sS o-/-/Sr7 op. D.. °.7 o Summary of Reports . . . -7_ 6 1976 Sheep and Wp61' Special Report 457 May 1976 Agricultural Experiment Station Oregon State University, Corvallis CONTENTS 1. CHEMICAL CURING OF SPECIAL USE PASTURES IN WESTERN OREGON 1 Glenn Savelle, David Tart, and William Hohenboken 2. WETHER REPORT II - Summary of 1975-76 OSU Sheep Breeding Research William D. Hohenboken 3. 12 RECYCLING ANIMAL WASTES D. T. ToreZZ 4. 20 FORAGE CROP BREEDING AND GENETICS Rod V. Frakes 5. 40 CURRENT STATUS OF RESEARCH ON FOOT ROT IN SHEEP John A. Schmitz 6. 42 PERFORMANCE DATA FROM A ROTATIONAL CROSSBREEDING PROGRAM Martin Vavra, W. D. Hohenboken, R. L. Phillips and M. M. Wing 46 CONTRIBUTORS Dr. Rod V. Frakes is a Professor of Plant Breeding at Oregon State University, Corvallis, Oregon. Dr. William D. Hohenboken is an Assistant Professor of Animal Science, and Acting Director of the Genetics Institute at Oregon State University, Corvallis, Oregon. Dr. Ralph L. Phillips is an Assistant Professor of Animal Science, Union, Oregon. Mr. Glenn D. Savelle is an Assistant Professor of Rangeland Resources at Oregon State University, Corvallis, Oregon. Dr. John A. Schmitz is an Associate Professor of Veterinary Medicine at Oregon State University, Corvallis, Oregon. Mr. David Tart is a Graduate Research Assistant at Oregon State University, Corvallis. Dr. Don T. Torell is an Animal Husbandry Range Nutritionist and Livestock Specialist, Hopland Field Station, University of California, Hopland, California. Dr. Martin Vavra is an Assistant Professor, Eastern Oregon Agricultural Research Center, Union, Oregon. Mr. Mark Wing is a Research Technician, Eastern Oregon Agricultural Research Center, Union, Oregon. CHEMICAL CURING OF SPECIAL USE PASTURES IN WESTERN OREGON Glenn Savelle, David Tart, and William Hohenboken CHEMICAL CURING WITH PARAQUAT 1/ A problem familiar to all livestock producers in Marine Temperate and Mediterranean environments is the feast or famine situation of having an overabundance of high quality native pasture for 4 or 5 months of the year and inadequate amounts of marginal or submarginal quality pasture for the other 7 months. Two special problems that sheep producers face are: 1) finishing lambs at a time when the growing season is ending and pasture quality is deteriorating and, 2) entering the reproductive season with native pasture that is approaching its lowest ebb in quality and quantity. Paraquat is an extremely fast acting herbicide which probably moves into leaves and stems through the minute orifices (stomata) that are normally the sites where gaseous interchange of oxygen and carbon dioxide is effected between the atmosphere and intercellular spaces. The apparent centers of herbicidal activity are the small granules (chloroplasts), within cells, which contain the green coloring matter of plants. Here, paraquat is able to substitute for an iron protein compound involved in photosynthesis. This proves lethal for the cell and no further redistribution or metabolic breakdown of nutrients or energy can occur. Only catalytic amounts of the herbicide are needed to kill most chlorophyllous tissue. Complete desiccation or drying out of the herbage generally occurs within 3 to 4 days. If the nutritional value of the species which dominate a pasture is high at the time of chemical application, the result should be a preserved standing forage of high quality. In the past few years several animal and range science groups at Oregon State University have been examining both these problems from different perspectives. This paper will report on progress in one of these areas - reproductive efficiency and the flushing of ewes, but will direct primary attention to the approach being used to obtain high quality special use pasture - chemical curing. It is our intent to provide a better understanding of: what chemical curing is, what has already been done in other geographical areas and, what the potentials and limitations are for this method of pasture conservation in western Oregon. At moderate and high concentrations paraquat is non-selective, that is, grasses and forbs and most herbaceous species in general are affected similarly. Some translocation of the 1/ Paraquat has not yet been cleared by the U.S. Environmental Protection Agency for the chemical curing of grasses. 1 chemical may occur in a few species but the principal herbicidal action occurs near the point of contact. For this reason perennials are usually defoliated but not killed. This also means that in very dense, thick, meadow-like conditions only the top part of the vegetation may be desiccated while the lower part remains green for a period of time. The use of paraquat to cure pasture vegetation in place, developed indirectly from research that USDA Agricultural Research Service scientists Don Hyder and Forrest Sneva were doing at the Squaw Butte Experiment Station. This work was directed towards improving the summer quality of crested wheatgrass pasture by developing a grazing system timed to remove the growing point of this grass and thereby help arrest the redistribution of nutrients. This led them to look also for a chemical that would accomplish the same end by suppressing growing point dominance. Paraquat was one of the chemicals tested in 1961 and although it did not accomplish the desired result, the curing qualities of the chemical were noted and Sneva began a series of experiments and grazing trials that are still active. activity on the chemical curing of pastures. A half dozen chemicals other than paraquat have been tested. The results are usually a little more erratic and not quite as satisfactory as with paraquat. CURRENT STATE OF KNOWLEDGE The potentials and limitations of chemical curing, with any of the suitable herbicides, will be better understood if we take a brief look at some of the effects on vegetation and animal performance thus far demonstrated for paraquat. Chemical composition and digestibility. A naturally matured pasture can lose 50 percent or more of its crude protein value by the end of summer, while the same pasture when chemically cured has the potential to retain 80 to 90 percent of that value. Other nutrients are also influenced. Phosphorus concentrations are always higher in paraquat treated forage. Potassium and sulfur are usually higher, while magnesium concentrations appear unchanged by treatment. Calcium, crude fiber, and lignin are variously affected. Cellulose is usually lower in sprayed forage. In 1966 scientists at the University of California, Davis, who had worked with paraquat as a weed control herbicide, began looking at its curative properties on annual rangeland. Simultaneously reports on similar types of experiments in Australia started appearing in the literature. During the past 10 years these have been the three main geographical centers of research 2 The initial nutrient value of chemically treated pasture, and the relative difference in nutrient status of sprayed and unsprayed forage at the end of summer depends on the time of treatment, and probably on some yet unidentified factors. Australian studies show that within the 3-week span from grass head emergence until pollination, protein retention can drop to 20 percent of maximum value. Results of less magnitude have been obtained in eastern Oregon and California. are improved or unaffected. In no case has any plant constituent except lignin been found to be less digestible after treatment. The fact that both live and dried plant material is susceptable to leaching has been known for some time, but the amount and pattern of precipitation that will cause significant leaching loses and the relative leachability of treated vs. untreated forage is unclear. One laboratory study of the leaching of cut-up and groundup herbage after immersion in water concluded that all nutrients except nitrogen are equally susceptable to leaching, whether chemically treated or not, but that some nutrients particularly phosphorus and potassium - may be leached at a faster rate if paraquat treated. It cannot be assumed that these same relationships will hold under field conditions. Botanical composition and yield. There is obviously going to be a reduction in total dry matter yield when pasture growth conditions continue to be favorable and biomass accumulates (in unsprayed pasture) beyond the dates of paraquat application. The magnitude of the reduction depends on the length of the growing season. In the Mediterranean environment of Western Australia, a dry spring and a consequently shortened growing season resulted in yield reduction of no more than 2 to 15 percent. In a wet spring this loss may increase to 40 percent. Eighty-five percent of all pasture experiments reported in the literature show some reduction in total yield. A complicating factor is the effect precipitation has on regrowth. In several instances regrowth has been greater on treated pasture, with a net result that the total mineral status of treated pasture and the difference between it and unsprayed pasture was greater than would have been expected without precipitation and greater than it had been in dry summers. A less obvious loss of biomass may occur because of the susceptibility of paraquat treated forage to fungus attack. Accelerated decomposition can be triggered by a combination of moist conditions and high temperature. This may have occurred in several studies, but the relative importance of such dry matter disappearance has never been separated from yield reductions due to continued pasture growth or from the possibility of accelerated physical shattering of chemically treated herbage. Less has been done with digestibility. All in vitro and in vivo trials have demonstrated the greater apparent crude protein digestibility of paraquat treated herbage, but there is lack of agreement as to whether the digestibility of dry matter, water soluable carbohydrates, gross energy, crude fiber, or cellulose In nearly every study in California and Australia, where subterraneum clover was substantial component of a paraquat sprayed pasture, there was a carry-over effect the second year in the form of increased clover dominance. This increase has been as much as 100 3 are most likely to present problems and which, therefore, should receive research priority. to 300 percent by weight. The same shift in species composition has also occurred in the first year when pastures were sprayed in mid-winter. There is almost certain to be a trade off of quantity for quality. Infrequently there may be an increase in nutrient yield per acre even when total dry matter yield is reduced. Generally, however, one must expect to carry fewer animals per pasture. This would probably not make chemical curing a viable management practice in situations where animal maintenance through the summer is the only requirement. Livestock performance. Reports from grazing trials are few and all have been limited to weaner lambs and yearling cattle with results rather evenly divided. Trials in Australia were not able to demonstrate absolute weight gains for animals on paraquat sprayed pasture, although lambs did maintain weight during summers that they were losing on untreated forage. Identifying the proper time to apply a chemical like paraquat is, for a number of reasons, one of the two most important technical. problems in western Oregon. First, the odds that a complex mixture of annual and perennial grasses will be found on most foothill range are good. Such a situation potentially increases the spread between calendar dates when individual species attain the same stage of phenological development. As this time interval widens, the probability increases that there will be one or more important vegetation components at peak nutritional quality when other equally important components have either not reached that stage or are already declining in value. Animal selectivity can also cause problems in a polyculture. If the preferred species coincide with those of highest nutritional value, the overall quality of the pasture can decline during the grazing period. A third point to consider is how proper timing of chemical application can be used to influence botanical composition, especially the establishment or restoration of clover dominance. An inherent danger is that annual On the other hand, yearling heifers grazing paraquat treated crested wheatgrass in eastern Oregon have averaged a 1/2 pound per day greater gain than on untreated pasture. In two of the three grazing trials, conducted on annual rangeland in California, lambs gained about 0.15 pound per day on treated summer pasture, which is double what they gained on uncured pasture. The Australians are the only ones to have looked at wool production. In several studies they have found a 20 to 30 percent higher grease wool weight from animals on paraquat pasture. CHEMICAL CURING IN WESTERN OREGON This review of past studies leaves unanswered many questions as to the relative advantages and limitations of chemical curing. But it does give insight into some of the more probable results to expect if this pasture management practice were employed in western Oregon. It also directs attention to those aspects of chemical curing that 4 weedy forbs may increase instead of clover. Lastly, the relationship of timing to subsequent regrowth is an unknown factor but possibly an important one. The second important problem is the amount or pattern of precipitation that can be tolerated in late May or June - or any other summer month - before excessive leaching of the vegetation occurs. Some investigators in Australia suggest that there is too much summer moisture in an area like western Oregon to make