Study Guide Evolution of Animals Chapter – 19 Animalia – Fig 19.1 and Table 19.2 are important. I will ask questions on main animals in each phylum. Concentrate on phylum characteristics or general characteristics of animals, protostomes, deuterstomes and chordates. 1. Trends in evolution of animals a) Symmetry Fig 19.5 – sessile or slow moving animals had radial symmetry (body parts arranged around a vertical axis) because food can come from any direction but most animals evolved Bilateral Symmetry (body parts arranged on 2 sides of a vertical plane because this favored in having a fixed anterior end b) Cephalization – Animals evolved head at anterior end because when moving to search for food, shelter or mate they required maximum information around them – this led to evolution of head having most sense organs to observe and brain to process and finally also mouth to feed inside head itself c) Fig 19.8 Coelom is the space between body organs =viscera and body wall (skin + muscles) lined by peritoneal membranes d) Complete digestive system has mouth to feed and anus to defecate e) Segmentation is the internal divisions of muscles, nervous tissue and excretory organs, also depicted externally in most animals. 2. Porifera = Sponges are the simplest animals. Fig 19.2. Sponges may have asymmetric or radial symmetry. No true tissues or organs evolved in them. Epidermal cells line the surface Collar cells establish water current and capture food with flagella. It is called Filter Feeding. Amoebocytes lie in between the 2 layers and secrete Spicules – spine like skeletal structures formed of lime or silica or sponging fibers. No nerve cells evolved yet. 3. Glass Sponges have spicules of silica and look like glass in a dead organism, found in deep water. 4. Cnidaria fig 19.3 (jelly fishes, corals) have a characteristic stinging cell nematoblast = cnidoblast. Most of the times it paralyses the prey but may trap it. 2 main body layers with a jelly layer in between. True tissues and organs evolved including the nerve cells. Incomplete digestive system and radial symmetry is observed in cnidarians. 5. 2 Main Body Forms = Polyp and Medusa - Polyp – has a cylindrical body with a mouth and a ring of tentacles armed with nematocysts surround the mouth. Incomplete digestive system because no anus present. Mostly polyp is sessile and medusa is motile. Medusa has an umbrella like structure, mouth is at the end of stick and tentacles cover the rim of umbrella. Fig 19.4 6. Physalia = Portugese man of war – has an air sac to float and has both specialized polyps and medusa in the colony. It can trap preys like fish and others with its long tentacles. 7. Aurelia = Jelly fish has a large medusa, mouth is surrounded by 4 arms, 4 horse-shoe shaped gonads having either eggs or sperms, can be seen through the bell. Polyp is reduced or absent in the life c 8. Sea Anemone = Metridium has advanced popyp with several rows of tentacles and gastrovascular cavity divided by partitions. 9. Corals are advanced polyps, mostly colonial, and deposit lime around them and look like stone having pores. When living each pore has a single individual formed of a polyp. Most corals harbor 1 algae in them and can grow better in clear waters. Pollution of oceans is resulting in elimination of corals from large areas of sea-beds. 10. Platyhelminthes = flat worms – Thin long bodies, bilateral symmetry, cephalization initiated, incomplete digestive system with only mouth, space between internal organs and body wall filled = Acoelomate, bisexual = monoecious animals. 11. Planaria – a member of free-living flat worms, head has 2 eye-spots, 2 auricles – lateral extensions of head, mouth in the middle, branching gut (digestive system), slow moving bottom dwellers which feed on small animals or dead animals. Fig 19.7 12. Parsitic flat worms include flukes and tapeworms. 13. Flukes are parasites living in the liver, intestines, lungs or blood vessels of vertebrates. Oral sucker around mouth and ventral sucker on ventral side help fluke to attach host and feed on the tissues and blood. Fresh water snail is an intermediate host in life cycle of most flukes. 14. Tape worm - It does not have any digestive system, absorbs food through body surface (is absorptive instead of normal Ingestive mode of animals). It is very long, has a very tiny scolex with hooks and suckers to attach to intestine. 15. Nematoda = Round worms – are amongst most widely distributed and most abundant animals found everywhere including soils, water, marine or fresh, animal and plant tissues. You observed the vinegar eels in undistilled vinegar and some are found in undistilled beer. Most round worms are free living but many are human parasites – Ascaris. Fig 19.9 16. Complete digestive system – 1st to develop tube within tube body plant and a complete digestive system. Pseudocoel – the false body cavity is present but is not lines by peritoneum membranes. Parasitic round worms lack attaching organs like suckers or hooks found in parasitic flat worms. Round worms are dioecious – sexes are separate, males are smaller and narrow and females are longer and broader. 