Kimberley Nguyen 35: Plant Structure, Growth, & Development Overview: No Two Plants are Alike Plasticity is when plant form is controlled by environmental and genetic factors 35.1: The plant body has a hierarchy of organs, tissues, & cells Plants have organs, which are made up of tissues, which are made up of cells Roots: - help anchor a plant in soil, absorb minerals and water, store organic nutrients - eudicots and gymnosperms have a taproot system and lateral roots branching off from it - monocots have small roots, called a fibrous root system - modified roots: prop roots, storage roots, “strangling” aerial roots, buttress roots, pneumatophores Stems: - a system of nodes (where leaves are attached) and internodes (segment between nodes) - between a leaf and stem, is an axillary bud, which are unlikely form into a branch, while terminal buds are at the tip of a branch and will develop since it gets all the nutrients this unbalanced spread of resources is called apical dominance - modified stems: stolons, bulbs, tubers, rhizomes Leaves: - main photosynthetic organ, consisting of a flat blade and petiole - angiosperms classified by leaf shape, spatial arrangement of leaves, and vein patterns - modified leaves: tendrils, spines, storage leaves, bracts, reproductive leaves Dermal Tissue: outer protective covering and prevents water loss; epidermis and periderm Vascular Tissue System: transports materials in xylem and phloem Ground Tissue System: a filler tissue and for storage, photosynthesis and support; pith and cortex Common plant cell types: parenchyma, collenchyma, sclerenchyma, water-conducting cells in xylem, and sugar-conducting cells in phloem 35.2 Meristems generate cells for new organs Animals undergo determinate growth, while plants undergo inderterminate growth - Due to plant’s meristems which divide frequently to replace or provide additional cells - Apical meristems: add cells to shoots and roots; process called primary growth - Lateral meristams: vascular cambium which adds layers to vascular tissue and cork cambium which replace epidermis and periderm cells; process called secondary growth Plants categorized on how long they live; annuals, biennials, and perennials 35.3 Primary growth lengthens roots & shoots primary growth: apical meristems produce the primary plant body roots: - root tip has a root cap to protect the apical meristem - Root grows behind in the tip in 3 zones: cell division, elongation, maturation - In young roots, the primary growth produces the epidermis, ground, and vascular tissues Kimberley Nguyen - Structure (exterior to centre): epidermis, cortex, endodermis, pericycle (where the lateral root forms), vascular cylinder with xylem and phloem in centre (pith) In shoots: - divides at the tip of a terminal bud and is dome-shaped and creates internodes and leaves called leaf primordia - Stem structure (exterior to centre): epidermis, collenchyma cells, vascular tissue, ground tissue - Leaf structure (top to bottom): upper epidermis with stomata, ground tissue with veins, lower epidermis 35.4 Secondary growth adds girth to stems & roots in woody plants secondary growth: lateral meristems produce the secondary plant body in woody plants Primary and secondary growth occur simultaneously The vascular cambium is responsible for the thickening of a root or stem - Made of fusiform initials and ray initials - Over time, secondary xylem (wood) is mainly tracheids and fibre - Each year, a boundary is created between early and late wood Trees have layers called heartwood and sapwood Epidermis replaced by periderm when it dies(2 cork cambium tissues: phelloderm and cork cells) Bark is made up of the secondary phloem, most recent periderm and older layers of periderm 35.5: Growth, morphogenesis, & differentiation produce the plant body There are 3 developmental processes: growth, morphogenesis and cellular differentiation. Growth: - the plane and symmetry of cell division determines plant form some undergo asymmetrical cell division the microtubule ring, preprophase band determines the plane of division - water in vacuoles make cells expand rapidly and up due to cellulose microfibrils - some cells expand in all directions since the microtubule arrangement is random Morphogenesis: - process of cells being organized into multicellular arrangements - Pattern formation is determined by positional information Polarity as positional information Cellular Differentiation: - cells contain same genome, but are different in function and structure due to gene expression - Depends on positional information Phase changes:- juvenile phase to adult vegetative phase to adult reproductive phase Most significant change is the leaves of juvenile to adult Most significant change is flowering of vegetative to reproductive due to floral meristem identity genes ABC model of flower formation depicts flower’s phenotype by showing which genes are active