Species Organism that has at least one characteristic that is not shared with any other organism. Characteristic Controlled by proteins Proteins Made of amino acids Order of amino acids Determined by a gene Gene Series of nitrogen bases in a DNA molecule Mutation Change in the nitrogen bases through any number of causes - Change gene - Change amino acid / sequence - Change protein - Change characteristic - Change organism Evolution •Change in gene frequency / species over time •Unifying theory of biology based on scientific observation •Occurs in populations – not individuals. Theories associated with evolution Gradualism Evolution in slow gradual steps Punctuated equilibrium short periods of rapid species formation followed by long periods of little evolution Coevolution Interactions between species can result in changes in the gene pool of both species Four Factors in Darwin’s Theory - all species reproduce in excess of the numbers that can survive - all organisms compete for limited resources - variations exist within a species - environment selects for the more advantageous variations Needed for evolution to occur 1. Means of increasing Variation Mutations – changes in nucleotide sequence of DNA Recombination – reshuffling of genetic material during meiosis Gene flow – gene frequency changes due to immigration from other populations 2. Means of decreasing variation Natural selection – organisms with less advantageous traits (genes) survive less often and leave fewer off-spring. Genetic drift – random events cause change in gene frequency (small Populations) founder effect population bottlenecks Gene flow – gene frequency changes due to emigration Microevolution Change in a population’s alleles over a period of time B. List and describe three possible outcomes of natural selection Snail coloration best adapted to conditions Average Natural selection Number of individuals Number of individuals Directional Natural Selection New average Coloration of snails Previous average Average shifts Coloration of snails Proportion of light-colored snails in population increases Figure 5-6 (1) Page 102 Slide 8 Light snails eliminated Dark snails eliminated Natural selection Number of individuals Number of individuals Stabilizing Natural Selection Snails with extreme coloration are eliminated Coloration of snails Coloration of snails Average remains the same, but the number of individuals with intermediate coloration increases Figure 5-6 (2) Page 102 Slide 9 Intermediate-colored snails are selected against Light coloration is favored Dark coloration is favored Natural selection Number of individuals Number of individuals Diversifying Natural Selection Coloration of snails Snails with light and dark colors dominate Coloration of snails Number of individuals with light and dark coloration increases, and the number with intermediate coloration decreases Figure 5-6 (3) Page 102 Slide 11 A. List limits of adaptation to change. - A change in environmental conditions can lead to adaptation only for traits already present in the gene pool of population - Even if a beneficial heritable trait is present in a population, that population’s ability to adapt can be limited by its reproductive capacity - Even if a favorable genetic trait is present in a population, most of the population would have to die or become sterile so individuals with the trait could predominate and pass the trait on. Summarize some common misconceptions about natural selection. Survival of the fittest is survival of the strongest Fittest is the measure of reproductive success not strength Evolution involves a “plan” that results in progressively more perfect species No plan or goal – only constant genetic adaptation to conditions Individuals will adapt to their environment Biological adaptations are the result of genetic characteristics not of conscious thought Individuals have no choice in their genetic makeup, therefore they either have the characteristic or not. Macroevolution Large changes recorded in the fossil record (large scale) Speciation: When a group of one species is cut off from the rest of the population and gene flow is stopped. Each group is acted on by local selective conditions. Kiabab Squirrel Abert Squirrel Conditions determine Characteristics •Related organisms becomes less alike •Distantly related organisms develop similar characteristics Observations that led to, and support evolution Fossils Any preserved remnant or impression left by organisms that lived in the past. Relative dating – superimposition of sedimentary rocks tells the relative age of fossils Absolute dating – dating using half-lives of radioactive isotopes (Potassium 40, Rubidium 87, Uranium 238, Carbon 14) Transitional forms – Archeoptryx and similar bird-like reptiles. Absence of some forms can also be used as an argument against evolution. Homologous structures Structures that share a common ancestry. Modified versions of the same structure. Vestigial structures Remnants of an organisms evolutionary past. The structure serves no major purpose at this time. DNA comparison Protein comparison