17. In Protostomes, Mouth (1st mouth) develops from blastopore. Fig 19.15. Protosomes include Flat worms, Round worms, Rotifera, Mollusks, Annelids and Arthropods. 18. Phylum Mollusca has animals with soft body dorsally covered by a skin fold called Mantle and have a single ventral foot. In majority of mollusks mantle secretes a shell. Most have a tongue like radula with rasping teeth. Fig 19.10 2 class shell Feed on Foot gastropods Conical spiral Most on plants/algae Extensible muscular bivalves 2 similar hinged parts Planktons, filter feeder Hatchet like, used for burrowing Body parts Head, tentacles, radula, jaws snails No head or tentacles or radula cephalopods Mostly reduced predators Divided into arms / tentacles fixed on head Head, radula, eyes, Clams, oysters, scallops Squids, octopus, nautillus example 19. Phylum Annelida includes segmented animals with coelom the true body cavity. Annelids have well developed digestive, circulatory, nervous and reproductive systems. But use skin as the respiratory system. Some of them have external gills. They have stiff bristles called setae or chaetae for locomotion. Fig 19.11 class Locomote with Live in Feed on character Sexes Neries (polychaetes) parapodia Marine water Small animals, trap planktons Tentacles on head separate Earthworms (oligochates) setae Burrowing in land Organic matter rich soil Leeches (Hirudinea) Oral and posterior sucker Water or land Blood, are ectoparasites Permanent clitellum bisexual Temporary clitellum bisexual 20. Phylum Arthropoda has animals with exoskeleton of hard plates of chitin and jointed appendages. These are the largest group of animals and were 1st to colonize land. Most have a respiratory system of cuticular tubes called Trachea which directly carry oxygen to the tissues. Fig 19.13-14 class Antenna Body regions Walking legs Live in Respiration by example crustaceans 2 pairs Cephalothorax and abdomen 5 pairs arachnids absent Cephalothorax And abdomen 4 pairs insects 1 pair Head, thorax and abdomen 3 pairs water gills Cray fish, crab, lobster, shrimp On land Book-lungs Spiders, scorpion, mites On land trachea Butterfly, wasp, beetles, Bugs, flies millipedes 1 pair Head, thorax and abdomen 2 pair/ segments On land trachea Cylindrical body centipedes 1 pair Head and long body 1 pair / segment On land trachea Flat body 21. In Deuterostomes Anus (2nd mouth) develops from the blastopore. Deuterostome animals have 2 major animal Phyla – Echinoderms and Chordates. 3 22. Echinoderm (spiny-skinned) – a) radial symmetry in adult but bilateral larva b) bone like endoskeleton or test c) water vascular system of canals and tube-feet for locomotion and capturing food d) regenerate broken arms and other body parts. Fig 19.16 23. Sea Stars – have 5 thick arms with viscera extended into them and feed on Clams or Oysters. 24. Brittle Stars – have 5 thin arms without viscera. The arms show snake like movements and get easily broken but help them swim. 25. Sea Urchins – have very large spines coming out of test and possess Aristotle’s Lantern a feeding device to eat sea weeds or kelps. No arms are present 26. Sea Cucumbers – have cylindrical body and pick food with their tentacles or are filter feeders. No arms present. 27. Feather Stars – have branched 5 arms and remain attached most of the time but can swim freely when needed. 28. Chordates possess 4 characters at least once during their life time (in embryo or larva or adult): a) Notochord – an elastic chord, is replaced by vertebrae in most chordates called Vertebrates b) Dorsal and hollow nerve tube – develops into brain and spinal cord in vertebrates c) Pharyngeal Gill Clefts or pouches – are slit like openings in sides of pharynx and help in respiration in lower chordates. d) Post-anal Tail – is present in all chordates. Urochordate – Sea Squirt Notochord and nerve tube in tail of larva. Uro = tail Body covered by test with spicules. No head or eyes. Fixed to sea rocks Filter feeder Fig 19.17 Cephalochordate – Amphioxus Notochord extends into anterior end. Cephalo = head Fish like body has no head. Burrowed or free swimming Filter feeder Fig 19.18 Vertebrates –fish mammal Notochord of embryo or larva replaced by vertebrae Distinct head, eyes, fins or limbs Free swimming Mostly Predators or herbivores Fig 19.20 29. Vertebrates have jawless fishes Cyclostomes or Agnatha and Gnathostomates the vertebrates with jaws in mouth. 