chemical curing of forage feasible, at least with the chemical products presently on the market. A closely related aspect of the moisture problem is the increased susceptability of herbage to fungus attack under conditions of high relative humidity. It was with this background knowledge that we decided to put to the test the practicality of chemically curing standing forage, given the climatic conditions that exist in western Oregon. FLUSHING TRIALS During the 1974-75 grazing season a combination flushing/creep feeding study was started by Dr. Hohenboken and his students. Flushing consisted of three treatments: two in drylot and one on hill pasture. The preliminary results of this segment were reported last Sheep Day. The complete results, together with an economic evaluation for the first year, are the subject of a Master's thesis and are presented elsewhere (Hohenboken and Taiwo, 1976). For the 1975-76 grazing season, a paraquatcured dryland pasture was added as a fourth treatment. The following is a summary of the first year flushing treatment design and results together with the modified design and partial results for the current season. On September 13, 1974, one hundred and two mature blackface crossbred ewes were randomly divided into three treatment groups. Ewes in group II were maintained, throughout the flushing period, in drylot where they were fed a daily ration of one pound alfalfa hay and one-half pound barley per head. Grass clover hay was provided "free choice". The group III ewes received the same treatment as group II until bred. They were then turned out to dryland hill pasture. Group I, which served as the control group, was kept on hill pasture for the entire flushing period without supplemental feed. Breeding started with the introduction of the rams to all groups on September 30. The flushing treatment was ended 17 days later at which time group II and all unbred group III ewes were placed on dry pasture with group I controls. Rams were left with the pooled ewes, for an additional 17 days, until November 5, 1974. From this date until lambing time all ewes were maintained together on hill pasture. The control group (I) exhibited a gradual increase in weight a subsequent leveling off, and then a slicAt drop in weight towards the end of the breeding season. Both drylot groups initially lost weight, regained it, and then leveled off. Despite this variation in ewe weight through time, overall after being bred. Group I, the dryland hill pasture ewes, gained about 0.15 pounds per day for the entire period. The paraquat ewes (group IV) gained 0.25 pounds a day until breeding but then lost weight. If we accept a figure of from 0.2 to 0.3 pounds per day gain, the range most frequently appearing in the literature, as signifying a flushing response, we can say that the drylot ewes were flushed; the paraquat ewes almost flushed; and the dryland pasture ewes were not flushed. weight performance for all groups gave little evidence that a flushing response had been achieved. We don't have an explicit explanation for this. It could have been due, in part, to the already good condition of the animals at the start of the experiment (weights ranged between 154 and 161 pounds). An initial miscalculation resulted in a marginally inadequate feed ration, according to NRC standards Dusty drylot conditions plus excessive handling for blood samples may also have been a contributing factor. The most obvious question, posed by the past season's results, is why the ewes began to exhibit a flushing response and then lost weight after the start of mating. The appearance of the treated pasture after three weeks of grazing suggested a lack of total available dry matter. This has occurred in some of the Australian work with weaner lambs and steers on summer pasture. However, there was close to 1000 pounds per acre of total biomass on the pasture when it was sampled September 17. If we discount 10 days of possible regrowth, and allow for reduced accessibility of some of the forage because of trampling, there should still have been 400 to 500 pounds per acre of readily available forage at the end of the flushing trial. The nutrient quality of the pasture, at the time of animal use, has yet to be determined. This, coupled with an initial selective curing effect combined with selective grazing, could have become an important factor towards the end of the flushing period. Lambing percentages are shown in Table 1. The differences between groups suggest that drylot treatments had some stimulating effect on ovulation rate, but that continued drylot feeding increased embryonic mortality. A similar experimental design was used during the 1975 breeding season except for the addition of the paraquat-cured pasture treatment and a fourth group of 33 ewes. Livestock procedures, handling facilities, and feed ration were improved. The entire schedule was also shifted forward 5 weeks so that flushing was started August 4 and rams were introduced August 22. As in the previous year, rams were left with the pooled ewes on dryland pasture for an additional 17 days beyond the end of the flushing period on September 8. Weight changes for each group are shown in Figure I. Drylot ewes gained about 0.4 pounds per day until the start of breeding. Group II ewes, which remained on drylot throughout flushing, continued to gain at a reduced rate while group III ewes barely maintained weight when they were turned onto dryland pasture There was a total of 1.5 inches of precipitation during a 6 Table 1. EFI4 ECTS OF FLUSHING MANAGEMENT ON REPRODUCTIVE PERFORMANCE OF EIES Year I Treatment 2 III II Number of Ewes 1 34 34 34 Fertility (%) 97 97 97 Multiple Birth (%) 65 59 77 167 161 182 Number of Ewes 1 31 29 32 30 Fertility (%) 87 90 91 80 Multiple Birth 44 70 38 46 Lamb Crop (-%) 1 148 174 145 154 Measurement 1974 Lamb Crop (%) 1 1975 1 Based on number of ewes surviving after lambing 2 I Dryland pasture (control) II Drylot until end of flushing III Drylot until bred IV Paraquat pasture IV Figure 1 WEIGHT CHANGE DURING FLUSHING PER TREATMENT GROUP ■••■ 1 2— until ----mibmismmwmaDrylot bred 41■11~111. I AP 101:00.741w AI ■se• AI 41: 411 401. 1 0— I," .14P AI ' III∎ 4■41\ Nik -4■ t • 40`,■ IWO* AO■ ■ 8— 0 CI_ 6— 4— 2— Drylot throughout flushing lo lb' 0 4 10" AteiP 401. AP' 40, 40110•400,110, 4, 1000• Asill° ol■•• INN& /111 11111141k I" 4:1.11' August 4 Begin flushing Dryland pasture August 22 Begin breeding 8 Paraquat pasture 17 day period in late June and early July. On two occasions within this interval, 0.4 inches fell in a 24 hour period. Although we do not feel at this time that this was a critical factor, shortly after the period of precipitation forage blackened considerably because of the presence of fungi. The chemical analysis will show if there was appreciable leaching. CONCLUSION Animal response to paraquatcured pasture, during this past years' flushing trial was not as good as we had hoped. But, it was encouraging enough to warrent continued research. We cannot expect maximum animal response until we have obtained the greatest possible vegetation response to the curing process. This will not be A complete economic analysis will accomplished until we learn be made of flushing and creep more about which species are feeding after this years crop of the key to the timing of chemical lambs is weaned. Table II gives application, which give a comparison of the first year's maximum regrowth after being extra-flushing costs for drylot chemically cured, and which feed, which will be a little are the least susceptible to higher this year, and the current leaching and fungus attack cost of chemically curing pasture • (if this latter factor should None of these figures take into prove important). Finally we account the extra labor cost for need a better idea of the drylot handling and feeding conditions and the threshold or for spraying the herbicide. point where precipitation becomes a factor. TABLE II FLUSHING COSTS FOR FEED AND PARAQUAT PER EWE BRED Group Treatment II Drylot 34 days The authors gratefully acknowledge the loan of the sheep by the Superior Packing Company of Ellensburg, Washington, and the donation of Paraquat by the Chevron Chemical CompanY that were used in these studies. Cost $ 3.72 III Drylot until bred 2.76 IV Paraquat pasture 1.41 The paraquat treatment was approximately one-third the cost of flushing in drylot for the entire 34 days (group II), and one-half the cost of flushing ewes until bred (group III). The cost for the paraquat treatment could probably be reduced to less than $1.00. An application rate of 0.5 pounds per acre was used and this is over twice the rate that several studies have shown to be necessary. 9 References Agbakoba, S.C.O., and J.R. Goodin. 1967. Effect of paraquat on the nitrogen content and regrowth of coastal bermudagrass (Cynodon dactylon (L.) Pers.). Agron,j. 59M:605-607 Akhavein, A.A., and D.L. Linscott. 1968. The dipyridylium herbicides, paraquat and diquat. Pages 29-145 in Francis A. Gunter, ed. Residue reviews. S p ringer-Verlag, New York. Arnold, G.W., D.W. Barrett, P. Lapins, and J. Whitehead. 1970. Some effects of paraquat on yield of dry matter and nutrients, and on sheep production from annual pastures. Pages 866-869 in Proc. 11th Intern. Grassi. Cong., Australia. Arnold, G.y., and D.W. Barrett. 1974. The effect of paraquat desiccation in spring on plant and animal production from annual pastures. Aust. J. EXD. Ag ric. Anim. Husb. 14(1): 23-32. Barrett, D.W., H.A. Campbell, and G.W. Arnold. 1973. The effects of pre-maturity desiccation with paraquat and leaching on the quality of dry Wimmera ryegrass. Aust. J. Exp. Agric. Anim. Husb. 13(63):411-417. Barrett, D.W., G.W. Arnold, and N.A. Campbell. 1973. Effects of tine and rate of paraquat application on yield and botanical composition of annual pastures containing subterranean clover in a Mediterranean climate. Aust. J. Exp. Agric. Anim. Husb. 13(63):55b-562. Freyman, S. 1970. Chemical curing of pinegrass with atrazine and paraquat. Can. J. Plant Sci. 50:195-197. Hohenboken, William, and Bolaji Taiwo. 197b. Drylot breeding, creep feeding in we s tern Oregon. Oregon Farmer-Stockman. 99(3):40-43• Kay, Burgess L. 1968. Effects of paraquat on yield and composition of a subclover-hardinggrass pasture. Weeds 16(1):66-68. Kay, Burgess L., and Donald T. Torell. 1970. Curing standing range forage with herbicides. J. Range. Manage. 23(1):34-41• Kay, Burgess L. 1970. Paraquat curing of seeded dryland pasture species. J. Range. Mana ge. 23(b):40(-411. Pullman, A.L., and W.G. Allden. 1Q71. Chemical curing of annual pastures in southern Australia for beef cattle and sheep. Aust. J. Agric. Res. 22( ):01-413. 1 0 Raleigh, R.J., F.A. Sneva, and H. A. Turner. 1968. Chemical curing range forage for fall grazing. Pages 283-287 in Proc. W. Sec. Am. Soc. Ani. Sci. 19. Romberg, Barbara, G.R. Pearce, and D.E. Tribe. 19b9. The effect of chemical curing with paraquat on the intake and digestibility of Phalaris pasture. Aust. J. Exp. Agric. Anim. Husb. 9(1):71-73. Sneva, Forrest A. 1967. Chemical curing of raage grasses with paraquat. J. Range. Manage. 20(6):389-394. Sneva, Forrest A. 1970. Paraquat - effects of growing season applications. J. Range. Manage. 23(6):451-452. Sneva, Forrest A,, R.J. Raleigh, andY!_.A. Turner. 1973. Paraquat cured herbage for late season grazing. J. Ani. Sci. 36(1): 107-113. Sneva, Forrest A. 1973. Nitrogen and paraquat saves range forage for fall grazing. J. Range. Manage. 26(4):294-295. Taiwo, Bolaji. 1975. Biological and economic effects of flushing and creep feeding in sheep. M.S. Thesis. Oregon State University, Corvallis. 05 pp. Wallace, Joe D., P.A. Sneva, R.J. Raleigh, and C.B. Rumburg. 1966. Digestibility of chemically cured range forage. Pages 385-390 in Proc. V. Sec. Am. Soc. Ani. Sci 17. 11 WETHER REPORT II Summary of 1975-76 OSU Sheep Breeding Research William D. Hohenboken As I begin this report, the month of March shows every indication that it intends to go out like a lion. We are in the path of a storm front featuring strong winds and driving rain. It's a good day for writing; it would be a poor day for working lambs. area at the irrigated pasture unit, installing an underground water system to pastures near the Sheep Barn, overseeding clover into several irrigated and dryland pastures and using hay making, wheat farming, pasture clipping and heavy summer grazing as parts of our pasture improvement program. Much of the maintenance and improvement work was completed using fence wire, lumber and materials from unused facilities. Our main problems of weeds, inadequate fencing and poor production from some pastures are gradually coming under control. In this OSU Wether Report, Second Edition, I will first comment on our animal, land and personnel resources. Then progress reports of four breeding projects will be presented. Finally, there will be a brief comment on work currently in progress. Resources Personnel. Bob Klinger and Lloyd Westcott continue as full-time OSU shepherds. In May of 1975, Bob was honored as the OSU Oregon State Employee's Association employee of the month. Bolaji Taiwo, graduate student in livestock management, completed his master's degree and returned to his native Nigeria in December, 1975. He had conducted the first year of the mating and lactation feeding management project for his research involvement. This year, David Tart, graduate student in Range, is continuing that work under the supervision of Mr. Glenn Savelle. Progress on the project is reported elsewhere in this bulletin. Miss Rose Mary Cedillo analyzed the lamb and wool production of crossbred ewe lambs for her graduate research. Joe Ngam, a master's student from the Cameroun, is working on the importance and the inheritance of appetite in ruminants for his research. Results from both of their experiments are presented later in this article. Animal. Purebred Hampshire and Suffolk flocks number 36 and 59 ewes, respectively, including replacements. These sheep support our undergraduate teaching program. Rams from both flocks breed over 400 ewes involved in Animal Science and Rangeland Resources research. The purebreds also support shortterm management, behavior, genetics and nutrition research. Our experiment on the effects of feeding during mating and lactation involves 123 grade blackface ewes on loan from Superior Packing Company and C2L, Inc. of Ellensburg, Washington. After this year, they will be replaced by a group of their daughters who will be used in continuing management research. The major OSU breeding experiment has 390 ewes of eight crossbred groups split between hill pasture and irrigated valley management systems. RESEARCH PROGRESS REPORTS Land. This past year we have continued efforts -to maintain and improve our physical facilities. Projects have included: moving division fences and replacing perimeter fences at the Wilson Farm, constructed a shelter and work Breed effects on lamb carcass merit. 