend aa groet great deal hand, eponC other hand, on tha the other oa Ia the flrst In reetaura:rtg' and reataurante hotcls and !n hotels as in as wg11 favorite well as streans, as favorLte streams, Ths The cotrelderabla spando considerable algo spends aagler also tho angler but the cheap, but not cheap, ata not nets nEts are lnis inar:gler ls tho angler by the takan by flgh taken cost trxr per fish fi:e oost sums for tackls, tackle. The slrlts for ',!0"69|ln WaahGrgan River Rtver in the Green on the th d on ls considered consldered that creased nhsn vthenttit is oreassd ingtoxi, l n g t o n , dduring u r t n g t hthe o l g 1940 4 0 s 3 darid 1 9 4 1941 1 e t g a lateelhead h e a d g 9 a 8 seasons, o 1 u t , 6 , 9 35,938 8 TtllIn TIllIn per angler. a:igler' fish per 0'1? flsh or 0.17 ouly 994 anglers 994ftoh, fish, or tsr-,k only aaglere took perlod from froo the period Durlng ttra better. During Cld bsttsr' ainookCcunty Countyti:'a theangler angler did anook checkad anglors checked 636 anglers 1942t the 626 91, 1942, November 23,1941 1q41toto }u'arch laroh 31, Novonber 25, por (O.ga) fiah ftsh per cnethird (0,32) abogt cna-thlrd or about 202 staslhead, had staelhead, or taksn 202 had taken angler, aagS.er n Recore:rCstio:rs Whatever T ' | h g t g ? s r t h the g f a t efate o f t h gof c othe w r c l oorcal . a l f l s h a r fishery - v f o r s t o efor l h asteelhead. ad. improve in it l t wwi-U l l l b o dbe e g ldesirable r g b l e t o l n g to rova t h o a n g t rthe ' t l g rangling o g u l a t l oregulations nela d c'32 cf :h more than the the 0.32 :sore thaa oat th raay cat angl-armay avsrag€ angler order the average that the order that rocommndaTherefore, the reconendaThsreforo' ti:e nenttoned' previouzly mentioned. atoclhaad provlously a steelhead hers' repeatad here ti-one made lnin 1942 1942 are are repeated tloae nada the bag bag fhay are, They are, thet that the reduced from 3 to 2 fish per day llimit l . n l t f ofor r a d adult u j ' t E t gateelhead e ] . i r e a d b ebe rgducadfrcreSto2fiohparday ltnltati.ong that limitatIons g€ak or possession)l that per week (and (and fron fromg9 toto 66 per or in in possession); cpeairg date date for for the opening and, that that the a::g!'lng3 and, on boat boat angling; be placed plaoed on be perslt a a to permit ay 15 15 (( to advanced to to ilay be advanced angling for 6-inch trout be 6-lnch trout apgllng for ocaan)' tlie ocean). to the steglhead to young steelhead of young greater grsat€r escapement ee capo:c*nt of *7- 1. ?ebLs 1. Table 3.943 Febn:ary, 1942 Jesuary" February, Steolha ad, Jaivary, Coms'srplal SteeThead, Age-1ngth, Ags-l$ngth, Coxnercia1 of aimuli an:rull No. $o. of TTT III ff IV V WT VI VII VII Pcrostrt TotaLg Percent Totala IX IX YtrII VIII leagth F'ork lenigth Fork lnohec Inches 22-28.9 22-23.9 5 I e 5 cl.l, .71 ?t 1 '!R a 3 77 77 11o 110 174 arz 62 1 ,G tt 272 ia 39* 98 58 a 8 2 2 181 18]. 28, 26. !{o 39 4s 43 1? 1? 1 104 104 LDr 15. 32-88.9 32-33.9 2 2 23 23 !J 14 1 1 3 4.t5.9 34.35.9 5 L5) 12 3€.3?.9 36.31.9 1* 9 4.25.9 24.25.9 !1 11 48 48 9€=E?*oB 26-27-.9 €t 35 28-29.9 28-29.9 17 gO-Sl.ng 30-31,9 A {! 