Chapter 12 Lecture Outline See separate PowerPoint slides for all figures and tables preinserted into PowerPoint without notes. Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. 1 Chapter 12 Blood 2 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. 12.1 Introduction 3 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. A. Blood, a type of connective tissue, is a complex mixture of cells, chemicals, and fluid. B. Blood transports substances throughout the body, helps to maintain a stable interstitial fluid environment, and distribute heat. C. The blood includes red blood cells, white blood cells, platelets, and plasma. 4 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. D. Blood Volume and Composition 1. Plasma is a mixture of water, amino acids, proteins, carbohydrates, lipids, vitamins, hormones, electrolytes, and cellular wastes. 2. A blood hematocrit (HCT) is normally 45% cells (mainly red blood cells) and 55% plasma. 3. An average-sized adult has a blood volume of about 5.3 quarts (5 liters). 5 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. Fig 12.1 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. 6 Fig 12.2 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. 7 12.2 Blood Cells 8 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. A. Red Blood Cells 1. Description a. Red blood cells (erythrocytes) are biconcave disks that contain one-third oxygen-carrying hemoglobin by volume. b. When oxygen combines with hemoglobin, bright red oxyhemoglobin results. c. Deoxygenated blood (deoxyhemoglobin) is darker. d. Red blood cells discard their nuclei during development and so cannot reproduce or produce proteins. 9 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. Red Blood Cells, Description, cont. e. Red blood cells produce ATP through glycolysis alone and can not use the oxygen they carry since they have no mitochondria. 10 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. Fig 12.3 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. 11 2. Red blood cells counts a. The typical red blood cell count is 4,700,000 - 6,100,000 cells per mm3 for males and 4,200,000 - 5,400,000 cells per mm3 for females. b. The number of red blood cells is a measure of the blood's oxygen-carrying capacity. 12 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. 3. Red blood cell production and its control a. In the embryo and fetus, red blood cell production (erythropoiesis) occurs in the yolk sac, liver, and spleen; after birth, it occurs in the red bone marrow. b. The average life span of a red blood cell is 120 days. c. The stages of red blood cell formation from hematopoietic stem cells (hemocytoblasts) is displayed in Figure 12.4 on the next slide. 13 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. Fig 12.4a Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. 14 Red blood cell production and its control, cont. d. The total number of red blood cells remains relatively constant due to a negative feedback mechanism utilizing the hormone erythropoietin, which is released from the kidneys and liver in response to the detection of low oxygen levels. e. Excessive increase in red blood cells is called polycythemia which causes viscous, slow moving blood. 15 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. Fig 12.5 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. 16 4. Dietary factors affecting red blood cell production a. Vitamins B12 and folic acid are needed for DNA synthesis, so they are necessary for the reproduction of all body cells, especially in hematopoietic tissue. b. Iron is needed for hemoglobin synthesis; most of the iron comes from recycling old red blood cells. c. A deficiency in red blood cells or quantity of hemoglobin results in anemia. 17 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. 5. Destruction of red blood cells a. With age, red blood cells become increasingly fragile and are damaged by passing through narrow capillaries. b. Macrophages in the liver and spleen phagocytize damaged red blood cells. c. Hemoglobin from the decomposed red blood cells is converted into heme and globin. d. Heme is decomposed into iron which is stored or converted into biliverdin and then bilirubin which are excreted in bile. e. The globin is broken down into amino acids and reused. 18 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. Fig 12.6 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. 19 B. White blood cells 1. Description a. White blood cells (leukocytes) help defend the body against disease. b. They are formed from hemocytoblasts (hematopoietic stem cells) in red bone marrow. c. Hormones that stimulate WBC production fall into two categories, interleukins and colony-stimulating factors (CSF’s). d. White blood cells can leave the bloodstream to fight infection. e. Life span is about 12 hours. 20 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. 2. Five types of white blood cells are in circulating blood and are distinguished by size, granular appearance of the cytoplasm, shape of the nucleus, and staining characteristics. a. The types of white blood cells are the granulocytes, (neutrophils, eosinophils, and basophils), and the agranulocytes (monocytes and lymphocytes). 21 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. b. Neutrophils have purple-stained fine cytoplasmic granules and a multi-lobed nucleus; they comprise 54-62% of leukocytes. Fig 12.7 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. 