1973 male lambs from eight crossbred groups (Dorset, North Country Cheviot, Romney or Finnsheep rams mated to Suffolk or whiteface, Columbia-type 12 Yield grade five lambs would be seriously overfinished. range ewes) were raised to weaning in June under typical western Oregon hill pasture conditions. They were then trucked to the Columbia Basin Branch Experiment Station in Hermiston where they were sorted by weight into lots, sheared, drenched for internal parasites, implanted and placed on feed. The RALGRO ® implants were furnished by Commercial Solvents Corporation, Terra Haute, Indiana, and their support is gratefully acknowledged. After six weeks, lambs were topped out for slaughter at Superior Packing Company, Ellensburg, Washington. All remaining lambs were sent to slaughter five weeks later. OSU personnel collected data on carcass quality and quantity of 136 lambs. All groups averaged middle choice (11 on the numerical scale) for USDA quality grade. Thus, lambs of all the crossbred groups, when carried to normal market weight on a feedlot ration, had no difficulty in grading acceptably. Important differences did exist for fat thickness. Columbia-type crosses were fatter than Suffolk crosses, and Romneys and Finnsheep were fatter than Cheviots and Dorsets. Differences in kidney fat percentage were also significant. For this trait, the more wasty groups were the Columbia-type, the Dorset and the Finnsheep crosses. This is reflected in poorer cutability or yield grade for those groups. Suffolk and Cheviot crossbreds had superior cutability. Results are presented in table 1. All lambs were two breed crosses, so the figures represent average performance of all lambs of 50% inheritance from each sire and each dam breed. Carcass weight per day of age (WDA) is important economically because it is a good measure of growth rate. High WDA means fewer days in the feedlot to reach market weight or more pounds of lamb available at the end of a constant feeding period. Differences among the groups were not large, but Cheviot, Suffolk and Dorset crosses were superior to Romney, Finnsheep and Columbia-type crosses. Likewise, differences in dressing percentage were not large. The difference between the highest and lowest dressing groups was only 1.2%. Two more years of carcass data have been collected and are awaiting analysis. At this stage we have not found large differences in WDA, dressing percent or USDA quality grade. There are indications that differences exist in fatness and in cutability that favor Suffolk and Cheviot crossbreds, with Columbia-type and Finnsheep cross breds being less desirable. Breed effects on wool production. Wool production data from ewes of the same eight crossbred groups were presented in the 1975 Wether Report. This will be an update of that information, with records on 1974 ewes added to the analysis. The ewes were born in February of 1973 and 1974 and weaned and sheared in June and July, respectively, each year. They were grown out during the summer on irrigated pastures and were bred each year starting in early October. Each group thus lambed in February at about 12 months of age. Fleeces were sampled at the britch and at midside before shearing in March or April. Wool data represent about 81/2 months of growth for each group. Lamb carcasses can now be federally graded for quality, using the primechoice-good standards that most producers are familiar with, and for yield of salable lean meat from the carcass. Yield or cutability is estimated by the USDA yield grade system. 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The same ewes from the same eight crossbred groups in the same two management systems, as were discussed in the previous section, furnished these data. All ewes had been bred to Hampshire rams beginning in late September of each year. Results are presented in table 3. For staple length, Dorset-sired ewes were shortest with small differences among the other breeds. Finnsheep crosses had the longest 81/2 months growth. Within Dorset and Finnsheep crosses, ewes with Columbia breeding had longer fleeces than ewes with Suffolk breeding. Columbia-type and Romney crosses had a clear-cut advantage for grease fleece weight, and Finnsheep crosses were poorest for that trait. Differences in clean wool yield were small and not consistent among breed groups and years. The high yields for all groups probably reflect our "pre-scoured" wool from winter rains. Clean weight was estimated by multiplying grease fleece weight by clean wool yield. Averages for clean wool followed the same pattern as averages for grease wool weight. Fertility refers to the number of ewes lambing as a percentage of the number of ewes mated. The overall level of fertility achieved was not satisfactory. For breeding of ewe lambs to be economically feasible, at least twothirds of the potential replacements should conceive during a 40-50 day breeding season. Part of the reason that we did not achieve this level was that we saved a much higher replacement percentage (about 85%) and thus a higher percentage of smaller ewes than a commercial producer normally would save. This was done to build up numbers and to allow the experiment to be run in a shorter amount of time. Ewe lambs with Suffolk breeding were more highly fertile than ewes with Columbia-type breeding. Finnsheep crosses were most fertile, Romney crosses the least, with Dorset and Cheviot crosses intermediate. The groups didn't differ greatly in twinning rate except that Finnsheep cross ewes were 40-50% above the others. Higher reproduction is their main potential contribution to American commercial sheep production. Clean wool prices were assigned according to fiber diameter grade, staple length and discount for medullated fibers. Medullation (harsh-feeling wool caused by faulty structure within individual fibers) was most severe in Cheviot crosses and least severe in Finn crosses. It occurred with higher incidence in Suffolk than in Columbia crosses. Price per pound of clean wool favored Columbia-type over Suffolk crossbred ewes. It was lowest for ewes with Cheviot breeding. Finnsheep crosses scored surprisingly well--on a par with the Romney crosses. For total gross income from wool, Columbia-type crosses surpassed Suffolk crosses, and Romney crosses surpassed the other sire breeds. All crossbred ewes were mated to Hampshire rams, so all of the lambs were 50% Hampshire breeding and 25% of each of two other breeds. Differences in average lamb weaning weight among groups reflect breed differences in growth potential and in milk production of the crossbred mothers. For average weaning weight per lamb. Suffolk crosses were superior to Columbia-type crosses. Average weaning weight also 15 4 a) 00 0 CIZS U $.4 0 0 N a) r-1 r-1 •ct tk .100 0 i N4 00 Ce) 0 v-4 0 N• 0 00 •Cr ri 1-1 t4) CNI Cs.] C•4 M Cs] •1' et N- tr) O 00 N- N 00 (N1 00 0 r•-4 00 00 0 lf) fn cd ■-1 N $-1 -C) rn $-+ 4-1 0 Sa.1 0 0 a) -0 g g 03 $-1 0 a) al, 0., C.) O $-1 U g 4-) b4) 1--4 • H U 0) -zr rn r•-4 •J• 1•4 ••:t 00 •,1• •Zr "Zt Cr) Cs/ V) CN1 Ci) br) 0 0 0 I-- ∎0 if) V) •ct N. 01 00 ri -0 cd o4.-) cr) c-0) cd Oa a) .ri $-■ (-7 3 V) 4 Lf) N.0 $.4 O 4-1 1-4 0 .)4), o ri cd C ■0 00 v-4 •r oo 0 Crs N N4 N ■0 .0 N. .0 .0 N. 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They will probably still lamb with a higher twinning rate than other groups, but the difference will not be as large as it was for ewe lamb production. The product of fertility, twinning rate, survival and weaning weight gives total pounds of lamb weaned per ewe bred. Pricing those pounds at 394 apiece gives lamb income per ewe. Adding wool income from table 2 gives total income generated per ewe bred per crossbred group. Inheritance and importance of appetite. Last year in the Wether Report I described experiments to determine whether appetite was related to rate of gain or feed efficiency in lambs. If it were, it might be cheaper, easier or more accurate to performance test lambs for appetite than for efficiency. Suffolk cross ewes were superior to Columbia cross ewes for lamb production, for lamb income and (despite poorer wool income) for total income. They were also heavier the fall after their first lambs were weaned. The Suffolk has great potential for commercial crossing systems in Oregon. The breed should be exploited not only for growth rate and carcass merit, but for reproductive and maternal characteristics as well. Of the breeds evaluated in this experiment, it comes closest to all-purpose, all-weather specifications. It lacks only in wool production and wool value, and these deficiencies are more than compensated for by increased lamb production. Trials similar to those reported last year were conducted on purebred Hampshire and Suffolk replacement ewes and rams during the summer of 1975. During the three-week test period, lambs were confined to individual 4 x 12 foot pens. They were allowed one week to acclimate to test conditions and to a pelleted ad libitum diet. Six appetite trials followed at two-day intervals. For each trial, feed was taken away from the lambs at 4:00 pm and reintroduced at 8:00 am the following day. Consumption was recorded after 30 minutes, after 60 minutes and after 24 hours. Appetite was estimated as the proportion of 24 hour feed consumption eaten in 30 minutes (FC30) and in 60 minutes (FC60) and as the average daily feed consumption per unit of body weight (F/W). Ewe lambs with Finnsheep breeding surpassed ewe lambs with Dorset,. Cheviot and Romney breeding (in that order) for lamb production, lamb income and total gross income per ewe bred. The higher fertility and twinning rate of Finncross ewes was responsible for most of the difference. They held their own for wool income but had below average weaning weights. Romney x Columbia-type ewe lambs suffered from low fertility; only part of the deficiency was made up by their above average wool income. For fall ewe weight, Finncross ewes were slightly lighter than other crossbred groups. Differences among the other three breed groups for ewe weight were not large. Average values for FC 30 for rams and ewes respectively were 23 and 25%. Comparable FC 60 percentages were 28 and 31%. Thus, after an overnight fast, sheep will eat over one-quarter of their next 24 hour feed consumption the first hour after feeding. For ewes the percentage is slightly higher than it is From these data, the Finncross ewe 17 a) cu cd 0 4, —I 0 06 4-J H4 cd 0 0 4-1 0 • rI 00 (N C \I e--1 e--4 CA v•4 N (N 0 V) 0 (- \I 1-4 •C • N O) 0 N (N N 1-1 N •:1• t4) N N Cr) 00 1-4 ■-1 (N \0 V) (N r-1 r--.1 (N 0 ■-I r-i Lf) (N r--1 Lf) r--I Hi O 3 a) a) E U.D. 0 t.) 4 0 ga4 E $-1 0) E-4 E-1 0 0 4 cd U a 0 •1-i 3 a) 4-) -0 0) 4 a) $•4 $.• Mg a) -0 ri cd 0., 0 a) a) 3 3 O t)0 cd E •H 0 o > 4-) o a\c' 00 ••,:• r--I V) N H4 (N NI M re) V) N CA N •cl• in t.4.) 