4 Totals 67 n?n 370 Prroant Percent IU 10 G? 53 tn' Ày. Fo length leagth in. Ac. F. * 38 inches tnchog * ao a 1 I 2 1 411 41 6" 2 21t 3o 3, : I 91q 213 s0 30 gg.5 e ? ' 6 29.5 e ? n L 27.6 27.]. 44 6 6 6 rUi .85 go*4 30,4 ttq 31.6 a 11 I .:.4 .14 I a .14 .14 34.0 32.5 $ 2 ' 5 34.0 ,?'o ) .La[ .14 *8- I ' Table Tabla 2. of .Ago Average Length Age GrouPa Gro1p3 Le:gth o±' Average Ia In Tsar Year Arrn1i Aanu1l stream etreaB rutu3* Annuli oc€&l ocean No provlouaprevioue Nc ly ly apawned oparnod ?otal. Total PsraE* Percent F ' L. Lo Ày. A v . F. lnshas inches 10. 10. 27.L 27.1 E 3 f I TT II 1 Ary 67 4 4 I III IIJ 14 14 4-& 44 6u 6. 80"8 30.8 II II ? T II 12 u 326a 3?Ss 47, 47u 27.2 27.2 5 .60 o60 !? 32.1 I15 11Sb t€ 16. 304 60n4 f f 1Y IV. 5R L! II T YT III AQ 87 lII III TT II 6 vo93° 13. 18' 27'.& 27.4 af 1 1 x1 .14 $?.6 32.5 II IX .lv IV 22 22 22 22 3 nl 3.1 3 1.4 31.4 Ltr III ?TT III Z .? 2.7 29 29.5 "S 5 6 ?f I tI I 8 VY 12 "1,4 "1. JJ ii 2 2 2 .28 n?8 A ?ng 27,8 TT II V 2 a 2 .28 66o in a .,g sO 32.8 ltt III IV ,iv 4t .A = 4 ,5 31.4 s:.,4& 8 I II 1l VI 1 1. .14 g2,5 32,5 I9 III lrI ?'T .t .L 1 ,l .14 .I! 34.0 E4aO lye xYe 7 a. 8o b. b. o. Or d. d,. e. 6r VI ! nr'l L Str€a$' are TtriiIidawater. I srg ttdewter. rre I I atreai, stres4' I: tldeg'&ter. 7 ? a"s tide-vater. 1 1Itres, I 'b{4sryater. Er€ lXgtfsg$' 8 I are t eter, tdwtcr. I1 tj.do flstrsss, 4 are a.rs IItrearn, I tlCspstEr' areII:strqeYn' I ttdwzter. *2 areI1Itrea, EE" ?cg' "1& -9* 3. Táb1 $" 3eb1s Melss S?atralng' Maies PreviouB Spwii, Ag8 .{ga an1 end Pravlcus In Ia Year Tcar s $ Aiuli Anaul"l s4rewn ltrsar[ ? 1 .rniu1 A:::tu15. ooen oD4t]3 TT II Ti tlws pated spewaed Number Sirs,bcr uot No 1lo. nct opamsd, epawrred 0 4]. II 4 { 1 I III TI1 I 12 1? 0 0 't TT II ?T 11 1 3 3 2 1 146 t&8 0 18 1 I 5 T ? II ?Y? III TT? III fl 33 !x 13 0 1 5 6 t 2 1 1 11 g8 33 0 '!I 6 TY II T TT 111 T1 I?S rj'l IV l",t -L 111 1 1 2 & 2 1 0 1 1 z2 2 "IT II 0 T] 4 2 ? , 4+ ?otal 809 PErcsnt of, ee3.asspawrod, 15 g6E 4 ?ab1a Tab1 4" Faraalaa Age and Age id Prarrlaus Prviou SBaw:r{ngn Spa1ng, Fea1s8 Annull Armuli rtroa:a strean Ag.ul.l Annuli oesgn ocean I 8 t 1 T T 11 U 0 * 4 ? 1 r.ll III n 0 In In Tcar Year' li.rnas Tini epams* 'l 1 ?T II 1. $rcabol Lcnbex So. not aot No, epamsd paed apamod, spaned 26 25 tr8 18 IC n 0 g 2 1 1 -, Y a ?-{t 11 I 0 I 2 a 4. I 2o ?? 11 I8 ??? I:I III C 1 $s 50 2 1* 1 III III ?T II n 0 ? I t7 V 3 1I ? ? II IV IV 1a 3 16 ; TT? III T?T Ill t1 C 1 2lL 8 R 8 f 8 1 I11 I V {r 2 3 1 I 1 III III rY IV I 2 o 3 I! E 2 1 1 II T: YI VI t 4 'rinr1 { 3 a a .Li.TD fotaL Tctal $e0 40 4a cf ferna1e 9ene.:.esepwrnedn Percent apawr.d, 28 Percaat of *1 etre, I tdawatr 30 30 49 t .2 1S8 188 gdB' tl qtr ll! 9 6' T a b l e 3. Table A + o at a* Ma'LurltY Sexual Maturity P,oaoh!.ng Sexual Age Reaching ..bv*v In In Ygar Year .tur::atl.i Arnuli strsgm stream Alr::'l { Anuli €cge]l oce 2 2 T I I I I 8 ? 1 TT II 4 I 1 TTI III $ 3 II II ? I *4. TT IT TT :: 3E at II 11r III t 4 a.L.t III 6 5 6 6 I 6 Susaborof of Thmber f3.sh fish t 2 8e 82 FErcod Perc .88 .28 11 11.88 "66 t! 33 4.70 4.70 6 A g:s 24 3 r43 3.42 6A 52 ?n41 7.41 I I 1n 10 1,d&3 1.42 T TT III TT 11 99 14.O9 14.09 lr.L III TTT III 8 1o14 1.14 3 II3 2t .?8 1T+ IV ffi 702 *i33 stream, *111 strsaa, I tLdewatar