THE ORIGIN AND EVOLUTIONARY HISTORY OF LIFE CONDITIONS OF EARLY EARTH • Age of Earth is ~4.6 billion years • Atmosphere had little free O 2 • Included CO2, H2O, CO, H2, N2 • Maybe also NH3, H2S, CH4 REQUIREMENTS FOR CHEMICAL EVOLUTION TO PRODUCE LIFE • Absence of oxygen • Energy • Chemical building blocks • Sufficient time PREBIOTIC SOUP HYPOTHESIS • Molecules formed near the Earth’s surface • Sugars, nucleotides, amino acids formed spontaneously IRON-SULFUR WORLD HYPOTHESIS • Organic molecules formed at hydrothermal vents on the ocean floor • Laboratory simulations show catalyst effects of iron and nickel sulfides ORIGIN OF CELLS • Spontaneous assembly of small organic molecules into macromolecules RNA WORLD • Self-replicating RNA molecules • Function as both enzyme and substrate for replication • Ribozyme is enzymatic RNA • First step in evolution of the DNA / RNA / protein system DIRECTED EVOLUTION • Large pool of RNA molecules with different sequences • Selected for ability to catalyze a reaction • Amplify / mutate / repeat DIRECTED EVOLUTION • In the RNA world, ribozymes catalyzed protein synthesis • DNA formed from double strands of RNA • DNA more stable than RNA THE FIRST CELLS • Heterotrophs that feed on organic molecules • Anaerobic fermentation process to obtain energy AUTOTROPHS • Selected after organic molecule food stock became scarce • Photosynthetic production of organic molecules • Cyanobacteria split water molecules and released oxygen FORMATION OF THE OZONE LAYER • Ultraviolet radiation forms O 3 from O2 in the upper atmosphere • Prevents UV from reaching Earth • Decreased mutagenesis • Enabled organisms to live in surface waters and on land OZONE FORMATION EUKARYOTES AROSE FROM PROKARYOTES • Endosymbiont theory • Mitochondria and chloroplasts derived from prokaryotes • Ingested but not digested • Reproduced along with host cell ENDOSYMBIONT THEORY GEOLOGICAL ERAS • Paleozoic • 543 mya - 251 mya • Mesozoic • 251 mya - 65 mya • Cenozoic • 65 mya - present PRECAMBRIAN TIME • • • • • Before 543 mya Bacteria Protists Fungi Simple multicellular animals CAMBRIAN EXPLOSION • All animal phyla established • Many new body plans • Bizarre, extinct phyla ORDOVICIAN PERIOD • Shallow seas covered land • Cephalopods • Coral reefs • Jawless fishes SILURIAN PERIOD • Jawed fishes • Terrestrial plants • Air-breathing animals DEVONIAN PERIOD • Bony fishes • Amphibians • Wingless insects • All major plant groups except for flowering plants established CARBONIFEROUS PERIOD • Swamp forests • Reptiles • Winged insects PERMIAN PERIOD • Therapsids • Reptilian ancestors of mammals • Seed plants dominant • Ended the Paleozoic with the greatest mass extinction • 90% of marine species • 70% of land vertebrates MESOZOIC ERA Divided into three time periods * Triassic (251-199.6 million years ago), * Jurassic (199.6145.5 million years ago), * Cretaceous (145.565.5 million years ago). TRIASSIC PERIOD • Thecodonts • Ancestors of dinosaurs and birds • Pleiosaurs and ichthyosaurs • Pterodonts • First mammals • Small insectivores JURASSIC AND CRETACEOUS PERIODS • Saurischians • Ancestors of lizards • Ornithischians • Ancestors of birds • Ended with mass extinction caused by a meteorite impact CENOZOIC ERA • Began about 65 million years ago • continues into the present • third documented era in the history of Earth. • Tertiary period • Diversification of flowering plants, birds, insects, mammals • Quaternary period • Genus Homo • Large mammals