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An interesting observation so far is that Dorset-cross ewes seem more willing to approach and challenge (by foot stamping) a tethered dog. The relationship between this and their life expectancy is not yet known! We are also about to begin experiments to determine whether ewes of the eight crossbred groups harbor different loads of internal parasites. After these and other trials and tribulations, our 390 crossbreds would be justified in feeling more like guinea pigs than sheep. We hope it doesn't damage their mental health. for rams. Both FC 30 and FC 60 were quite repeatable from day to day. In other words, a lamb who started out above average was likely to stay above average for the remaining five appetite trials. Both sexes consumed pellets equal to 4% of their body weight per day. FC 30 and FC 60 were highly correlated to one another. Both were moderately negatively related to F/W, however indicating that the two traits are not the same biologically. For appetite to be a useful trait to measure on sheep, it should predict some economically important characteristic. FC 30 and FC 60 were not consistently related to the lambs' weaning weight, their weight at the start of the test or their rate of gain during the test. This is similar to results from the winter, 1974 trials. Thus, these traits do not look promising as useful future selection criteria. More extensive analyses will be necessary before they can be eliminated with certainty, however. F/W did have a consistent negative relationship with weaning weight. Lambs heavier at weaning tended to consume a smaller percentage of their body weight in feed than did lambs lighter at weaning. The full relevance and significance of that observation is not yet known. Feed, air and water are a lamb's only raw materials for meat and wool production. The amount of feed that he consumes is bound to be important in determining how much animal product he produces. We suspect that the rate and greediness with which the feed is consumed might also be important, but concrete results and conclusions have eluded us to this point. Mr. Ngam, the graduate student working on this project, should finish by fall of 1976. More definitive answers will be available at that time. Work in progress. The OSU cross19 RECYCLING ANIMAL WASTES D. T. Torell University of California, Hopland, California greatly exceeds the demand. Present ewe population is approximately 13,000,000. INTRODUCTION "Waste" is perhaps not the best choice of words to describe what I am going to discuss, as the dictionary defines "waste" as something that is "rejected as useless or worthless." Since anything with a potential use cannot be considered useless or worthless, a product which can be recycled cannot be considered waste. A better nomenclature might be the old-fashioned term, "by product," which is defined as a "secondary or incidental product." Since there are many by-products that can be recycled or fed as livestock feed and I haven't the time nor the inclination to cover all of these, I shall limit my presentation to "manure"— not an especially enticing title! So, we may as well stick with the misnomer, "wastes." This report will be further limited to the manure of laying hens, broilers, and feedlot cattle. The categories may be limited, but not so the manure; it is indeed plentiful. According to the Federal State Market News Service, there were 382,793,000 hens in the United States in 1974. If one hen excretes .07 pounds (air dry) of manure daily, and if one ewe's diet consists of 50% dry poultry waste, this is enough to feed 19 million ewes (Forsht et al. 1974). Likewise, the 2,992,844,000 broilers raised during 1974 in six fills per year would mean that there were approximately 500,000,000 broilers in existence each day. Their manure would weigh 17.5 million pounds (air dry), so if one-fourth of a ewe's diet were from broiler manure, one-fourth from the bedding, and one-half from an energy source (grains or pulps), there would be enough broiler litter to feed 25 million ewes. The amount of ewe feed that could be furnished by beef feedlot manure is astronomical. During the 1973-74 feeding year, 96,625,000 tons of feed were fed to cattle in the feedlot. If this were 65 percent digested, there would be 33,818,750 tons of dried manure or enough to feed 70 to 80 million ewes. So, the availability PROTEIN The chemical composition of broiler litter, cage manure, and feedlot manure depends upon several factors such as presence of litter, type of litter (shavings, peanut hulls, rice hulls, almond hulls, etc.), number of broods per change of litter, age and type of stock, floor area per bird, and the ration they have been fed. Table 1 shows the variability of these products. Feedlot manure is not included in this table as there have been few papers published on its use as a livestock feed. Ratio of roughage to concentrate in the ration would greatly influence the chemical composition of the manure. Several similarities exist in the reported chemical composition of the broiler litter. Crude protein is variable with a minimum of 20.9% and a maximum of 32.5%. It is estimated that 40 to 50% is true protein, 30 to 40% uric acid, and the remainder ammonia, urea, creatine, and other non-protein nitrogens. Uric acid has an advantage over urea in that it is 10,000 times less soluble. Belasco (1954) found that the purines—uric acid and allantoin (a product of partial breakdown of uric acid by enzymic pathway of degradation to ammonia)—are utilized by ruminal organisms as nitrogen sources. This occurs at a much slower rate than the breakdown of urea to ammonia, so the toxicity danger is almost nonexistant. The microorganisms are able to utilize all of the NH 3 , whereas the peak production of ammonia from urea degradation is greater than can be utilized by the organisms. The crude protein content of layer manure is higher than broiler litter, primarily because of the dilution of the broiler manure with the litter. Table 2 (Flegal, 1971) shows the amino acid composition of dried layer waste. Table 3 (Forsht et al., 1974) compares the ingredients in dried 20 may limit the activity of ruminal microorganisms. Lowman et al. (1970) found that the copper content of dried poultry excreta was almost double that of barley, but the digestibility was less than half as great, hence the available copper content of barley (24.5 ppm) was higher than that of dried poultry excreta (17.6 ppm). A local turkey producer feeds turkey manure mixed with either almond hulls or apple pulp to his ewes from June to December with no problems. After having fed beef for approximately 180 days, Oliphant (1974) analyzed their livers for copper and found controls with 333 ppm; those fed deep litter poultry manure, 282 ppm; and those fed battery poultry manure, 273 ppm. Fontenot et al. (1971) reports the average copper levels of ewe livers at death or slaughter for lots fed 0, 25, and 50% litter were 648, 1,993, and 2,671 ppm respectively. The average copper content of the litter was 195 ppm. Copper is more toxic to sheep than to cattle. He also states, "Feeding litter had no consistent effect on total red blood cells, total and differential white blood cell numbers, blood urea, blood ammonia, hemoglobin, or appearance of glucose, ketones, protein, occult blood or bilinibin in the urine. No treatment-related abnormalities other than those of copper toxicity were evident from gross observations, detailed necropies and studies of histological sections." layer waste and other primary poultry rations. It is interesting to note the high percentage of cystine in dry layer waste. Cystine and methionine are the two amino acids containing the sulphur atom used for wool growth. Experimental evidence of greater wool growth when feeding poultry waste is not documented but experience has indicated this. Whether it is because of the cystine and methionine or better over-all nutrition is not known. Crude protein content of feedlot manure was quite high-18.8% when the cattle were fed a ration of 32.5% alfalfa hay, 12.5% oat hay, 7.0% beet pulp, 39.0% rolled barley, 2.5% cottonseed meal, 6% molasses, 1% salt (Hull et al., 1974). Torell (1975) fed ewe lambs with a 60% feedlot manure, 40% almond hull pellet which contained 14.4% crude protein. Discounting the contribution of the almond hulls in the ration, the feedlot manure contributed 21.3% crude protein. ASH Soil contamination from chicken scratching and cleanup with scrapers etc. will account for some of the ash. Since most rations fed to chickens are highly digestible, the mineral component of the diet, mostly nondigestible, will be concentrated. The wide variation in total ash reflects variation in individual mineral constituents. Table 4 shows the ranges which have been recorded for dry poultry manure. Layers are fed high-calcium diets for egg shell formation so the manure would also be high in calcium. Phosphorus and magnesium are moderately high. Manganese and zinc are very high but would not cause problems to livestock. Copper is an element which needs monitoring, especially when sheep are fed in a feedlot for long periods. Fontenot et al. (1971) reported the occurrence of copper toxicity in one ewe after it was fed a diet containing 50% poultry litter for 137 days. When the experiment terminated (254 days), 64% of the ewes fed the ration containing 50% litter and 55% of those fed the 25% litter had died of copper toxicity.. The amount of copper used in the broiler's ration as a growth stimulant, then, would be a point of concern. A diet containing a high percentage of broiler litter or dried waste products tends to be unpalatable and of lowered digestibility (Bhattachary et al., 1964, 1965). The high levels of zinc and copper ENERGY Metabolizable energy from dried poultry waste was found to be 500 Kcal/lb. by Pryor and Connor (1964) and 300 Kcal/lb. by Nesheim (1972). These values should be compared with an average metabolizable energy figure for barley of 1,250 Kcal/lb. Broiler litter would have similarly low values because of the low digestibility of most of the litters such as rice hulls, wood shavings, peanut hulls, etc. As the feed passes through the digestive system, most of the energy is removed. APPARENT COEFFICIENT OF DIGESTION Table 5 shows results of some digestion trials which have included hen or broiler manure in the rations. The digestibility cannot be compared from one to another because of the variations among treatments. Each author considered the compari21 sanction the use of poultry litter as a feed or as a component of feed for animals. Poultry litter subject to the jurisdiction of the Federal Food, Drug, and Cosmetic Act and offered for use as animal feed may be considered as adulterated within the meaning of section 402(a) (1), (2) (C), and/or (3) of the act. At the time of this writing, the release of broiler litter as animal feed is imminent. sons that he made to be valid, so I have taken the liberty of condensing the ration ingredients to the pertinent differences between treatments. Dry matter and organic matter digestibility decrease as the percent of poultry manure increases in the ration. Bhattacharya and Fontenot (1965) found the protein digestibility to be reduced with increasing manure; however, Lowman and Knight (1970) and Harmon et al. (1975) showed an increase. Crude fiber digestibility changes slightly (Bhattacharya and Fontenot, 1966; Harmon and Fontenot, 1975) or decreases (Bhattacharya and Fontenot, 1965) with increased amounts of manure, as does energy digestion (Bhattacharya and Fontenot, 1965; Lowman and Knight, 1970). DISEASES With the FDA regulation in mind, Caswell et al. (1975) tried several methods for reducing or eliminating the bacteria in the broiler litter. Dry heat, auto-claving, dry heat and paraformaldehyde, and ethylene oxide were used. All samples tested negative for Salmonella, the bacteria of most concern especially for young lambs, as well as Shigella and Proteus. However, he did find more than 30,000 colonies of total bacteria per gram of untreated litter. Coliforms were completely eliminated by all processing methods except fumigation with ethylene oxide for thirty minutes. Dry heat plus paraformaldehyde was very effective, especially at 4 g. per 100 g. litter. Fumigation with ethylene oxide for 2 hours was also very effective. Jacobs and Leibholz (1974) state, "Pomeroy (personal communication, 1972) has studied litter ensiled in a `Harvestore' on a commercial feedlot property in Minnesota. He detected Salmonella in the litter going into storage, but none in the `poultry silage' (ensiled about 30 days)." Harmon et al. (1975) conclude, "The trend for lower coliform numbers in litter silage than controls suggests that ensiling may be an economical means of eliminating potential hazards from the possible presence of pathogens in litter." The magnitude of the problem associated with feeding litter to livestock, on the basis of potentially infectious microorganisms found in litter is well documented by bacteriologic studies (Halbrook et al., 1951; Whitehead and Corson, 1962; Schefforle, 1965; Tucker, 1967; Alexander et al., 1968; Messer et al., 1971). However, Alexander et al. (1968) also state that the diseases of poultry, apart from Salmonellosis, are not thought to be of major importance in the feeding of ruminants. Salmonella REGULATIONS The Food and Drug Administration has issued a policy statement regarding the use of litter as animal feed (Kirk, 1967). (a) Poultry rations used today generally contain drugs including antibiotic drugs used individually or in combination. The levels of drug use vary from very small quantities of antibiotic drugs for growth promotion to relatively large quantities of drugs for treatment of diseases. Consequently, poultry litter can be expected to contain drugs and antibiotic drugs or their metabolites. It is not practical to determine, or feasible to estimate, the nature and levels of the drugs and their metabolites in litter. Therefore, it is not possible to conclude that poultry litter is safe as a feed or as a component of feed for animals, nor is it possible to conclude that there will be no drug residues in the tissues and by-products of animals fed poultry litter. (b) Disease organisms may be transmitted from poultry to other animals through the use of poultry litter as animal feed. There are several diseases affecting poultry that can also affect cattle, hogs, and sheep as well as man. Thus such transmission of disease organisms from poultry to other animals and possibly to man constitutes a hazard to animals and to the public health. (c) Therefore, the Food and Drug Administration has not sanctioned and does not 22 for the litter groups. When fattening steers (Table 11), there was little difference in gain between the steers fed the peanut hull poultry litter ration and those fed the control ration. Feed efficiency was highest for the steers fed the poultry litter ration. Noland et al. (1955) also fattened steers and found that steers fed ground chicken litter as a source of nitrogen gained more slowly than those fed cottonseed meal. This difference was significant during the first 56 days when the nitrogen intake was