Biology ECO SC.912.L.15.13 Sophiaa Lopez, P2. • As powerful as natural selection is, unfortunately, it is sometimes misunderstood. A persistent misconception is that natural selection occurs mainly through differences between organisms in death rates, or differential mortality. Selection normally proceeds in a much more subtle and inconspicuous way. Whenever one organism leaves more successful offspring than another, in time its genes will come to dominate the population gene pool. Eventually, the genotype leaving fewer offspring must become extinct in a stable population, unless concomitant changes confer an advantage on it as it becomes scarcer. Ultimately, natural selection operates only by differential reproductive success. Differential mortality can be selective but only to the degree that it creates differences between individuals in the number of reproductive progeny they produce. Hence, phrases such as “the struggle for existence” and “survival of the fittest” have had an unfortunate consequence. Simpson Evolution Example Natural Selection Individuals with favorable traits are more likely to leave more offspring better suited for their environment Example: English peppered moth (Biston betularia) Peppered Moth Lab Peppered Moth Natural Selection Simulation MUTATION #1 A mutation can create a new allele in an individual. MECHANISMS When this happens, the OF population experiences a EVOLUTION change in its allele frequencies and, consequently, experiences evolution. Despite mutation’s vital role in the generation of variation, mutations almost always cause early death or lower the reproductive success of an organism. EVOLUTIONARY CHANGE: MUTATION Mutagen DNA Normal basepair sequence Mutated basepair sequence Normal protein Mutated protein Normal phenotype Mutated phenotype Mutations • Are rare because you have self correcting enzymes • Natural Process that produces genetic diversity • Not all mutations are bad – Some won’t affect the body at all • Blood types/ear lobes – Some are advantageous (thumb) five digit s wrist bone GENETIC DRIFT POPULATION BEFORE GENETIC DRIFT Allele frequencies: cleft chin (dominant) smooth chin (recessive) Neither allele is related to reproductive success. Inheritance is based solely on chance. POPULATION AFTER GENETIC DRIFT There are now more recessive alleles in the population than before. MECHANISMS OF EVOLUTION GENETIC DRIFT A population can experience random changes in allele frequency that do not influence reproductive success and, consequently, the population experiences evolution. REPRODUCTION In this example, a heterozygous couple (Cc) could have two children that are homozygous recessive (cc), causing an increase in the proportion of recessive alleles in the population. FIXATION Genetic drift leads to fixation when an allele’s frequency becomes 100% in a population. If this occurs, there is no longer genetic variation for the gene. GENETIC DRIFT: FOUNDER EFFECT SOURCE POPULATION Allele frequencies: 5 digits per hand (recessive) >5 digits per hand (dominant) A group of individuals may leave a population and become the founding members of a new, isolated population. NEWLY FOUNDED POPULATION The new population will be dominated by the genetic features present in the founding members. MECHANISMS OF EVOLUTION FOUNDER EFFECT The founding members of a new population can have different allele frequencies than the original source population and, consequently, the new population experiences evolution. AMISH MIGRATION (GENE FLOW) 1 BEFORE MIGRATION Two populations of the same species exist in separate locations. In this example, they are separated by a mountain range. Population 1 MECHANISMS OF EVOLUTION MIGRATION After a group of individuals migrates from one population to another, both populations can experience a change in their allele frequencies and, consequently, experience evolution. Population 2 MIGRATION (GENE FLOW) 2 MIGRATION A group of individuals from Population 1 migrates over the mountain range. Population 1 MECHANISMS OF EVOLUTION MIGRATION After a group of individuals migrates from one population to another, both populations can experience a change in their allele frequencies and, consequently, experience evolution. Population 2 MIGRATION (GENE FLOW) MECHANISMS OF EVOLUTION MIGRATION After a group of individuals migrates from one population to another, both populations can experience a change in their allele frequencies and, consequently, experience evolution. 3 AFTER MIGRATION The migrating individuals are able to survive and reproduce in the new population and they may experience evolutionary changes from population 1. Population 1 Population 2 Mechanism of evolution Natural Selection 1. VARIATION OF A TRAIT IN A POPULATION 3 Conditions that must occur for Natural Selection The tiniest dog in a litter has reduced differential reproductive success. Its more robust siblings prevent access to the food it needs to grow and thrive. 2. The trait must be inheritable 3. One version of the trait must be in greater abundance than a different version of the trait. EVOLUTION BY NATURAL SELECTION: A SUMMARY 1 VARIATION FOR A TRAIT Different traits are present in individuals of the same species 2 HERITABILITY Traits are passed on from parents to their children. MECHANISMS OF EVOLUTION NATURAL SELECTION When these three conditions are satisfied, the population’s allele frequencies change and, consequently, evolution by natural selection occurs. 3 DIFFERENTIAL REPRODUCTIVE SUCCESS In a population, individuals with traits most suited to reproduction in their environment generally leave more offspring than individuals with other traits.