Chapter 12 The Earliest Dispersal of the Genus Homo: Homo erectus and Contemporaries Chapter Outline A New Kind of Hominid The Morphology of Homo erectus Who Were the Earliest African Immigrants? Historical Overview of Homo erectus Discoveries Technological and Population Trends in Homo erectus Interpretations of Homo erectus: Continuing Uncertainties Homo erectus First hominid to expand into new regions of the Old World. Existed over 1 million years. The behavioral capacities & morphological changes. Homo erectus African discoveries established Homo erectus by 1.8 m.y.a. Some researchers see anatomical differences: African fossils as Homo ergaster. H. erectus/ergaster represents a different grade of evolution than their African predecessors. Morphology of Homo erectus Brain size is related to overall body size. Body size dramatically increased. Cranium had a distinctive shape with a thick cranial bone and large brow ridges. Shovel-shaped incisors suggest an adaptation in hunter-gatherers. The Dmanisi Hominids Early 1990s discovery of the Dmanisi. The most informative specimens are four wellpreserved . Best-preserved hominids of this age outside of Africa Mixed pattern characteristics. The Dmanisi Hominids The Dmanisi crania have some similarities to H. erectus, while some characteristics are different from other hominid finds outside of Africa. The most complete specimen has a less robust and thinner browridge, a projecting lower face, and a large upper canine. All three Dmanisi crania have small cranial capacities. A number of stone tools, similar to early ones from Africa, have been recovered at Dmanisi. Questions Raised by the Dmansi Discoveries 1. 2. 3. Was Homo erectus the first hominid to leave Africa—or was it an earlier form of Homo? Did hominids require a large brain and sophisticated stone tool culture to disperse out of Africa? Was the large, robust body build of H. erectus a necessary adaptation for the initial occupation of Eurasia? Discoveries in Java Six sites in eastern Java have yielded all the H. erectus fossils found on this island. Dates range from 1.8 m.y.a. to 1.6 m.y.a. The Ngandong individuals date from 50,000 to 25,000 y.a. If the Ngandong dates are correct it would make Homo erectus and Homo sapiens contemporaries. In Java, no artifacts have been found that can be associated with Homo erectus. Discoveries in Peking “Dragon bones” used as medicine and aphrodisiacs were ancient bones. 40 male and female adults and children have been found near Zhoukoudian. The site was occupied for 250,000 years. 40% of the bones were from individuals less than 14 years old, 2.6% were from individuals between 50-60 years. Chinese Tools From Middle Pleistocene Sites Discoveries in East Africa Louis Leakey unearthed a fossil skull at Olduvai. An almost complete skull was discovered in east Turkana. The most complete H. erectus skeleton ever found was uncovered in west Turkana. In Ethiopia, an abundance of Acheulian tools have been found as well as a robust mandible dating to 1.3 m.y.a. East African Homo erectus East African specimens have thinner cranial bones than those found in Asia. Some scientists argue that the African and Asian erectus finds should be classified as separate species. The African and Asian populations are separated by more than one million years. Technological Trends in Homo erectus Expansion of the brain enabled H. erectus to develop sophisticated tools: Biface - stone that was worked on both sides and used to cut, scrape, pound, and dig. Thousands of Acheulian hand axes have been found with remains of large animals. Homo erectus is seen as a potential hunter and scavenger. Small Tools of the Acheulian Industry (a) Side scraper (b) Point (c) End scraper (d) Burin Trends in Homo erectus Homo erectus liked to travel. Stone tools found on the island of Flores, suggest that H. erectus constructed ocean-going vessels. Homo erectus embraced culture as a strategy of adaptation. Key Hominid Fossils Asia Site Dates (y.a.) Taxonomic Designation Java (6 locales) 1,8000,000– 25,000 Homo erectus China (6 locales; most significant is at Zhoukoudian); 400,000+– 200,000? Homo erectus Key Hominid Fossils Europe Site Dates (y.a.) Taxonomic Designation Ceprano 900,000– 800,000 Homo erectus 780,000? Quite likely not H. erectus, referred to by discoverers as “Homo antecessor” Gran Dolina (Atapuerca, Spain) Dmanisi (Republic of Georgia) 1,800,0001,700,000 Homo erectus/ Homo ergaster Key Hominid Fossils Africa Site Dates (y.a.) Taxonomic Designation Bouri(Ethiopia) 1,000,000 Homo erectus 1,600,000 Homo erectus, also frequently referred to Homo ergaster 1,800,000 Homo erectus, also frequently referred to Homo ergaster Nariokotome (West Turkana, Kenya) East Turkana(Kenya)