Chapter 40: Basic Principles of Animal Form & Function Cell size limited by surface area :volume Cells must be in aqueous solution for nutrient & waste exchange tissues Groups of cells with common structure & function Types: Epithelial Cover body, line organs, & body cavities Tightly packed; held together by tight junctions Barrier against microbes, fluid loss, or mechanical injury Categorized by numbers of layers & shape Shape Cuboidal- dice-like Columnar- brick-like Squamous- floor tile-like Types: Simple epithelium Single layer “leaky”- allows for diffusion Ex. Alveoli, capillaries Stratified epithelium Multiple tiers of cells Glandular epithelia Absorb or secrete solutions Ciliated- lining of respiratory system Mucus membrane- oral cavity, nasal passages; moisten & lubricate Connective tissue Bind to & support other tissues Sparse number of cells scattered through an extra-cellular matrix Consist of 3 protein fibers Collagen: non-elastic; strength; resists stretching Elastin: rubbery; retain shape Reticular: branched to connect adjacent tissues Types of connective tissue Loose connective Fibroblasts- secrete proteins of extracellular fibers Macrophages- immune response Adipose Storage of fat For energy & insulation Fibrous connective Tendons- attach muscle to bone Ligaments- attach bone to bone Cartilage Part of all of skeleton in vertebrates Bone Mineralized connective tissue Blood Plasma- liquid matrix of water, salts, & proteins Leukocytes- white blood cells Erythrocytes- red blood cells Platelets- blood clotting Form in red bone marrow at ends of long bone Nervous tissue Sense stimuli & transmit signals within the animal Neuron=nerve cell specialized to conduct impulses dendrites= extensions that conduct impulses to the cell body Axons= extensions that transmit impulses away from the cell body Muscle tissue Capable of contraction when stimulated Consists of contractile proteins actin & myosin Types: Skeletal muscle Attached to bone by tendons Voluntary Striated pattern Cardiac muscle Contractile walls of heart Striated pattern Smooth muscle Lines walls of internal organs Involuntary Tapered, non-striated appearance Organ systems Are inter-dependant; organs with separate functions act in coordinated manner Tissues organized into organs except in simple organisms Some organs are layered (ex. Vertebrate stomach) Mesentery=sheets of connective tissue in which organs are suspended Bioenergetics Flow of energy through an organism Limits animal’s behavior, growth, & reproduction Determines food requirements Animals are heterotrophs Energy Input (ingested food) Digestion (enzymatic hydrolysis of food) Absorption (small energy molecules by body cells) Catabolism (cellular respiration & fermentation harvest chemical energy from food molecules) some Energy Stored in ATP some Energy Lost as heat is to surroundings Energy Used - chemical energy of ATP powers cellular work. After the needs of staying alive are met, leftover chemical energy and carbon skeletons from food molecules can be used in biosynthesis. Energy Lost - cellular work generates heat, which is lost to surroundings or used to maintain body temperature (endotherms) Metabolic rate Total amount of energy an animal uses to stay alive Measured in Calories (kilocalories) Calculated by measuring: oxygen used Amount of heat loss per unit of time Heat loss is by-product of cellular work Heat loss is measured with calorimeter Range of metabolic rates Minimal rate for life support (breathing, sleeping….) Maximal rates occur during peak activity (all out exercise) Vary depending on: Age, sex, size Body and environmental temperature Food quality & quantity Amount of available oxygen Hormonal balance Time of day & activity level Endotherms Animals that generate their own body heat metabolically Includes birds, mammals Many endotherms are homeothermic Maintain temperature within narrow limits Basal metabolic rate Endothermic animal’s metabolic rate measured under resting, fasting, & stress-free conditions Average BMR for humans: Males….1600-1800 kcal/day Females….1300-1500 kcal/day Ectotherms Animals that acquire most of their body heat from the environment Includes fish, amphibian, reptiles, & invertebrates Minimal metabolic rate must be determined at a specific temperature Standard metabolic rate (SMR) Ectotherm’s minimal metabolic rate measured under controlled temperature as well as resting, fasting, & stress-free conditions Metabolic rate/gram is inversely related to body size among similar animals Smaller animals consume more calories/gram than larger animals Correlated to higher metabolic rate & need for faster oxygen delivery to tissues Small animals have a higher Breathing rate Blood volume Heart rate Inverse relationship holds true for both endotherms & ectotherms Body plans & the external environment single celled organisms Multi-cellular animals body plan must have sufficient surface area of plasma membrane to service the entire volume of cytoplasm two-layered sac ... e.g. coelenterates / cnideria flat-shaped body ... max. surface area exposed to water (e.g. the flat worms) most complex animals have a small SA : volume and lack adequate exchange area on the outer surface but….. Highly folded, moist, internal surfaces for material exchange. Circulatory system shuttles materials between the inner & outer surfaces Regulating the internal environment Mechanisms interstitial fluid Regulators: Internal controls used to maintain homeostasis if external environment changes Conformers: internal conditions vary with external environmental changes **animal may be regulate according to 1 variable & conform to another Homeostasis depends on feedback circuits (i.e. nervous system) receptors - detects internal change control center - processes information from receptors & directs the effectors to respond effector provides a response feedback (common method of regulation in animals) positive feedback enhancement of a response Rare Ex. Blood clotting, childbirth contractions negative feedback reduced response examples: insulin regulation in mammal, body temp. regulation hypothalamus ... detects high blood temperature impulse to sweat gland sweat gland increases output evaporative cooling normal temperature obtained stops sending impulse Thermo-regulators Animal maintains internal temperature within a tolerable range Can be ectotherm or endotherm Poikilotherm Homeotherm Internal temps vary widely Relatively stable internal temps Modes of heat exchange between organism & the environment: Conduction Convection Radiation Evaporation Adaptations for thermo-regulation Insulation Hair, feathers, & fat layers Reduce flow of heat between organism & the environment Circulatory adaptations Vaso-dilation Vasoconstriction Increase diameter of surface blood vessels Warms skin & transfers heat to environment by radiation, convection, & conduction Decreases diameter of surface blood cells Reduces blood flow to the surface Retain heat Countercurrent heat exchange Arrangement of blood vessels to help trap heat in body core Reduces heat loss Cooling by evaporative heat loss Sweating Breathing Panting Increase mucus production Spread of saliva on body surface Behavioral responses Move between shaded & sunny locales Huddling Shivering Torpor Hibernation- long term (cold) Estivation/summer torpor- slow in summer to survive high temps & low water levels Daily torpor- based on feeding times Acclimatization Physiological response occurring over days or weeks to environmental temperature change Occurs by: Increasing or decreasing insulation (fur or fat) Changing functional enzymes Changing proportions of unsaturated fats & cholesterol in the cell membrane Rapid adjustments to temperature change involve stress-induced proteins i.e. heat shock proteins- protect other proteins from denaturization at high temperatures