22 c. Eosinophils have coarse granules that stain deep red, a bilobed nucleus, and make up only 1-3% of circulating leukocytes. Fig 12.8 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. 23 d. Basophils have fewer granules that stain blue; they account for fewer than 1% of leukocytes. Fig 12.9 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. 24 e. Monocytes are the largest blood cells, have variably-shaped nuclei, and make up 3-9% of circulating leukocytes. Fig 12.10 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. 25 f. Lymphocytes are long-lived, have a large, round nucleus, and account for 25-33% of circulating leukocytes. Fig 12.11 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. 26 3. Functions of white blood cells a. Leukocytes can squeeze between cells lining walls of blood vessels by diapedesis and attack bacteria and debris. b. Neutrophils and monocytes are phagocytic, with monocytes (become macrophages) engulfing the larger particles. Both of these contain many lysosomes that help breakdown organic molecules. c. Eosinophils moderate inflammation and allergic reactions, as well as defend against parasitic infections. 27 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. Fig 12.12 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. 28 Functions of white blood cells, cont. d. Basophils migrate to damaged tissues and release histamine to promote inflammation and heparin to inhibit blood clotting. e. Lymphocytes are the major players in specific immune reactions; B cells produce antibodies. 29 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. 4. White blood cell counts a. Normally a cubic millimeter of blood contains 3,500 to 10,500 white blood cells. b. Leukocytosis occurs after an infection when excess numbers of leukocytes are present. c. Leukopenia, too few white blood cells, occurs from a variety of conditions, including AIDS. 30 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. White blood cell counts, cont. d. A differential white blood cell count (DIFF) can help pinpoint the nature of an illness, indicating whether it is caused by bacteria or viruses. A differential white blood cell count lists the percentages of the types of leukocytes in a blood sample. 31 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. C. Blood platelets 1. Blood platelets or thrombocytes are fragments of megakaryocytes, developed from hematopoietic stem cells in response to thrombopoietin. 2. Platelets help repair damaged blood vessels by adhering to their broken edges. 3. Normal platelet counts vary from 150,000 to 350,000 platelets per mm3. 32 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. Table 12.1 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. 33 12.3 Blood Plasma 34 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. A. Plasma is the clear, straw-colored fluid portion of the blood. 1. Plasma is mostly water (92%) but contains a variety of substances. 2. Plasma functions to transport nutrients and gases, regulate fluid and electrolyte balance, and maintain a favorable pH. 35 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. B. Plasma Proteins 1. The plasma proteins are the most abundant dissolved substances in the plasma. 2. Plasma proteins are not used for energy and fall into three groups-albumins, globulins, and fibrinogen. a. The albumins help maintain the colloid osmotic pressure of the blood and account for 60% of the plasma proteins. 36 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. Plasma Proteins, cont. b. The globulins, comprising 36% of the plasma proteins, are designated as alpha, beta, and gamma globulins. 1) Alpha and beta globulins function in transporting lipids and fat-soluble vitamins. 2) Gamma globulins are a type of antibody. c. Fibrinogen (4%) plays a primary role in blood coagulation. 37 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. Table 12.2 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. 38 C. Gases and Nutrients 1. The most important blood gases are oxygen and carbon dioxide. 2. The plasma nutrients include amino acids, monosaccharides, nucleotides, and lipids from the digestive tract. 3. Since lipids are not soluble in the water of the plasma, they are surrounded by protein molecules for transport through the bloodstream as lipoproteins. 39 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. D. Nonprotein Nitrogenous Substances 1. Nonprotein nitrogenous substances generally include amino acids, urea, and uric acid, creatine and creatinine. 2. Urea and uric acid are the by-products of protein and nucleic acid catabolism. 3. Amino acids come from the breakdown of proteins. 4. Creatinine comes from creatine (creatine phosphate in muscles) 40 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. E. Plasma Electrolytes 1. Plasma electrolytes are absorbed by the intestine or are by-products of cellular metabolism. 2. They include sodium, potassium, calcium, magnesium, chloride, bicarbonate, phosphate, and sulfate ions. 3. Some of these ions are important in maintaining osmotic pressure and pH of the plasma. 41 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. Fig 12.2 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. 42 12.4 Hemostasis 43 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. A. Hemostasis refers to the stoppage of bleeding. 