Foundations of Genetics Chapter 7 Copyright © McGraw-Hill Companies Permission required for reproduction or display Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Outline • • • • • • Gregor Mendel Pea Experiments Inheritance Theory Punnett Squares Epistasis Multiple Alleles Sex Linkage Human Heredity Mutations Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Gregor Mendel • Monk studied science and mathematics at University of Vienna. Worked with garden peas to study heredity. - Many varieties available - Infrequent version of a trait - Small and easy to grow - Left alone, flowers do not open Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Mendel’s Design • • • Let each variety self-fertilize for several generations (True breeding). P generation Crossed two pea varieties exhibiting alternative traits such as white versus purple flowers. Offspring resulted in F1 generation. Allowed plants produced in crosses to self fertilize and counted numbers of each type of offspring. F2 Generation Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Copyright © McGraw-Hill Companies Permission required for reproduction or display Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Mendel’s Results • • F1 Generation All F1 flowers were purple. - Called expressed trait dominant and unexpressed trait (white flowers) recessive. F2 Generation Approximately 25% of individuals exhibited white flowers. Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Mendel’s Results • F3 Generation Ratio of purple:white flowers was 3:1. - Concluded 3:1 ratio in F2 generation was actually a 1:2:1 ratio. - (1) True-breeding dominant: (2) not-true breeding dominant: (1) true-breeding recessive. Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Mendel’s Proposed Theory • • Parents transmit information about traits to offspring (genes). Each parent contains two copies of the factor governing each trait. Homozygous - same copies Heterozygous - different copies Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Mendel’s Proposed Theory • • • Alternative forms of a factor lead to alternative traits (alleles). Phenotype - Appearance (Physical expression of genes). Genotype - Alleles governing appearance. Alleles inherited from each parent do not affect each other. Presence of allele does not ensure trait expression. Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Punnett Squares • Punnett Squares can be used to visualize crosses between two individuals. Possible gametes of individual are listed along horizontal side. And possible gametes of the other individual are listed along the vertical side. - Genotypes of potential offspring are represented by cells within square. Frequency expressed as probability. Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Punnett Squares Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Testcross • Test unknown individual with a homozygous recessive yields two alternatives: Unknown individual homozygous (PP) PP x pp: all offspring have purple flowers. Unknown individual heterozygous (Pp) Pp x pp: one-half offspring will have white flowers (pp) and one-half have purple flowers (Pp). Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Mendel’s Laws • • First Law - Segregation Only one allele specifying an alternative trait can be carried in a particular gamete, and gametes combine randomly in forming offspring. Second Law - Independent Assortment Genes located on different chromosomes are inherited independently of one another. - Dihybrid Crosses Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Epistasis • Epistasis - Interaction between products of two genes where one gene modifies the other gene’s phenotypic expression. Emerson - To produce pigment, a Zea mays plant must possess at least one functional copy of each enzyme gene. Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Epistasis Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Multiple Alleles • Often each allele has its own effect and the alleles are considered codominant. Human ABO Blood type - Type A individuals only galactosamine - Type B individuals add only galactose - Type AB individuals add both sugars - Type O individuals add neither sugar Rh Blood Group - Rh cell surface marker Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies ABO Blood Groups Copyright © McGraw-Hill Companies Permission required for reproduction or display Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Genotypic Modifications • • Pleiotrophic alleles have more than one effect on the phenotype. May have different effects on different phenotypic consequences. Characteristic of many inherited disorders. Incomplete Dominance Some alleles produce a heterozygote phenotype that is intermediate between those of the parents. Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Sex Linkage • A trait determined by a gene on the sex chromosome is said to be sex-linked. In Drosophila, sex is determined by the number of copies of the x chromosome. Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Human Chromosomes • An individual’s particular array of chromosomes is known as a karyotype. Failure of chromosomes to separate correctly during meiosis I or II is called nondisjunction, which leads to aneuploidy (abnormal number of chromosomes). - Down Syndrome caused by trisomy 21. Overall odds are 1 in 750. Much higher in women over 45. Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Nondisjunction in Sex Chromosomes • • X Chromosome XXX or XXY yields Klinefelter Syndrome XO yields Turner Syndrome Y Chromosome XYY - Antisocial ? Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Mutations in Human Heredity • Mutations are accidental changes in genes. Rare, random, and usually result in recessive alleles. - Pedigrees used to study heredity. Hemophilia - Inherited condition where blood is slow to clot or does not clot at all. Only expressed when individual has no copies of the normal allele. Royal Hempohilia - Sex-linked Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Sickle-Cell Anemia • Sickle-Cell Anemia is a recessive inherited disorder in which afflicted individuals have defective hemoglobin, and thus are unable to properly transport oxygen to tissues. - Heterozygotes usually appear normal. - Homozygotes have Sickle-Cell, but are resistant to malaria. Copyright © McGraw-Hill Companies Permission required for reproduction or display Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Other Disorders • • Tay-Sachs Incurable heredity disorder which causes brain to deteriorates. - Nonfunctional form of hexosaminidas A enzyme. Huntington’s Disease Inherited condition caused by dominant allele causing progressive brain deterioration. - Symptoms usually develop late in life. Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Genetic Counseling and Therapy • Process of identifying parents at risk of producing children with genetic defects and assessing genetic state of early embryos. Amniocentesis Ultrasound Chorionic Villi Sampling Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Review • • • • • • Gregor Mendel Pea Experiments Inheritance Theory Punnett Squares Epistasis Multiple Alleles Sex Linkage Human Heredity Mutations Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Copyright © McGraw-Hill Companies Permission required for reproduction or display Copyright © McGraw-Hill Companies Permission required for reproduction or display Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies