Color Vision, Part 2 Lecture 9. Thurs, 10/01 Jonathan Pillow Perception (PSY 323), Fall 2009 The University of Texas at Austin Basic Principles of Color Perception energy an illuminant with most power at medium wavelengths (i.e., a greenish light source) 13 measurements of power spectrum (example) Basic Principles of Color Perception energy an illuminant with power at all visible wavelengths (a neutral light source, or “white light”) 13 measurements of power spectrum (example) Basic Principles of Color Perception Q: How many measurements of this same spectrum does the human eye take (in bright conditions?) Basic Principles of Color Perception photoreceptor response Q: How many measurements of this same spectrum does the human eye take (in bright conditions?) A: Only 3! One measurement from each cone class Basic Principles of Color Perception photoreceptor response Q: How many measurements of this same spectrum does the human eye take (in bright conditions?) A: Only 3! One measurement from each cone class Think of each of these curves as a “receptive field” (over wavelength) for a single cone • tell how to “add up” the energy from different parts of spectrum in order to generate cone response Basic Principles of Color Perception photoreceptor response Q: How many measurements of this same spectrum does the human eye take (in bright conditions?) could also call this axis “absorption” or “sensitivity” A: Only 3! One measurement from each cone class Think of each of these curves as a “receptive field” (over wavelength) for a single cone • tell how to “add up” the energy from different parts of spectrum in order to generate cone response More terminology: photoreceptor response absorption spectra - describe response (or “light absorption”) of a photoreceptor as a function of frequency Absorption spectrum for “L” (red) cone A single photoreceptor doesn’t “see” color; it gives greater response to some frequencies than others single cone absorption spectrum single cone absorption spectrum • All the photoreceptor gives you is a “response” • Cant tell which light frequency gave rise to this response (blue or orange) Problem is actually much worse: cant tell a weak signal at the peak sensitivity from a strong signal at an off-peak intensity single cone absorption spectrum +0.5 spectral power +1 • Response to a +2 single-wavelength light source is the power times the absorption spectrum curve. (Just like the receptive fields we considered earlier) • All three of these lights give the same response from this cone Problem of univariance: An infinite set of different wavelength–intensity combinations can elicit exactly the same response from a single type of photoreceptor single cone absorption spectrum +2 • Therefore, one +0.5 spectral power +1 type of photoreceptor cannot make color discriminations based on wavelength So a single cone cant tell you anything about the color of light! Colored stimulus Response of your “S” cones sensitivity 1 0.8 0.6 0.4 0.2 0 400 450 500 550 600 650 700 energy illuminant #1 wavelength cone responses: 40 175 240 sensitivity 1 0.8 0.6 0.4 0.2 0 400 450 500 550 600 650 700 energy illuminant #1 wavelength cone responses: 40 175 240 sensitivity 1 percept 0.8 0.6 0.4 0.2 0 400 450 500 550 600 650 700 energy illuminant #1 wavelength cone responses: 40 175 240 sensitivity 1 percept 0.8 0.6 0.4 0.2 0 400 450 500 550 600 650 700 energy illuminant #1 #2 wavelength cone responses: 40 175 240 sensitivity 1 percept 0.8 0.6 0.4 0.2 0 400 450 500 550 600 650 700 energy illuminant #1 #3 #2 wavelength cone responses: 40 175 240 sensitivity 1 percept 0.8 0.6 0.4 0.2 0 400 450 500 550 600 650 700 energy illuminant #1 #4 #3 #2 wavelength cone responses: 40 175 240 sensitivity 1 percept 0.8 0.6 0.4 0.2 0 400 450 500 550 600 650 700 illuminant #1 energy Metamers #4 #3 #2 wavelength - Illuminants that are physically distinct but perceptually indistinguishable Implication: tons of things in the natural world have different spectral properties, but look the same to us. Implication: tons of things in the natural world have different spectral properties, but look the same to us. But, great news for the makers of TVs and Monitors: any three lights can be combined to approximate any color. Single-frequency spectra produced by (hypothetical) monitor phosphors illuminant #1 energy Monitor phosphors produce “metameric match” to illuminant #1 (or any other possible illuminant). wavelength Close-up of computer monitor, showing three phosphors, (which can approximate any light color) This wouldnt be the case if we had more cone classes. hyperspectral marvel: mantis shrimp (stomatopod) • 12 different cone classes • sensitivity extending into UV range This wouldnt be the case if we had more cone classes. hyperspectral marvel: mantis shrimp (stomatopod) • 12 different cone classes • sensitivity extending into UV range • No surprise that they never invented color TV! Real vs. Conterfeit $$ Output of hyper-spectral camera (colorized artificially) Worth knowing about if you end up seeking employment in the “informal” sector following graduation. (or law enforcement) Color vision photoreceptor response Our color vision relies on comparing the responses of three cone classes Cone responses entirely determine our color percepts: S M L 100 100 100 50 50 50 0 0 0 Cone responses entirely determine our color percepts: S M L 100 100 100 50 50 50 0 0 0 100 0 0 0 100 0 0 0 100 Cone responses entirely determine our color percepts: S M L 100 100 100 50 50 50 0 0 0 100 0 0 0 100 0 0 0 100 100 100 0 0 100 100 100 0 100 How did they figure this out? -3 “primary” paints -color mixture of lights: any color can be made three suitable lights... How did they figure this out? Late 17th Century: Isaac Newton Late 17th Century: Isaac Newton “The rays themselves, to speak properly, are not coloured” Newtons Spectrum: R O Y G B I V Newtons Theory: seven kinds of light -> seven kinds of photoreceptor James Maxwell: color-matching experiment Given any “test” light, you can match it by adjusting the intensities of any three other lights (2 is not enough; 4 is more than enough) Trichromacy Color space: A three-dimensional space that describes all colors. There are several possible color spaces (i.e., different coordinates for describing the same space) • RGB color space: Defined by the outputs of long, medium, and short wavelength lights • HSB color space: Defined by hue, saturation, and brightness Hue: The chromatic (color) aspect of light Saturation: The chromatic strength of a hue Brightness: The distance from black in color space •hue around the edge •saturation increasing from center to edge Trichromatic color vision is called the Young-Helmholz theory of color vision • Thomas Young (1773–1829) and Hermann von Helmholtz (1821–1894) independently discovered the trichromatic nature of color perception • This is why trichromatic theory is sometimes called the “Young–Helmholtz theory” • James Maxwell (1831–1879) developed a colormatching technique that is still being used today Notice the variability between individuals! Young-Helmholz theory of color vision However, theory doesnt completely explain everything (E.G. Why is red the after-image of green?) response Some Retinal ganglion Cells have center-surround receptive fields with “color-opponency” space M L M M M response M M wavelength Red-Green (L-M) Color-Opponent cell Opponent Channels: • L-M (red - green) • S - (L+M) (blue -yellow) • L+M - (L+M) (black - white) Opponent Processes Opponent color theory: perception of color is based on the output of three mechanisms, each of them based on an opponency between two colors: Red– green, blue–yellow, and black–white • Some LGN cells are excited by L-cone onset in center, inhibited by M-cone onsets in their surround (and vice-versa) • Red versus green • Other cells are excited by S-cone onset in center, inhibited by (L + M)-cone onsets in their surround (and vice-versa) Blue versus yellow Opponent Processes Afterimages: A visual image seen after a stimulus has been removed Negative afterimage: An afterimage whose polarity is the opposite of the original stimulus • Light stimuli produce dark negative afterimages • Colors are complementary. Red produces green afterimages and blue produces yellow afterimages (and vice-versa) • This is a way to see opponent colors in action