1. Following injury to a vessel, three steps occur in hemostasis: blood vessel spasm, platelet plug formation, and blood coagulation. 44 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. B. Blood Vessel Spasm 1. Cutting a blood vessel causes the muscle in its walls to contract in a reflex, or engage in vasospasm. 2. This reflex lasts only a few minutes, but it lasts long enough (30 minutes) to initiate the second and third steps of hemostasis. 3. Platelets release serotonin at this time constricting smooth muscles in the blood vessel walls. 45 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. C. Platelet Plug Formation 1. Platelets stick to the exposed edges of damaged blood vessels, forming a net with spiny processes protruding from their membranes. 2. A platelet plug is most effective on a small vessel. 46 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. Fig 12.13 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. 47 D. Blood Coagulation 1. Blood coagulation is the most effective means of hemostasis. 2. Blood coagulation is very complex and uses many clotting factors as well as calcium. 3. Damaged tissues release a chemical called tissue thromboplastin, which activates the first in a series of factors leading to the production of prothrombin activator. 4. Prothrombin activator converts prothrombin in the plasma into thrombin. This in turn, catalyzes a reaction that converts fibrinogen into fibrin. 48 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. Fig 12.14 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. 49 Blood Coagulation, cont. 5. The major event in blood clot formation is the conversion of soluble fibrinogen into net like insoluble fibrin causing the blood cells to catch. 6. Serum is plasma minus the clotting factors. 7. The amount of prothrombin activator formed is proportional to the amount of tissue damage. 8. Once a blood clot forms, it promotes still more clotting through a positive feedback system. 50 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. Blood Coagulation, cont. 9. After a clot forms, plasminogen is converted to plasmin that digests the fibrin. 10. Fibroblasts then enter the area and begin repair by forming fibrous connective tissue to seal the break. 11. A clot that forms abnormally in a vessel is a thrombus; if it dislodges, it is an embolus. 51 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. Fig 12.16 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. 52 12.5 Blood Groups and Transfusions 53 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. A. After mixed success with transfusions, scientists determined that blood was of different types and only certain combinations were compatible. 1. Before a transfusion is given, the blood must be typed and cross-matched. 54 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. B. Antigens and antibodies 1. Clumping of red blood cells following transfusion is called agglutination. 2. Agglutination is due to the interaction of proteins on the surfaces of red blood cells (antigens) with certain antibodies carried in the plasma. 3. Only a few of the antigens on red blood cells produce transfusion reactions; these include the ABO group and Rh group. 55 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. C. ABO Blood Group 1. Type A blood has A antigens on red blood cells and anti-B antibodies in the plasma. 2. Type B blood has B antigens on red blood cells and anti-A antibodies in the plasma. 3. Type AB blood has both A and B antigens, but no antibodies in the plasma; universal recipient. 4. Type O blood has neither antigen, but both types of antibodies in the plasma; universal donor. 56 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. Fig 12.17 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. 57 ABO Blood Group, cont. 5. Adverse transfusion reactions (agglutination of red blood cells) are avoided by preventing the mixing of blood that contains matching antigens and antibodies. 58 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. Table 12.3 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. 59 Fig 12.18a&b Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. 60 Fig 12.18c&d Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. 61 D. Rh Blood Group 1. The Rh blood group was named after the rhesus monkey. 2. If the Rh factor surface protein is present on red blood cells, the blood is Rh positive; if it is absent, the blood is Rh negative. 3. There are no corresponding antibodies in the plasma unless a person with Rh-negative blood has physical contact with Rh-positive blood; the person will then develop antibodies for the Rh factor. 62 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. Rh Blood Group, cont. 4. There are two ways in which an Rh-negative individual can have contact with Rh-positive blood – a transfusion or pregnancy. 5. If an Rh-negative woman carries an Rh-positive baby, she may be exposed to the positive blood during delivery. a. The mother will now make antibodies against Rh that could attack the blood of a second Rh-positive baby – called erythroblastosis fetalis. b. The problem can be prevented by giving the mother the drug, RhoGAM. 63 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. Fig 12.19 Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. 64 The End Copyright © 2015 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education.