equal but there was no significant difference when the amount of litter ration was increased (15%) to nearly equal the energy intake of the two groups. Oliphant (1974) conducted three years of experiments with British Friesian steers from 150 kg. live weight to slaughter at 400 kg. The treatment rations were made isonitrogenous (14.5% crude protein). When the crude protein of the poultry manure was low (24%), poor performances resulted. For manures of 30% crude protein, performance was not markedly different from that shown with control diets (Table 13). El-Sabban et al. (1970) used poultry waste subjected to three different heat treatments to produce autoclaved, cooked and dried products which he fed to steers for a 139-day feeding trial. All of the supplemental nitrogen was provided by soybean meal, urea or the poultry waste. Rate of gain and feed efficiency were not significantly different amount steers fed rations in which supplemental nitrogen was provided by soybean meal, autoclaved and dried poultry waste (Table 14). Carcass characteristics and meat acceptability were not significantly different among the steers fed the different rations. In a sheep fattening experiment (Table 15) Thomas et al. (1972) found sheep fed rations containing 25 and 50% dehydrated feces gained .16 and .15 kg./day which was significantly less than those fed a control corn-corncob-soybean meal ration (.21 kg./day). "Results indicate the feasibility of using relatively large amounts of dehydrated caged layer feces as a nitrogen and energy source for cows and lambs." Galmez et al. (1970) fed lambs varying amounts of a rice hull, broiler litter ration (27.4% crude protein) with the remainder of the ration containing one-half beet pulp and one-half ground oats plus 2% salt (Table 16). These rations were compared with baled alfalfa hay. All lambs on the organisms may be transferred to ruminants, but these should not cause difficulty in adult animals unless the management is poor (Gibson, 1965). Of the many experiments with unprocessed broiler litter, none have reported any disease problems. Torell (unpublished data) has fed unprocessed litter to from 600 to 1,000 ewes during flushing and breeding for four years and to 1,200 ewes for two to four weeks just prior to lambing without any disease problem caused by the litter. During this period, one ewe did not eliminate the sand contained in the litter from her rumen and had to be sacrificed. Broiler litter-apple pulp has also been fed to replacement ewe lambs with no disease problem (Torell, 1975). PRODUCTION We have seen that broiler litter and layer manures are high in nitrogen, low in energy, are fairly well digested, and can be made safe to feed. Tables 6 through 21 present the results of some studies measuring the feeding potential of this feed when used as a nitrogen (protein) source. These data are based primarily on trials with beef cattle but because of the similarity of the rumen the results can be applied to sheep as well. Drake et al. (1965) fattened beef on broiler litter derived from four different litters and compared results with the control ration (Table 6) of soybean meal and chopped hay (protein and fiber equalized to litter mixtures). The average daily gain for the broiler litter groups were less than the control groups, but significance was not reported. Ray and Child conducted a number of experiments feeding broiler litter rations for wintering beef cows and calves (1965), wintering steers (1964), and fattening steers (1964). The results of their experiments are shown in Tables 7, 8, and 9. For wintering cows, the litter ration prevented as much weight loss to the cows and increased the calf gains at less expense. Wintered calves and yearlings fed litter rations gained more with about the same feed to gain ratio as the prairie hay groups. The same type of results were obtained in finishing steers. Fontenot et al. (1964), wintering steers and heifers (Table 10), found that the feeding of a wood-base broiler litter had no marked effect on the daily gain of the cattle—I.25 lbs. per day for the cottonseed meal groups and 1.22 lbs. per day 23 the cottonseed meal supplement, but intake of supplement was much greater for those given the manure-barley pellet." litter rations gained more than the alfalfa group as well as being more efficient. Fairly high levels of broiler litter can be fed (68%) with good gains (170 g.). They also fed ewes, during the last third of gestation and approximately 90 days of lactation, a ration containing 63% broiler litter, 35% beet pulp, 2% salt and one composed of 87% alfalfa, 13% beet pulp. Both rations were calculated to contain 52% TDN. Table 17 shows the results of these trials. There were no significant differences in lamb birth weights, 90-day weights, or ewe consumption between the rations although the litter group's lambs were heavier at birth and at 90 days. Toren (1975) fed three groups of ewe lambs rations containing either 50% broiler litter plus 50% apple pulp, 60% feedlot manure plus 40% almond hulls as a pellet or 60% alfalfa plus 40% almond hulls as a pellet for 97 days prior to joining with rams and for another 49 days of breeding. The regulated feed consumption was approximately the same for all three groups to give similar nitrogen intake. The 97-day gains were 6.2 kg., 2.2 kg., and 14.4 kg. respectively. The percent breeding as lambs and lambing as yearlings was highest for the alfalfa-almond hull group (90%), followed by the broiler litter and apple pulp group (68%), and lowest for the feedlot manure almond hull group (50%). Noland et al. (1955) fed ewes during gestation and lactation on rations of oat hay and supplemented with either soybean meal, ground chicken litter, or ammoniated molasses. Table 18 summarized the results. The group fed ground chicken litter performed as well as those fed soybean meal. Ensiling broiler litter with either corn forage (Harmon et al., 1975) or with grain (Jacobs and Leibholz, 1974) looks very promising. Table 19 shows the results of ensiling grain with broiler litter. Dry matter intake and nitrogen balance were significantly improved by ensiling the grain with the broiler litter. COMMENT In view of the FDA regulation, one might question the practicality to the sheep industry of proceeding with this line of investigation. But the Federal Food and Drug Administration is concerned only with human food products transported across state lines. Since breeding ewes are the class of sheep most likely to be fed these waste products, rather than lambs, and since too often ewes are culled only "when they have been dead three days," it is unlikely that FDA laws would be violated. Harmon et al. (1975) investigated the nutritive value and palatability of ensiling broiler litter and corn forage cut at two stages of maturity. Dry matter digestibility averaged 64.5% and was similar for all silages. Voluntary dry matter intake was higher, approximately 70% for silages containing broiler litter than for the control (corn silage only) group. Even though poultry waste does not change the flavor of the meat (Drake et al., 1965; El-Sabban et al., 1970; and Fontenot et al., 1964) lamb produced on this type of feed would probably not be so popular with the public. Ironically, the public is blissfully unaware that millions of hogs have been fattened behind steers in the feedlot. If this fact were publicized, many pork consumers would undoubtedly alter their meat preferences somewhat. Lamb producers must protect their image but at the same time operate efficiently, to stay in business and feeding poultry manure to breeding ewes can be considered an efficient management practice. FEEDLOT MANURE Hull et al. (1974) supplemented beef cows grazing dry native annual range for a period of 84 days with pelleted cottonseed meal (0.90 kg./head daily) or a pelleted mixture of 75% feedlot manure and 25% barley (ad lib.). A third group received no supplementation. "Cows given the manure-barley pellet had a higher body weight than cows given 24 Chance, C. M. 1965. Non-protein nitrogen and poultry litter in ruminant diets. Proc. Md. Nutri. Conf., p. 8. REFERENCES Alexander, D. C., J. A. J. Carriere, and K. A. McKay. 1968. 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Evaluacion nutritiva y ensayo de consumo de cama de broiler. Proc. XX Jornadas Agronomicas, Chillan. Hart, S. S. 1960. The management of livestock manure. Transactions, Amer. Soc. of Agri. Eng. 3:78-80. Morgan, D. E. 1970. Feeding of dry poultry manure. Proc. N.A.A.S. Conf: of Advisory Nutr. Chems. (Edinburgh). Howes, J. R. 1966. On-site composting of poultry manure. Management of farm animal wastes. Proc. National Symposium, Am. Soc. Agric. Eng. p. 68. Morris, W. H. M. 1966. Economics of liquid manure disposal from confined livestock. Natl. Symposium on Animal Waste Mangt. Proc., Amer. Soc. Agr. Eng. Pub. No. SP-0366. St. Joseph, Mich., 126-131. Howes, J. R. 1968. Management and utilization of poultry wastes. Feedstuffs 40(50):22-23. Hull, J. L., C. A. Raguse, J. G. Morris, and R. Delmas. 1974. Feedlot animal waste compared with cottonseed meal as a supplement for pregnant range cows. J. Range Mgt. 27(3): 192. Morris, W.H.M. 1971. Economics of waste disposal from confined livestock. Internatl. Symposium on Livestock Wastes Proc., Columbus, Ohio. 195-196. Muhrer, M. E., and E. J. Carroll. 1964. Urea utilizing microorganisms in the rumen. J. Animal Sci. 23:885. Jacobs, G. J. L., and Jane Leibholz. 1974. The digestibility of ensiled broiler house litter/barley rations for sheep. Proc. Aust. Soc. Prod. 10:400. Nesheim, M. C. 1972. Evaluation of dehydrated poultry manure as a potential poultry feed ingredient. Cornell Agr. Waste Mangt. Conf., Syracuse, N. Y. Kirk, J. K. 1967. Use of poultry litter as animal feed. Fed. Register 32(171):12714. Noland, P. R., B. F. Ford, and M. L. Ray. 1955. The use of ground chicken litter as a source of nitrogen for gestatinglactating ewes and fattening steers. J. Animal Sci. 14:860. Leibholz, Jane. 1969. Poultry manure and meat meal as a source of dietary nitrogen for sheep. Aust. J. Exp. Agr. and An. Husb. 9:589. Linn, A. 1966. Whipping the manure problem. Farm Quarterly, Winter, 56-59. Oliphant, J. M. 1970. Feeding poultry waste for intensive beef. Prog. Rep. Exp. Husb. Fms Exp. Hort. Stas. N.A.A.S. 11:94. Linton, R. E. 1966. The economics of poultry manure disposal. Cornell Agr. Exp. Sta. Bul. 1195, Cornell Univ., Ithaca, N .Y. Oliphant, J. M. 1974. Feeding dried poultry waste for intensive beef production. An. Prod. 18:211-217. Livshutz, A. 1964. Aerobic digestion (composting) of poultry manure. World's Poultry Sci. Jour. 20:212-215. Palafox, A. L., and M. M. Rosenberg. 1951. Dried cow manure as a supplement in a layer and breeder ration. Poultry Sci. 30:136-142. Loehr, R. C. 1968. Pollutional implications of animal wastes-a foreward oriented view. Fed. Water Pollut. Control Admin., Environ. Protection Agency, Ada, Okla., July. Parker, M. B. 1966. Chicken manure on orchard grass-ladino clover. Ga. Agr. Exp. Sta. Bul. N.S. 159, Univ. of Ga., Athens, May. Loehr, R. C. 1972. Animal waste management-problems and guidelines for solutions. J. Environmental Quality 1(1):71-78. Perkins, H. F., and M. L. Walker. 1964. Chicken manure-its production, composition, and use as a fertilizer. Ga. Agr. Exp. Sta. Bul. N.S. 123, Univ. of Ga., Athens, Dec. 26 Perkins, H. F., and M. B. Parker. 1971. Chemical composition of broiler and hen manures. Ga. Agr. Exp. Sta. Res. Bul. 90, Univ. of Ga., Athens, Mar. Taylor, J. C. 1971. Regulatory aspects of recycled livestock. and poultry wastes. Proc. Internatl. Sym. on Livestock Wastes. A.S.A.E. Pub. PROC-271-291. Pryor, W., and J. K. Connor. 1964. A note on the utilization by chickens of energy from faeces. Poultry Sci. 43:883-834. Taylor, J. C. 1972. Residues and heavy metal contamination problems caused by refeeding animal wastes. Proc. Maryland Nutrition Conference. Washington, D. C., 10 pp. Quisenberry, J. H., and J. W. Bradley. 1968. Nutrient recycling. Second Natl. Poultry Litter and Waste Mangt. Seminar, College Station, Tex., 96-106. Thomas, J. W., and H. C. Zindel. 1971. Feeding dehydrated poultry waste to dairy cows. Mich. Agri. Exp. Sta. Farm Sci. Res. Rpt. 152, Mich. State Univ., East Lansing, Nov. 8-11. Ray, M. L., and R. D. Child. 1964. Wintering steers on broiler house litter. Ark. Farm Research 13:5. Thomas, J. W., Yu Yu, P. Tinnimitt, and H. C. Zindel. 1972. Dehydrated poultry waste as a feed for milking cows and growing sheep. J. Dairy Science 55 (9):1261. Ray, M. L., and R. D. Child. 1964. Broiler house litter used as roughage in finishing rations for steers. Ark. Farm Research 13:3. Thornber, Cyril. 1969. British cite value of poultry manure in ruminant feeds. Feedstuffs 41:321. Ray, M. L., and R. D. Child. 1965. Chicken litter as a supplement in wintering beef cows and calves on pasture. Arkansas Farm Research XIV:4. Toren, D. T. 1975. Broiler litter and feedlot manure for ewe lambs. J. Animal Sci., Vol. 41. Tucker, J. F. 1967. Survival of salmonellae in built-up litter for housing of rearing and laying fowls. Brit. Vet. J. 123-92. Ray, M. L. The digestion and utilization of chicken litter as a nitrogen source in beef calf wintering rations. Mimeograph. Univ. Arkansas. Walsh, L. M., and R. F. Hensler. 