QuickTime™ and a TIFF (Uncompressed) decompressor are needed to see this picture. Chapter 15 Darwin became a naturalist on the HMS QuickTime™ Beagle in 1831 and took aand 5 ayear journey TIFF (LZW) decompressor are needed to see this picture. around the world. Darwin made many observations and collected evidence that led him to his theory of evolution. Darwin collected many plant and animal specimens. He found many fossils of organisms once alive that resembled organisms presently living. QuickTime™ and a TIFF (Uncompressed) decompressor are needed to see this picture. Fossils from the San Francisco Bay region. Mesozoic fossils are all marine, (J) Jurassic "clams" , (K) a Cretaceous ammonite, and (L) a Cretaceous ichthyosaur. Land-animal fossils are all Cenozoic in age, such as (M) part of the Miocene beaver skull, (N) a Pleistocene saber-toothed cat, and (O) a Pleistocene mammoth. Darwin observed that the characteristics of many animals and plants varied noticeably among the different Galapagos Islands. QuickTime™ and a TIFF (Uncompressed) decompressor are needed to see this picture. QuickTime™ and a TIFF (Uncompressed) decompressor are needed to see this picture. Saddle back shell Hood Island Dome shaped shell Isabela Island QuickTime™ and a TIFF (Uncompressed) decompressor are needed to see this picture. Darwin observed the similarities and differences amongst these finches and noticed how their beaks varied and were suited for the type of food they ate. QuickTime™ and a TIFF (Uncompressed) decompressor are needed to see this picture. (1785) James Hutton - Earth is shaped by geological forces over long periods of time. (1798) Thomas Malthus - more offspring are born than survive, limited by food and space. (1809) Jean-Baptiste Lamarck - organism’s acquire or lose traits by selective use or disuse passing on the trait to the next generation. QuickTime™ and a TIFF (Uncompressed) decompressor are needed to see this picture. Lamarck’s reasoning for his law of use and disuse (1831) Charles Darwin - evidence from voyage around the world laid the groundwork for his theory of evolution. (1833) Charles Lyell - processes occurring now shaped Earth’s geological features over long periods of time. (1858) Alfred Wallace - he also speculated that evolution by natural selection occurs. This spurred Darwin on to publish his theory. Origin of Species Darwin proposed that new species originate from ancestral forms through the gradual accumulation of adaptations over very long periods of time. In his book, Darwin describes descent with modification. Some heritable variations in populations are better suited QuickTime™ and a TIFF (Uncompressed) decompressor than others and since organisms tend to produce more are needed to see this picture. offspring than the environment can support, competition for resources favors those better suited. These individuals reproduce and pass on their traits. Darwin observed artificial selection in which nature provided variation and humans selected variations they found useful. QuickTime™ and a TIFF (LZW) decompressor are needed to see this picture. QuickTime™ and a TIFF (LZW) decompressor are needed to see this picture. QuickTime™ and a TIFF (LZW) decompressor are needed to see this picture. Variation leading to Natural Selection Insect Resistance Artificial Selection QuickTime™ and a TIFF (LZW) decompressor are needed to see this picture. Darwin proposed this idea. Descent with modification implies a common ancestor with a single tree of life linking all living things. In humans: use of drugs selects for pathogens that through chance mutations are resistant to the drugs’ effects. Bacteria and viruses evolve rapidly due to rapid rates of mutation and reproductive, posing a challenge to our society. HIV Resistance to 3TC QuickTime™ and a TIFF (LZW) decompressor are needed to see this picture. 1. 2. 3. 4. Fossil Record Geographical distribution of living species Homologous structures Similarities in embryological development Fossils provide the strongest evidence of evolution. Sedimentary strata reveal the relative ages of fossils. QuickTime™ and a TIFF (LZW) decompressor are needed to see this picture. Absolute ages are determined by radiometric dating. During his voyage, Darwin observed animals in similar environments on different continents that had similar anatomy and behavior. All species of finch on the Galapagos Islands evolved from a common ancestor from Ecuador. Differing selective pressures on the each Island was the driving force for the evolution of beaks. QuickTime™ and a TIFF (Uncompressed) decompressor are needed to see this picture. QuickTime™ and a TIFF (Uncompressed) decompressor are needed to see this picture. QuickTime™ and a TIFF (Uncompressed) decompressor are needed to see this picture. During the past few decades scientists have demonstrated that what is now called South America was part of a large land mass called Gondwana, which included Australia and Antarctica. QuickTime™ and a TIFF (Uncompressed) decompressor are needed to see this picture. Horses have a complex evolutionary history. The earliest horses evolved in North America; many lineages arose and died out, and ancestors of several of these lineages crossed into Asia over the Bering land bridge and into South America over the Central America land bridge. Organisms living in different geographical locations under similar ecological conditions are exposed to similar pressures of natural selection. Evolution of shared traits in unrelated species is convergent evolution. Regardless of the appearance of organisms, two species are closely related when they share common ancestry. Homologous structures have different functions but evolved from common ancestries. Forelimbs of all mammals show the same arrangement of bones with different functions. Analogous structures have a similar function but are anatomically different. Humans also have vestigial features, evidence of our own evolutionary history. The appendix is believed to be a remnant of a larger, plantdigesting structure found in our ancestors QuickTime™ and a TIFF (Uncompressed) decompressor are needed to see this picture. QuickTime™ and a TIFF (Uncompressed) decompressor are needed to see this picture. Cave-dwelling tetra fish are blind; they have small vestigial eyes that do not work. Why have them? Biologists have found a possible answer: genetic mutations that hamper eye development also may increase the number of taste buds. Thus, mutations that happened to give the fish an advantage in tasting and smelling—a huge benefit in a dark environment— might also have inadvertently, and harmlessly, caused the degeneration of their eyes. . QuickTime™ and a TIFF (Uncompressed) decompressor are needed to see this picture. Pythons and boa constrictors have tiny hind leg bones buried in muscles toward their tail ends. These are vestigial. Vestigial legs are a clue that snakes descended from lizards. Over 100 million years ago, some lizards happened to be born with smaller legs, which, in certain environments, helped them move about unencumbered. As generation after generation survived and reproduced, this new form flourished. QuickTime™ and a TIFF (Uncompressed) decompressor are needed to see this picture. All vertebrate embryos have a tail and pharyngeal pouches. This indicates a common ancestor. Genes are not active at the same time. Those that are active during early development are less subject to change than genes that are active later. This is because mutations occurring early in development have a far greater chance of being lethal and would not be passed on to future generations. Mutations occurring later in development have more limited effects, are less likely to be lethal and are more likely to be passed on. That’s why humans and chickens are so different at later stages despite similarities as embryos. Comparing DNA sequences is the most direct way to determine evolutionary relationships. The sequence of nitrogenous bases is more similar in closely related species than in species that are not as closely related. Scientists can use this information, with knowledge of mutation rates, to estimate how long it’s been since 2 species shared a common ancestor. 1. 2. 3. 4. 5. 6. Organisms differ and these differences may be inheritable. Organisms produce more offspring than survive. Organisms compete for limited resources. Organisms best suited to their environment survive and reproduce, passing on these traits. This process of natural selection causes species to change over time. Species alive today descended with modification from a common ancestor uniting all organisms in a tree of life.