Nonvertebrate Chordates, Fishes, and Amphibians What Is a Chordate? Members of the phylum Chordata are called chordates. A chordate is an animal that has, for at least some stage of its life, a dorsal, hollow nerve cord; a notochord; pharyngeal pouches; and a tail that extends beyond the anus. Characteristics of Chordates Muscle segments Tail Anus Notochord Hollow nerve cord Mouth Pharyngeal pouches Characteristics of Chordates The notochord is a long supporting rod that runs through the body just below the nerve cord. Pharyngeal pouches are paired structures in the throat (pharynx) region. The tail can contain bone and muscle and is used for swimming by many aquatic species. What Is a Chordate? Most Chordates Are Vertebrates About 96 percent of all chordate species are vertebrates. Most vertebrates have a vertebral column, or backbone. In vertebrates, the dorsal, hollow nerve cord is called the spinal cord. What Is a Chordate? As a vertebrate embryo develops, the front end of the spinal cord grows into a brain. The backbone is made of individual segments called vertebrae. In addition to support, vertebrae enclose and protect the spinal cord. Phylogeny of Chordates Sharks & their Jawless relatives fishes Nonvertebrate chordates Bony fishes Invertebrate ancestor Reptiles Birds Amphibians Mammals Nonvertebrate Chordates The two groups of nonvertebrate chordates are tunicates and lancelets. Similarities in anatomy and embryological development indicate that vertebrates and nonvertebrate chordates evolved from a common ancestor. Both tunicates and lancelets are soft-bodied marine organisms. Tunicates (Larva) The larval form of filterfeeding tunicates have all of the chordate characteristics. Tail Hollow nerve cord Wasteeliminating organ Notochord Mouth Intestine Pharynx with gill slits Stomach Heart Larva Tunicates (Adult) Siphon to mouth Pharynx with gill slits Siphon from anus Anus Tunic Intestine Reproductive organs Heart Stomach Adult Lancelets Lancelets are small, fishlike creatures that live on the ocean bottom. Notochord Hollow nerve cord Segmented muscles Mouth Tail Anus Pharynx with gill slits Intestine Reproductive organs Lancelets An adult lancelet has a definite head region that contains a mouth. As water passes through the pharynx, a sticky mucus catches food particles. The lancelet swallows the mucus into the digestive tract. Lancelets Lancelets use the pharynx for gas exchange. Lancelets are thin enough to exchange gases through their body surface. Lancelets have a closed circulatory system and do not have a true heart.