1971. Manage manure for its value. Wisc. Coop. Ext. Serv. Pamphlet, Univ. Wisc., Madison, June. Rusnak, J. J., T. A. Long, and T. B. King. 1966. Value of hydrolyzed poultry waste as a protein supplement for beef cattle. Dept. Anim. Sci. Rpt. 4-66, Pa. State Univ., University Park (mimeograph). Wehunt, K. E., H. L. Fuller, and H. M. Edwards, Jr. 1960. The nutritional value of hydrolyzed poultry manure for broiler chickens. Poultry Sci. 39:1057-1063. Saylor, W. W., and T. A. Long. 1972. Nutritive value of ensiled animal wastes, J. Animal Sci. 35:288. Whipple, R. 1971. Twenty-eight percent "safe" protein from broiler litter, Broiler Industry, 34(1):38-43. Schefferle, H. E. 1965. The microbiology of built-up poultry litter. J. Appl. Bact. 28:403. Whitehead, G. L., and 0. D. Corson. 1962. Tuberculin sensitivity from oral administration of killed Mycobacteria tuberculosis. Cornell Vet. 52:36. Scholz, H. G. 1971. Systems for the dehydration of livestock wastes-a technical and economic review. Internatl. Symposium on Livestock Waste Proc., Columbus, Ohio, Apr. 27-29. Yoshida, M., and H. Hoshu. 1968. The nutritive value of poultry manure. Jap. Poult. Sci. 5:37-39. Zindel, H. C., and C. J. Flegal. 1971. Economics of dried poultry waste (DPW) as a feed ingredient or a fertilizer. Mich. Agr. Exp. Sta. Farm Sci. Res. Rpt. 117, Mich. State Univ., East Lansing, Nov. 4-7. Smith, L. W. 1971. Feeding value of animal wastes, in animal waste reuse-nutritive value and potential problems from feed additives: a review. U.S. Dept. Agr., ARS 44-224, Feb. Zorita, Tomillo E., Guedas J. Rodriguez, and Martin J. Balboa. 1966. Rev. Nutrition Animal, Madrid, 4, 11, 111, 128. Snell, J. R. 1971. A new economic approach to the treatment and utilization of cattle feedlot wastes. Report prepared for Cobey Environ. Controls Co., Inc., Lansing, Mich. Sobel, A. T., and D. C. Ludington. 1966. Destruction of chicken manure by incineration. Natl. Symposium on Animal Waste Mangt. Proc., Amer. Soc. of Agr. Engin. Pub. No. SP-0366, St. Joseph, Mich., 95-98. Southwell, B. L., 0. M. Hale, and W. C. McCormick. 1958. Poultry house litter as a protein supplement in steer fattening rations. Georgia Agr. Exp. Sta. Mimeo Service M.S. 55. Surbrook, T. C., J. S. Boyd, and H. C. Zindel. 1971. Drying animal waste. Mich. Ag. Exp. Sta. Farm Sci. Res. Rpt. 117, Mich. State Univ., East Lansing, July, 16-20. Taiganides, E. P., and T. E. Hazen. 1966. Properties of farm animal excreta. Transactions, Amer. Soc. Agr. Eng. 9:374-376. 27 Table 1. Some chemical compositions of broiler litter and caged layer manure (dry poultry waste). • • Investigator • • E.E. C.P. C.F. N.F.E. ASH 15.2 14.6 9.7 12.7 20.8 16.0 30.7 Broiler Litter Fontenot et al. (1964): Galmez et al. (1970): Manterola (1969): English (unpublished) Forsht et a/. (1974) Evans et al. (unpublished) Means of 20 samples Torell (1975) Peanut hulls Shavings Shavings Rice hulls Shavings 32.5 32.4 21.7 27.4 21.6 24.4 29.9 3.3 2.5 16.3 16.7 31.1 30.7 31.8 2.9 1.3 3.1 24.8 26.0 18.2 29.9 32.3 17.9 25.8 20.9 1.4 24.5 24.8 23.4 25.2 Caged layer manure English (unpublished): Maximum Minimum Morgan (unpublished) Thomber (1969) Flegal, et al. (1971) Leibholz (1969) 39.1 24.7 32.7 26.6 24.3 28.0 1.1 0.5 1.8 14.3 12.4 11.9 26.1 2.1 0.9 13.7 7.6 39.6 21.8 Table 2. Amino acid composition of dried layer waste. Amino acid True protein Crude protein Grams/100 grams Alanine Arginine Aspartic acid Cystine Glutamic acid Glycine Histidine Isoleucine Leucine Lysine Methionine Phenylalanine Proline Serine Threonine Tyrosine Valine 4.38 1.93 4.38 4.51 6.35 3.40 .82 2.05 3.32 2.01 .37 1.84 2.21 2.13 2.05 1.07 2.58 Source: Flegal et al., 1971. 28 9.73 4.27 9.73 10.00 14.09 7.54 1.82 4.54 7.36 4.45 .82 4.09 4.91 4.73 4.54 2.36 5.73 39.4 29.5 27.5 15.2 26.9 33.8 Table 3. Comparison of ingredients in dried layer waste and other primary poultry rations. DLW Ingredient Alfalfa Corn Soybean mean (44%) • Ground oats Dried whey Wheat m iddlings Milo Percent Amino Acid: Arginine 0.48 1.13 .85 Cystine Glycine Lysine Methionine .50 .09 7.5 2.5 20.0 Tryptophan Calcium Phosphorus Fiber 0.82 .33 .88 .95 .30 .29 1.4 .3 24.2 0.43 .15 .32 .27 .16 .08 .02 .30 2.00 3.21 .63 0.74 .20 1.94 2.93 .59 .43 .47 .16 .61 .3 .6 5.9 .15 .1 .4 11.0 0.37 .33 .70 0.98 .19 .43 .90 .67 .18 .30 .29 .16 .19 .16 .18 .9 .8 .2 .21 .1 .8 6.7 .11 .02 .3 2.7 885 1,480 0.31 1,000 calories per pound Metabolizable 636 energy - 555 1,500 1,020 1,165 868 = not analyzed Source: Forsht et al., 1974 Table 4. Variations in ash content of dried poultry manure. PPM in dry matter Percent Calcium Phosphorus Magnesium Sodium Copper Manganese 2.4 - 11.5 1.4 - 2.8 0.4 - 0.6 0.3 - 1.0 Zinc Cobalt 29 60 40 300 - 500 350 - 1200 0.4 - 0.9 Table 5. Apparent coefficient of digestibility. Investigator and feed tested Brugman (1964) Layer litter Anthony (1963) Control Layer litter 1 Brood B.L Several brood B.L. Bhattacharya (1966) Control, alfalfa & corn 25% of control replaced by peanut hull litter 50% of control replaced by peanut hull litter 25% replaced by wood shaving litter 50% replaced by wood shaving litter Bhattacharya (1965) Control, SBOM 25% of control replaced by peanut hull litter 50% of control replaced by peanut hull litter 100% of control replaced by peanut hull litter El-Sabban (1970) Control, Soybean meal Autoclaved poultry waste Cooked poultry waste Lowman (1970) Control, barley 25% of control replaced by dried layer litter 50% of control replaced by dried layer litter 75% of control replaced by dried layer litter 100% of control replaced by dried layer litter Harmon (1975) Silage: Corn maturity Treatment 30% DM 30% DM 30% DM 30% DM 40% DM 40% DM 40% DM 40% DM Harmon (1974) 10.35% Soybean meal 14.84% Autoclaved B.L 13.25% Heat dried B.L 12.20% Acid & heat dried B.L Control Urea B.L. 15% B.L. 30% Control Urea B.L. 15% B.L. 30% : D.M. : O.M. 58.8 62.0 53.6 57.5 76.7 73.1 70.1 74.1 69.1 : C.P. 77.8 - - 78.4 79.1 76.6 71.2 : C.F. 59.2 75.0 74.5 74.0 74.6 72.0 59.6 62.6 63.2 62.2 62.9 76.4 72.7 69.2 72.7 69.4 67.1 64.8 57.7 73.9 76.8 74.1 71.0 60.4 58.4 40.5 41.3 - 74.3 65.5 69.4 77.9 73.6 68.7 62.9 56.6 80.7 77.3 74.0 70.3 66.5 68.4 69.5 76.9 76.5 77.2 - 64.3 66.7 65.0 63.4 65.0 64.9 64.1 62.7 - 57.1 71.5 59.9 67.0 48.0 61.9 54.9 64.0 53.5 58.6 54.5 54.8 44.0 44.0 46.1 52.7 63.3 59.9 58.6 56.3 65.5 57.9 62.4 54.3 72.9 66.8 68.8 65.1 30 - Energy 91.0 75.3 72.1 76.2 - : - - 74.7 73.5 76.3 80.0 74.8 72.1 66.1 60.3 Table 6. Beef finishing trials with various litters. Average dail y gain Broiler litter base 25% : Broiler litter : 40% Broiler litter .91 .96 1.02 .92 1.07 1.00 1.01 1.06 0.94 1.21 25% Peanut hull litter 25% Corn cob litter 25% Chopped hay litter 25% Soybean hull litter Control Source: Drake et al., 1965. Table 7. Chicken litter as a supplement in wintering beef cows and calves on pasture. Ration Feed cost per cow and calf Calf weight change Cow weight change Dollars Pounds 21.98 23.81 22.72 10.65 +120 + 82 +137 +124 -155 -206 -116 -118 Litter* free choice + all hay All hay Litter* (2 hrs.) + all hay Free choice litter* All on pasture. *Litter = 4 parts litter to 1 part corn. Source: Ray and Child, 1965. Table 8. Wintering steers on broiler house litter. Yearlings Calves Ration ADG Feed/gain ADG Feed/gain Pounds Broiler litter (cottonseed hull) + 2 lb. prairie hay Prairie hay 6.81 6.65 2.61 2.49 All received 75 to 85% of their ration as 9 parts ground corn and 1 part cottonseed meal. Source: Ray and Child, 1964. 31 3.25 2.05 7.39 8.84 Table 9. Results of finishing steers by feeding broiler litter and rice hull rations. Ration Gain Feed/gain Pounds 25% broiler litter 25% broiler litter + 4.5% fat 25% - 12% protein rice hulls (ammoniated) 25% - 12% protein rice hulls + 4.5% fat 288 305 309 298 8.17 8.20 7.97 7.43 Source: Ray and Child, 1971. Table 10. Performance of wintering cattle fed broiler litter on cottonseed meal - 56 days. Lot Cattle Supplement 1 2 3 4 5 6 Steers Heifers CSM Litter CSM Litter CSM Litter 23.4 3.0 1.0 23.1 3.0 23.5 3.0 1.0 23.3 3.0 23.4 2.9 1.0 23.2 2.9 Feed intake Lbs/head/day Corn silage Hay Cottonseed meal Broiler litter 1.4 Daily gain, lbs. 1.18 1.4 1.07 1.09 1.21 1.4 1.48 1.39 Source: Fontenot, Drake et al., 1964. Table 11. Feedlot performance of steers fed poultry litter rations (123 days). • • • • Peanut hulls Wood shavings Daily feed:* Mixture Long hay Feed/gain Mixture Long hay Total Daily gain • • • • Control Pounds 26.2 2.2 26.3 2.2 29.8 2.2 9.34 0.79 10.13 9.91 0.84 10.75 10.40 0.78 11.18 2.81 2.65 2.87 *Salt and a mineral mixture of 3 parts defluorinated pho stone and 1 part salt were provided, in addition. Source: Fontenot, Drake et al., 1964. 32 Table 12. Summary of finishing steer trial. 2.14 Av. daily gain (lb.) Period 3* 42 days • Cotton- Chicken seed litter meal Period 2* 56 days CottonChicken : seed litter : meal Period 1* 56 days •. Cotton- • Chicken : seed : : litter : meal 1.86 1.81 2.08 1.32 Av. daily feed intake per steer (lb.): 17.3 5.8 Concentrate mixture Prairie hay Feed per cwt. gain (lb.): Concentrate mixture Prairie hay 808 270 17.3 19.5 5.8 6.5 978 326 1049 350 19.5 6.5 1.92 23.8 27.4 7.0 9.1 1148 382 1472 475 1472 491 *Steers fed cottonseed meal ration in period 1 were fed chicken litter ration in 2 and 3 and vice versa. Source: Noland et al., 1955. Table 13. Mean performance of cattle. Year Control Deep litter poultry manure Mixture Battery poultry manure Daily live-weight gain (kg) 1969 1970 1971 1.13 1.29 1.18 0.93 1.23 1.04 1.09 1.23 1.26 1.14 1.19 Daily intake of dry matter (kg) 1969 1070 1971 5.6 6.1 5.6 5.2 6.0 5.3 5.6 6.2 5.4 6.1 5.4 Dry-matter conversion ratios (kg food dry matter/kg live-weight gain) 1969 1970 1971 4.95 4.71 4.72 5.67 5.14 4.90 5.12 5.00 4.74 Source: Oliphant, 1974. 33 4.86 4.90 Table 14. Average feedlot performance of steers fed rations containing processed poultry waste. Ration Nitrogen source No. of steers Daily gain, kg Daily feed consumption, kg Kg feed/kg gain 1-Soybean meal 6 2-Autoclaved poultry waste 7 3-Dried poultry waste 6 4Urea 6 1.22 12.53 10.27 1.22 12.12 10.02 1.15 12.45 10.83 1.43 11.66 8.15 Source: El-Sabban et al, 1970. Table 15. Performance of sheep fed dehydrated caged layer feces rations. Broiler litter Item Weight gain (kg/day) Intake grain mix (kg/day) Feed/Gain 0% 25% 50% .21 1.39 7.10 .16 1.41 10.00 .15 1.65 12.50 Source: Thomas et al.,1972. Table 16. Lamb fattening results. Ration Lot ADG(gm) Feed/gain Broiler litter Broiler litter Broiler litter Broiler litter Baled alfalfa 68% 58% 48% 38% 170 174 186 208 84 5.4 5.3 4.5 4.0 25.0 Rice hull base with 27.4% crude protein. Ration also contained 1/2 beet pulp, 1/2 ground oats, and 2% salt. Source: Galmez et al., 1970. 34 Table 17. Pregnancy and lactation results. Lamb data 63% Broiler litter 87% Alfalfa 4.78 19.70 171.00 4.39 17.60 146.00 2.4 2.5 Birth weight (kg) 90-day weight (kg) ADG (gm) Ewe consumption pre-partum (kg) Beet pulp + 2% salt composed remainder of ration. Source: Galmez et al., 1970. Table 18. Summary of gestating-lactating ewe trail. Item Initial weight per ewe (lb.) Final weight per ewe (lb.) No. lambs dropped No. lambs raised Birth weight per lamb (lb.) Birth weight, twins (lb.) Birth weight, singles (lb.) Av. daily gain birth to termination of trial: All lambs (lb.) Singles (lb.) Twins (lb.) Av. daily conc. intake of ewes (lb.) Oat hay fed per ewe daily (lb.) Oat hay consumed per ewe daily (lb.) Total conc. intake per lamb from creep (lb.) Lot I Soybean oil meal lot Lot 2 Chicken litter lot Lot 3 Ammoniated molasses lot 135 111 12 10 10.2 7.4 10.7 134 112 12 10 9.4 6.2 10.1 132 106 14 13 9.4 8.0 10.4 0.44 0.44 0.42 0.42 0.42 0.42 0.34 0.39 0.28 1.13 3.68 3.10 1.13 3.68 3.26 3.68 3.18 85.6 Source: Noland et at, 1955. 35 86.8 1.13 65.8 Table 19. Dry matter intake, dry matter digestibility, nitrogen digestibility. Diet Poultry litter + barley Poultry litter ensiled 3 wks. + barley Poultry litter + barley ensiled 3 wks. Poultry litter ensiled 6 wks. + barley Poultry litter + barley ensiled 6 wks. Dry matter intake Dry matter digestibility Nitrogen digestibility Nitrogen balance Gm/day % % Gm/day 932 888 1177 930 1104 65.8 66.4 64.5 66.8 66.3 69.8 72.5 72.9 72.5 73.7 5.9 6.7 11.4 6.9 10.5 Source: Jacobs and Leibholz, 1974. Table 20. Daily silage dry matter intake by sheep, palatability study. Silage • • Maturity 30% DM 30% DM 30% DM 30% DM 40% DM 40% DM 40% DM 40% DM Intake Treatment Grams DM Control Urea Litter-15% Litter-30% Control Urea Litter-15% Litter-30% 881 844 1317 1581 816 1006 1573 1348 Source: Harmon et al., 1975. Table 21. Mean body weight changes of pregnant range cows grazing dry annual range and given supplements of cottonseed meal (CSM) and manure-barley pellets. Item None Number of cows Length of supplemental period (days) Gain or loss during supplementation (kg) Supplement consumed (kg/hd/day) 8 84 -36.3 0 Source: Hull et al., 1974. 36 Supplement CSM 8 84 +14.5 0.90 Manure-barley 8 84 +34.9 7.75 contained 16.4% crude protein and 26.1% crude fiber on a dry matter basis. At each feeding level one group was fed daily; another was given its week's ration divided into two feedings (Monday and Thursday), while the third group received its entire week's ration at one feeding (Monday). The ewes were classified as thin, medium, or fat at the beginning of flushing and weighed after an overnight shrink at the beginning of flushing and when they left the feedlot. The ewes were sheared on the 11th day of breeding. The time at which the cubes were all consumed was recorded. At breeding two mature whiteface rams were placed with each group of ewes and given the same ration as the group of ewes to be bred. After 32 days in the feedlot, all ewes were returned to the same range pasture and the rams remained with the ewes for another 34 days. PART II. OBSERVATIONS ON FREQUENCY OF FEEDING Over the years we have developed the custom of eating three times a day and, if a meal is not served at the customary time, we get hunger pains. Likewise, we have assumed that our livestock must be fed at least once a day. In many parts of the world experiments have been conducted to examine the possibility of feeding ruminants at less frequent intervals than daily. Australian researchers have carried on many studies in drought feeding. They have found that, with daily feeding, the strong ewe gets stronger and the weak ewe gets weaker because the strong ewe eats more than her share of the daily ration while the weak ewe is timid and takes less than her share. Whereas, when feeding the whole week's ration in one day, they found that the strong ewe got her fill and left the feeding area so that the weak ewe had her turn at the feed and ate as much as she wanted. They both felt the pangs of hunger before the next feeding. The ewes were brought to a barn for lambing and the collection of data. Lambing data reported are for the first 26 days of lambing. RESULTS and DISCUSSION The two levels of feeding had a significant effect (P<0.01) on body weight change during the 32 days in the feedlot. Table 1 shows these changes in body weight corrected for fleece weight. All groups of ewes given the 4.0 pounds per day of cubes gained weight (mean 6.2 pounds) while all groups given 2.2 pounds of cubesper day lost body weight (mean -6.3 pounds). Frequency of feeding had less effect on body weight than level of feeding. At Hopland we have supplementally fed replacement ewe lambs daily and once and twice per week and have found that the lambs do as well or better when fed less frequently than when fed daily. This was also true when ewes were supplemented on the range during the latter part of gestation. The following experiment was conducted to determine the effect of frequency of feeding during flushing and breeding upon lambing percentage. The difference in body weight change between groups of ewes given the equivalent of 2.2 and 4.0 pounds of cubes daily decreased as the time interval between feedings increased. Two hundred eighty-eight Targhee type ewes were taken from range on August 13, 1973, and assigned by age and condition into 6 groups, each of which were fed in pens for 14 days of flushing, then 18 days of breeding. A 3x2 factorial allocation of treatments was used. Three groups were fed the equivalent of 2.2 pounds alfalfa cubes per day and three the equivalent of 4.0 pounds alfalfa cubes per day. The alfalfa cubes Changes in body weight during the period in the feedlot were not related to the initial condition of the ewe. Initial condition and body weight at the commencement of feeding were related. The mean body weights of the thin, medium and fat 37 ewes were 123, 130, and 141 pounds respectively. they were without any feed for one day between feedings for both the 2.2 and 4.0 pound groups at the beginning of the experiment, but this time was increased to 1.5 to 2 days by the end of the experiment. When ewes at Hopland Field Station are bred on the range, approximately 1.15 lambs are born per ewe lambing. Using lambing percent as an index of nutrition, it would appear that all groups in this experiment were receiving better nutrition than was available on the range, even though the groups given 2.2 pounds of cubes per day lost weight (table 1). There were 6.2% dry ewes in the groups given 2.2 pounds and 1.6% in those given 4.0 pounds per day. Considering all of the groups, 40% of the dries were in the daily, 10% in the twice-per-week, and 50% in the once-per-week groups. As the differences in lambing percentages between groups fed at the three frequencies of feeding and two planes of nutrition were not statistically significant, it would be advantageous from a management aspect to feed once or twice weekly. A level of 2.2 pounds alfalfa cubes per day appeared adequate for medium and fat ewes with possibly 4.0 pounds being superior to 2.2 pounds for thin ewes. Feeding a week's ration either once or twice per week resulted in similar lambing percentage as daily feeding but from practical management intermittent feeding would require less labor. The number of days sheep were without food between feedings was greater in the group given 2.2 pounds (3.5-4 days) than the group given 4.0 pounds (3-3.5 days). When the ewes were fed twice per week, 38 0 A - co • • 111• 00 0 0\ c1 c- a) `..0 • EN- K \ 1 1 • • E"-0 v- c-• K•\ -1-• K \ -1N1 ai 0 O'N 00 CO • z • • Cr' 0 •• o • H CT • CYN \ . CO • Kl CO • 1 A r24 0 0 •=C c- • N • K •\ • N LI\ N1 z •• •• ;11 Kl• K\• •r-I rd 0" 0 z a) • N• L(\• N-• r•rN r• U\• t•C te\ 1 •1 1 • 0 • LC\• ifl• 0• N- N 0 N 1 A a) 0 Z AH 0 A W • 44 CH • 0• .,-1 N -t A •• •• • N• vv- co ...7.; co N • 1 N• 00• -1- -.1- • 0 • N •• 1.4 - 0■ 1 . 4-4 cH •H N. 0• N -t A K1 . v■.0 LI\• • •• • 4-1 4-1 • NI 0. N -t A H CH 0 LO Z •• •-1 CD 95 > a) (I.) (L) r-I - N L(1• Ei A N H 0 A H • a) • "1-4 H HH r- p4 0 a) 0 Z 0 C/2 • r1 K1 P4 E-I c- N P E-I 0 \ 9 O pa Cr2 •=4 if\• C1/4.•1 0 I-1 4-1 a) o-1 a) a) N 0 39 CO 0 .0 H H N. 0• CD0 C N --t ..:-. .0 Forage Crop Breeding and Genetics Rod. V. Frakes Oregon Agricultural Experiment Station It is hoped that selection will identify alfalfa plants that can be made available to public and private alfalfa breeders interested in breeding for weevil tolerance. Forage breeding research at Oregon State University is concerned primarily with tall fescue, orchardgrass, alfalfa and birdsfoot trefoil. Currently, eighty varieties of alfalfa are being evaluated for forage production in the Willamette Valley. The 1976 harvest season will represent the third production year for this test. Alfalfa varieties are being released and marketed faster than they can be evaluated by the experiment stations. In an effort to learn about germplasm response in alfalfa producing areas, we are involved in an uniform testing program for the evaluation of certain germplasm. To date, the same germplasm is established at eleven location; Corvallis, Medford, Hermiston, Burns, Redmond, Caldwell, Aberdeen, Kimberly, Moscow, Prosser, and Puyallup. Data from this study will be of value in making inferences about potential adaptation of newly released varieties, provided the parental germplasm source is known. Thirty-eight varieties and experimental lines of birdsfoot trefoil are being evaluated for forage production in the It is suggested Willamette Valley. that in trefoil the first harvest of the year can be delayed until after spring rains, without a loss in forage quality. This is being tested during the 1976 season by sampling several alfalfa and trefoil varieties periodically from early spring until early July, to determine the relative change in percent protein, carbohydrate level, and digestibility, as the two legume species age. Fifty-two orchardgrass varieties and sixty-eight tall fescues are being evaluated for forage production and rated for seed producing potential. Part of the breeding program objective is to cooperate with grass breeders in the consuming areas by evaluating selected germplasm for seed production in Oregon. Twenty-five tall fescue genotypes from a private breeding program in Indiana, and twenty-one from the public program in Alabama are being examined for relative seed producing characteristics. Eighteen genotypes of Phalaris aquatica from Alabama, two experimental lines of Pennisetum from North Carolina, and twenty hybrid orchardgrasses from a private program in Minnesota are also being evaluated. In a cooperative study with the United States Department of Agriculture, we have selected desirable genotypes from long established alfalfa fields, in an attempt to identify plant material tolerant to natural pests. These selected plants were cross pollinated, and the progeny again crossed to elite alfalfa germplasm from the USDA program. Seed increase of three experimental lines were carried out by O. J. Hunt in Reno, Nevada and they are now ready for testing in the area of selection. The alfalfa weevil continues to be a major problem in certain areas of our state. Several thousand alfalfa plants representing germplasm believed to have some resistance to other alfalfa pests, have been established in weevil invested fields at Klamath Falls and Redmond. The breeding and genetics of tall fescue has been of primary interest in our forage breeding program. Fawn tall fescue, a synthetic variety with eight parent plants was released twelve years ago. 40 We recently initiated an experiment to determine (a) the influence of regrowth period, and season of growth on perloline content in tall fescue, (b) the effect of stage of growth, and plant part at maturity on the the genetic perloline level, (c) differences and level of heritability of perloline content. Data from this research will be used to develop a selection procedure for identifying tall fescue genotypes to be used in public and private breeding programs. Over five million pounds of Fawn seed It is a are produced annually. variety with good seedling vigor, higher forage yield in the spring and fall, and growers have suggested it may be more acceptable to the animal than other tall fescue varieties. During the 1975 season, however, Fawn was severely damaged by stem and leaf rust and it did not produce as well as other varieties in our forage production tests. Plant breeders have been successful in crossing tall fescue with ryegrass, and selecting plants that resulted in a stable variety. Kenwell is one of these, and it seems to be producing well in Oregon and other areas of the United States. This is a USDA variety, to be released this fall, and seed of it should be available to growers within a couple of years. Nearly one hundred million pounds of tall fescue seed are distributed each year. Most of it is used in the transitional zone for warm vs. cool season grasses; an area centered around Missouri and Kentucky. One of the reasons for its high use is the ability for the species to respond relatively well to adverse soil and climatic conditions. Tall fescue has high carrying capacity, but has been critized for its low palatability and low animal performance, when performance is measured in terms of daily gain. It has been observed that the alkaloid perloline is present in relatively high concentrations in tall fescue. Scientists of the University of Kentucky have experimental evidence to suggest that perloline level in tall fescue may be sufficiently high to reduce cellulose digestion within the rumen, causing a reduced rate of digestion, which could contribute to lower daily gain. 41 convincingly indicated that vaccinated sheep had fewer and less severe cases of foot rot than unvaccinated controls, although the vaccine failed to comp letely prevent the disease. CURRENT STATUS OF RESEARCH ON FOOT ROT IN SHEEP John A. Schmitz, D.V.M., Ph.D. Joseph L. Gradin, M.S. School of Veterinary Medicine Oregon State University Isolation and Classification of the Foot Rot Organism The primary research effort at Oregon State University has been directed toward the isolation and characterization of the B nodosus strains infecting sheep in Oregon. Sheep foot rot is a highly contagious disease and, considering the ease with which B nodosus is transmitted from a carrier to a susceptible sheep, it is a surprising phenomenon that the organism is extremely difficult to grow under laboratory conditions. Australian workers have sent cultures of B nodosus to scientists all over the world and the recipient laboratories have frequently had difficulty in successfullypropagating the organism. Approximately two years of culture attempts passed at OSU before the first isolation of B nodosus was made. However, the succeeding two years have been considerably more productive with 42 strains of B nodosus being isolated from 13 different flocks of sheep. During this time, a special medium has been developed that will inhibit the growth of unwanted bacteria while at the same time allowing the growth of B nodosus. This medium has greatly facilitated the isolation of B nodosus from samples collected from infected feet. Introduction During the past three decades, the major research effort on the cause, prevention and control of foot rot has been conducted in Australia and New Zealand. Investigators in these countries have quite conclusively shown that foot rot is caused by the combined activity of at least two anaerobic (oxygen sensitive) bacteria, namely, Bacteroides (formerly Fusiformis) nodosus and Fusobacterium necrophorum (formerly Sphaerophorous necrophorous). Numerous additional microorganisms have been incriminated and their significance is still not completely elucidated. Based on the Australian work which demonstrated the etiologic roles of Bacteroides (B) nodosus and Fusobacterium necrophorum, the emphasis of the foot rot research at Oregon State University has been toward the isolation and characterization of the B nodosus strains that infect sheep in Oregon. Australian workers have shown that while Fusobacterium necrophorum is readily available in any sheep environment (apparently derived from feces), B nodosus can persist only in the tissue making up the hoof wall. It has been demonstrated experimentally that B nodosus can survive in the environment, that is, in soil etc., for only ten days to two weeks. Based on this information research on the prevention of foot rot has been directed toward controlling the B nodosus infection by use of a vaccine. Reports from Australia, New Zealand and England on a B nodosus vaccine quite Work is presently in progress on the B nodosus isolates to determine their antigenic (chemical) make-up. Based on this information, the isolates can be classified into seperate groups, or "serotypes" of organisms, with identical or similar composition. Recent work in Australia has shown that for a foot rot vaccine to be successful it is important to produce the vaccine from a 42 The vaccine used in this field trial was produced by an American company using the same