Unit 4 – Taxonomy • In this unit, we look at how biologists classify organisms into groups. Taxonomy • When communicating in science, it is very important to be specific. Vague terms lead to misinterpretations, confusion, and disaster. • For living things, classifying them specifically allows researchers around the world to be certain they are dealing with the exact same organism, not a variant species or alternate strain. • As well, we can create clearly-defined categories for life based on their genetic/evolutionary history and not just appearance. Linnaean Taxonomy • The modern system for naming organisms owes much to the work of 18th century naturalist Carl Linnaeus. • This system is known as binomial naming, as each species has a two-word name. Linnaean Taxonomy • Binomial names include a name of the species in particular as well as a name of their genus. A genus is a grouping for closely related species. • The order goes genus then species. • • • • Human – Homo sapiens Neanderthal - Homo neanderthalensis Panther – Panthera pardus Sometimes the genus will be abbreviated - E. coli Linnaean Taxonomy • Using this system we can easily identify closelyrelated organisms from their genus. • The system goes beyond this binomial nomenclature, there are as many as 8 levels in this system: • Domain, Kingdom, Phylum (plural: phyla), Class, Order, Family, Genus, Species, (Sub-species) • All of these terms are called taxa (sing. taxon). Linnaean Taxonomy • Human: o o o o o o o o Domain – Eukaryota Kingdom – Animalila Phylum – Chordata Class - Mammalia Order – Primates Family – Hominidae Genus – Homo Species – Sapiens • Neanderthal: o Domain – Eukaryota o Kingdom – Animalila o Phylum – Chordata o Class - Mammalia o Order – Primates o Family – Hominidae o Genus – Homo • Species – neanderthalensis Linnaean Taxonomy • Human: o o o o o o o o Domain – Eukaryota Kingdom – Animalila Phylum – Chordata Class - Mammalia Order – Primates Family – Hominidae Genus – Homo Species – Sapiens • Dog: o o o o o o o o Domain – Eukaryota Kingdom – Animalila Phylum – Chordata Class - Mammalia Order – Carnivora Family – Canidae Genus – Canis Species - Lupus Linnaean Taxonomy • Human: o o o o o o o o Domain – Eukaryota Kingdom – Animalila Phylum – Chordata Class - Mammalia Order – Primates Family – Hominidae Genus – Homo Species – Sapiens • E. coli: o o o o o o o o Domain – Bacteria Kingdom – Bacteria Phylum – Proteobacteria Class - Gammaproteobacteria Order – Enterobacteriales Family – Enterobacteriaceae Genus – Escherichia Species - Coli Models of Diversity • One of the oldest questions of biology involves how species have changed and diverged from each other throughout time. • Most traditional models show life originating from a common ancestor which over time diverged into all organisms we now know. Models of Diversity • Scientists often organize the divergence of life in a tree diagram. • However, it is important to realize that species near the top are not inherently superior, they are just simply more recent occurances. Five Kingdom Model (Old) • One model to group organisms is the Five Kingdom Model. Up until recently, this was the generally accepted classification system. Here, life was grouped at the highest level into five kingdoms. • • • • • Monera Protista Fungi Plantae Animalia Five Kingdom Model (Old) • Monera: all prokaryotic organisms. • Fungi: eukaryotes that decompose their food externally and absorb it. • Plantae: photosynthetic eukaryotes that build foodstuffs using light energy. • Animalia: eukaryotes that feed by consuming other organisms. • Protista: all single-celled eukaryotic organisms. (basically all eukaryotes that are not fungi, plants or animals) Five Kingdom Model (Old) • The five kingdom model had two main problems. • First, the kingdom monera contained all prokaryotes, despite the fact that prokaryotes are a very diverse group. • Grouping all of these diverse organisms into one group was forcing species together who are actually unrelated. Five Kingdom Model (Old) • In particular, the two major groups of prokaryotes, bacteria and archaea were discovered to have very different evolutionary histories. • Archaea have been discovered to have more in common with eukaryotes than bacteria, evolutionarily despite still being prokaryotes. Five Kingdom Model (Old) • The modern interpretation is that bacteria diverged off from other life earlier than archaea, leaving more in common between archaeans and eukaryotes. Five Kingdom Model (Old) • In the end, archaea are not only very different from their fellow prokaryotes the bacteria but are simply unique in general. • Thus, the kingdom monera is a bad label as it groups two very different groups together. Five Kingdom Model (Old) • The other main problem with kingdom system involves the kingdom Protista. • Again, this category involves combining too many diverse creatures into one group – it gives a false sense of relatedness. • Protists are mashed together despite very different metabolisms, life cycles and development. Three Domain Model • To address these issues, a new three domain model was introduced. Domains are now the highest level, with kingdoms below. Three Domain Model • In the three domain system, the large evolutionary differences between prokaryotes is addressed by replacing monera with the Archaea and Bacteria domains. Three Domain Model • The four eukaryotic kingdoms now belong under the Eukarya domain. • Protists still remain somewhat forced into a group, but they are more often grouped by their phyla now rather than by the group “protist” as a catch-all. Ring of Life Model • Recently, the idea that early life evolved linearly with species branching off and off has come in question. • Many scientists now propose that early life would have been more similar than it is now and thus sharing DNA between species was more common. • Thus rather than branching straight off, early species made be interrelated on many, many levels. Ring of Life Model • The ring of life model basically proposes that very early on life forms shared genetic information significantly more than now. This forms a interconnected ring of relationships between early life forms. • Later, species would begin to diverge away from the ring and into the separate categories we have today. Origin(s) of Life on Earth • In the end, finding a model to fit the development of life on earth may involve mixing one or more theories. The fact that early life does not as readily fossilize as large, modern species only further makes the past harder to observe. • There are also some evidence that life on Earth may not have began on Earth but instead have been seeded by small microbes arriving by meteors. • Given the problems in finding evidence of these old life forms, we may never know the specifics of the early life on this planet. Video • http://www.youtube.com/watch?v=F38BmgPcZ_I