30. Agnatha – Cyclostomes are jawless fishes only having unpaired fins. The circular mouth act as a sucker a rasping tongue helps in feeding. Lamprey is an ectoparasite of sharks. Hag fish is slimy and usually enters dead or dying animals and feeds on the soft viscera. Cyclostomes lack jaws and paired fins/limbs of other vertebrates. 4 31. Gnathostomates include fishes – animals with paired fins and Tetrapods – animals with 2 pairs of limbs. Fishes live in water, respire with gills, skin covered with scales, have 2 chambered heart both paired and unpaired fins. Fishes have 2 main classes Cartilaginous fishes and Bony fishes. Fish Class Position of mouth Pharyngeal gill-slits Scales Tail fin Swim bladder – to regulate buoyancy For example Chondrichthyes=cartilaginous fishes Ventral, below the snout Exposed and easily seen Indistinct, small Lobes asymmetrical absent Sharks, rays, Fig 19.21 Osteichthyes = bony fishes terminal Covered by gill cover Distinct, large Lobes symmetrical or unlobed Present in many fishes, helps to gain or lose depth in water Salmon, Tuna, gold fish, Fig 19.22 32. Amphibians (amphi = both, bios = living, can breathe in air as well as water) are naked vertebrates lacking scales/feathers / hair. They use skin as respiratory organ and developed lungs. Amphibians moved freely on land but have to return to water to release eggs. Fertilization is external and eggs hatch to form larva. The larva is the 1st feeding stage and is fish like having gills. It undergoes metamorphosis to become adult amphibian. Frog Egg tadpole larva Frog. For example Frogs, Toads, Salamanders, mud puppies. Class Fishes Amphibians Reptiles Aves = birds Mammals Skin cover scales absent scales feathers hair heart 2 chambered 3 chambered 3 chambers 4 chambered 4 chambered Respiration by gills Skin, lungs lungs lungs lungs 2 pairs of fins legs legs legs legs Teeth used for Holding food holding holding Absent replaced by bill or beak Chewing food ears absent Ear drum at skin level Ear drum in Ear canal Ear drum in Ear canal Pinna outside ear canal Eggs Need water Need water Laid on land Laid on land Give birth 33. Reptiles developed the amniotic egg and became 1st true land vertebrates. Amnion is a fluid filled sac and protects embryo from desiccation (drying up) on land. This eliminated the need of external water, like amphibians, for fertilization. They have scales on skin to prevent water loss, and also have claws on digits and a 3-chambered heart. Eggs with lot of yolk and hatch into adults. No larval 5 stage present. Reptiles include turtles, lizards, snakes, alligators and crocodiles. Dinosaurs, the most famous extinct animals, belonged to reptiles. Birds evolved as a direct line from dinosaurs. 34. Birds are endotherms and maintain high body temperature and can live in extreme cold climates like Antarctica. The bones are very light but strong. Body has feathers and wings. Feet are covered with scales to indicate close relationship to reptiles. Birds also lay amniotic eggs with large yolk and no larva needed. Due to marked parental care for the eggs and hatchlings most birds lay only 2-3 eggs. The evolution of beak or bill helped them to specialize for different kinds of food. Sternum developed a high flat vertical bone, keel, to accommodate bulky flight muscles. Lungs are supported by air sacs to get oxygen even during expiration (breathing out). Due to brilliance of colors Huxley called them ‘Glorified Reptiles’. 35. Mammals – body covered by hair, mammary glands to feed milk, specialized teeth (incisiors for cutting; canines for tearing; premolars and molars for chewing), ear canal covered with Pinna, 7 vertebrae in neck. 36. Mammals include egg laying mammals, marsupials and placental mammals. 37. Monotremes = the egg laying (Oviparous) mammal – live only in Australia, lay eggs and feed hatchlings with milk. Duck bill platypus and spiny ant eater are only 2 examples. They are the connecting link between early reptiles and mammals. 38. Marsupials live only in Australian and South American regions. They give birth Viviparous) to premature babies and many carry them in an abdominal pouch = marsupium. Mammary glands are present in the pouch. Well known examples are Kangaroo, Koala and Opossum. 39. Placental Mammals – are most successful and most evolved animals including humans. Give birth (viviparous) to mature babies. Endotherms – maintain body temperature. Most live on land. Aquatic mammals include Whales, dolphins, seals etc and flying mammals are bats. In viviparous animals developing embryo draws nutrition and oxygen from mother’s blood and pass out wastes to it. Bear is omnivore but lion, tiger, wolf, cat and dog are carnivore mammals. Cattle, deer, zebra , giraffe are grazing animals. Monkeys, apes and humans are primate mammals. 6