B nodosus strains used to produce the commercial Australian vaccine. The trial was conducted as a "double-blind" study in which 500 sheep were injected with the vaccine and 505 sheep were injected with placebo (a fluid that looked like vaccine but in reality was only colored water). The feet of all sheep were examined seven times during the trial period and the presence or absence of foot rot was recorded. Since the sheep were vaccinated and inspected by coded numbers, there was no knowledge as to whether the sheep were injected with the vaccine or the placebo, and the judgement of the foot examination was not prejudiced. If there was no foot rot, the foot was designated as "0" for that examination. If foot rot was present, a grade was assigned to indicate the severity of the case--a "1" denoted an early, very mild case while a "4" denoted the most severe lesion where there was severe underrunning of the sole and separation of the wall of the hoof. Some cases, such as chronics, hoof deformity or recently healed cases did not fit into the "1-4" grading system and were arbitrarily assigned an "X" grade. Eight flocks of sheep were utilized in the in the field trial. strain of B nodosus that is identical with or cross-immunizes against the serotypes infecting the particular flock of sheep. Thus far, the major B nodosus isolates at OSU have been classified into either of two serotypes. Besides these two major serotypes other minor serotypes have been identified. Only five of the isolates belong to any of the major serotypes identified in Australia thus verifying dissimilar antigenic composition between B nodosus strains in the United States and Australia. Vaccination Against Foot Rot The relationship between antigenic composition (serotype) of a strain of B nodosus and its immunizing ability against different serotypes is not well understood. Vaccination trials in Australia have shown that their serotype "A" will immunize against serotypes "A" and "B" but not against serotype "C" and certain "untyped" strains. Presently, these relationships can be determined only by vaccine trials--prolonged, tedious and expensive endeavors. Such trials have not yet been conducted with OSU isolates. Foot rot vaccines are produced commercially in Australia, New Zealand and England but have not been authorized in the United States because there is no proof that the vaccine is effective under U.S. conditions. Two field trials of Australian-produced foot rot vaccine, one in California conducted by Dr. Blaine McGowan and the other in Oregon by Dr. Stanley Snyder, have failed to demonstrate any reduction in severity or incidence of foot rot in vaccinated sheep as compared to nonvaccinated sheep. A summarization of the vaccine field trial is given in Table I. In compiling this summary the five examinations, which were conducted following the second of the two vaccinations, were considered. If a sheep had a "4" grade foot rot lesion in any foot at any of the five postvaccination examinations, it was placed in the "4" category in Table I. Similarly, if the most severe foot rot lesion during this period was a "1", "2" or "3", the sheep was designated in the appropriate category in the summation table. The "X" grade was arbitrarily placed between a "2" and a "3" when deciding the most severe lesion, that is, if a particular sheep had both "2" and "X" grades, the Foot Rot Vaccine Field Trial A foot rot vaccine field trial was conducted by the authors in 1974-1975. 43 sheep was designated "X" in the table, if on the other hand it had grades "3" and "X", it was designated "3" in the table. The results of this vaccine field trial indicate that the vaccine was not totally effective in preventing foot rot in any flock, indeed in five of the flocks there appeared to be no difference in the incidence or severity of foot rot between the vaccinates and nonvaccinates. However, in three of the flocks (B, D, and H, Table I) there appeared to be considerable reduction in both the incidence and severity of foot rot in the vaccinated sheep. The results in these three flocks give hope for ultimate success and encouragement for continued research. Perhaps the three flocks beneficially affected by the vaccine all contained the same serotype of B nodosus which crossreacted with B nodosus strains in the vaccine. Perhaps one or two strains of B nodosus will cross-immunize against all or most of the B nodosus strains in Oregon. Only diligent research will answer these and many other questions relating to one factor of improved sheep production--that of prevention and control of sheep foot rot. * ***** * ***** * The authors gratefully acknowledge the assistace of the following men in the vaccination trial: Don Gnos, Irvin Jones, Jack Kalina, Ed and Roger Leabo, Jeff Rice, Wayne Swango, Claude Swanson, Wayne Townsend and Alfred Wheeler; also the technical assistance of Chris Fredricks and Sheryl Nied. ************** 44 -1C 0 a co c....) em w L.L. 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Wing Introduction 7 months of age were bred to a Montadale ram to decrease lambing difficulty. The mating scheme is illustrated in table 1. Observations were made on fertility, prolificacy, wool production, lamb growth rate, longevity and general health of ewes and lambs. Abbreviations used in the text are Col for Columbia, Finn for Finnish Landrace and Hamp for Hampshire. Crosses are designated with the sire breed first and the ewe breed or breeds second (Hamp X Finn, Col would be an animal which is 1/2 Hampshire and k Finnish Landrace and 1/4 Columbia and resulted from mating a Hampshire ram with a 1/2 Finn and 1/2 Columbia ewe). Increasing production of farm flock sheep has been a high priority research item in recent years. Farm flock sheep are kept under rather confined and hence controllable conditions. The opportunity to manage for maximum production potential is greatly enhanced. With the introduction of the Finnish Landrace breed in 1968, increasing the number of lambs produced per ewe became more feasible. Besides her high twinning rate, the Finn ewe is a good mother and produces a good supply of milk. Finn lambs are quite hardy at birth and are known for their "livability". However, Finn sheep produce poor quality wool, gain weight slowly and produce carcasses of poor conformation and cutability. Through a crossbreeding program it is hoped that strengths and efficiencies of the Finn and of available breeds can be combined to produce a ewe with acceptable wool production and quality that will produce more lambs with the capability to gain fast and produce good carcasses. Results and Discussion The performance of ewes as yearlings and 2-year-olds is presented in table 2. Crossbreeding had the greatest impact on production when the ewes lambed as yearlings. Finn X Col ewes had the highest conception rate as yearlings and Columbias had the lowest. All ewes that failed to lamb as yearlings were culled. Conception rates were more uniform when the ewes lambed as 2-year-olds. Experimental Procedure The limited data collected thus far indicate that inclusion of Finn blood has increased the number of lambs born per ewe. As with conception rate, the greatest increase in lambs born per ewe occurred when the ewes were yearlings. Triplets occurred only in the Finn X Col group. As 2-year-olds the 1/2 Finn ewes produced more triplets (25% and 20% for the 2 groups); additionally, one set of quadruplets was produced. However, triplet production results in more problems than the added lamb is worth. Triplet lambs are smaller than twin or single lambs and have a lower survival rate at birth. Also, Grade Columbia ewes formed the base population for the rotational crosses to be described. They also served as straightbred controls. In the fall of 1972 and 1973, 40 Columbia ewes were randomly allotted into two groups of 20 each which were bred to either a Hampshire or a Finn ram. The female progeny were bred at 7 months of age to a ram of the third breed. The resulting three-breed-cross females were then bred to a ram of the most distantly related breed. Concurrently, straightbred Columbia matings were made on mature ewes. Ewes bred at 46 Table 1. Crossbreeding scheme. Line 1 Hampshire Line Line II Finn Line *H X C F F X(1- H L c 2 C X I C2 H C F 1 Line III Straightbred Control C c) 12F2 4-21 H J. C X C 1972 breeding C X C 1973 lambing 1973 breeding F 1974 lambing 1974 breeding 4, 5 1 1 5 1 ) HX(-8-C-4-Fe-H) FX(7 C-4-H-g F FXrH—F4CHX o■ 16 16 F 16 , 4' 1975 lambing 1975 breeding 2 u 16 - 1976 lambing 1976 breeding 1 18 u r), ,(18 u 9 55 r) 432 L. 32 " 32 ' 32 " 32 3 , C X C *H = Hampshire F = Finnish Landrace C = Columbia --Ram breed is listed first. 47 C X C 1977 lambing 1977 breeding rn 0) 10 to u-) CO 0) 0 01 0 CO NHOC) H H H H 01 • N 0 • CO 10 . N ti CO 1.0. lO • 0 CO • 01 0. CO I I I I cc) • 1 I I I I I I co H ti CO 0 LO c0 l0 N 0) CO it) c0 0 In 0 0 H rl 0 0 cq c0 0 • co• • CR • CO -P 43 4-) E--1 Ca 0) 4-, • •r1 • cJ CO • o) E-1 • I LO • 01 0) co • N. I I N in di 0 0 0 0 0 0 0 0 CO N . ID 0) 4-) N N 0) H N di N 0) 0 0 0 0 c\i H in N 0 L0 di 0 (1) 0 •ri P LO H CO N 4) a, Fi 04 O 4-) /4 g m of H NHN H H H l0 H 01 H 0 N CO H N P-1 F-1 0 4-) 4-) • a, (D O 0 • O p 0 00 00 co 0 0 co 00 00 10 0 0 0 0 0) 0 0 0 0 rl H H H 0 0 r0 H Si 0 4-) (L) H 4-) 04 .0 0 0) 1-4 •r4 T3 a) al 3 CD ti 0 0 H A 48 one lamb of a set of triplets has to be removed from the ewe either fostered or raised artificially, increasing costs. All of the crossbred groups produced more twins as yearlings than did the Columbias. Ewes that were 1/2 Finn produced more twins than did 1/4 Finn ewes. As 2-year-olds the 4 Finn ewes produced the highest incidence of twinning (86%) but not any triplets. This is the ideal result of the inclusion of Finn breeding if it is repeatable in coming years and generations. Yearling ewes that were 1/2 Hamp produced the biggest lambs at birth even though the Finn X Col ewes had been bred to Hamp rams and the Finn X Hamp, Col ewes had been bred to Columbia ra ms. Finn breeding in the ewe apparently influences birth weight. All lambs born from Finn X Col yearling ewes gave birth unassisted even though lambs were 1/2 Hemp. (Hamp Col); Hamp X (Finn Col); and Montadale X Col lambs were compared in another study. The lambs in the second study were from yearling ewes. Columbia ewe lambs at the Union Station are normally bred to a ram of a smaller breed to decrease the incidence of lambing difficulty. In the first study lambs were weaned at about 65 days of age and put in the feedlot. Hamp X Col lambs were about 10 lb. heavier at weaning than were lambs of the other 2 groups (table 4). Finn X Col and Columbia lambs performed about the same during the suckling stage. Once in the feedlot all 3 groups of lambs performed equally well. Hamp X Col lambs had the heaviest weight at slaughter because of their greater suckling gains. In the second study the lambs were weaned when the average weight per group was approximately 50 lb. Lambs were then put in the feedlot and fed until finished. Montadale X Col lambs did not gain as well during the suckling period or in the feedlot as the three-breed-cross lambs. Carcasses of all groups in both studies were of similar quality. Wool production was decreased by the addition of Finn breeding (table 3). Thus far the reduction in production has been greatest in the 3-breed crosses. Hamp X Col ewes did not show a decrease in pounds of wool produced. Only yearling wool production is presented. All crossbred groups have slightly lower wool grades than the Columbias. As the number of crossbred lambs in the experiment increases,selection pressure may be applied so that the poorest wool producers are culled. The decrease in wool production due to crossbreeding then could be lessened. Summary Use of the Finn in a crossbreeding program has effectively increased the number of lambs produced per ewe. The greatest overall increase in fertility and prolificacy due to crossbreeding occurred when the ewes lambed as yearlings. However, Finn X (Hamp, Col) ewes performed better for reproduction than ewes of all other groups when they lambed as 2-yearolds. Finn X (Hamp, Col) ewes produced a large number of triplets which are somewhat harder to manage. However, 4 Finn ewes produced the greatest incidence of twins. Wool production was decreased by crossbreeding, but a careful selection program should rectify this in a few generations. Crossbred lambs performed In 1974, wether lambs from straight and crossbred ewes were weaned and put into the feedlot. Weaning weights, daily suckling gains and daily feedlot gains were recorded. The feedlot lambs were separated into 2 studies. Columbia, Finn X Col and Hamp X Col lambs from mature ewes were compared in one study. Finn X 49 well in the feedlot. Ewes with Finn breeding tend to be as hardy as contemporary ewes. Based on data gathered to date, use of the Finnish Landrace in a crossbreeding program can increase the number of lambs produced. Ewes of k Finn blood seem to be the optimum thus far because of the higher incidence of twinning but lower frequency (compared to 1/2 Finn) of triplets. Table 3. Yearling wool production and grade by breed. Ewe Breed Ave. An. Wool Prod Ave. Yearl. Wool Grade Columbia 11.2 54.0 Hamp X Col 12.8 52.3 Finn X Col 9.8 52.6 Hamp X Finn, Col 7.6 53.1 Finn X Hamp, Col 7.5 50.2 Table 4. Feedlot performance of lambs. Lamb breed Weaning Weight Ave. Daily Suckling Gain. Final Weight Ave. Daily Feedlot Gain Study 1 Columbia 57.3 0.55 99.8 0.83 Finn X Col 58.1 0.55 100.8 0.83 Hamp X Col 67.7 0.64 110.7 0.83 Mont X Col 51.2 0.40 100.8 0.50 Finn X (Hamp Col) 59.6 0.57 111.0 0.65 Hamp X (Finn. Col) 56.0 0.57 101.6 0.61 Study 2 50