Plant Nutrition and Soils Chapter 29 Plant Nutrition • Plant Nutrition- uptake from the environment of all raw materials required for essential biochemical processes. • More than 60 elements have been identified in plants. • 1880s- ten chemical elements were designated as essential for plant growth. – Carbon, hydrogen, oxygen, potassium, calcium, magnesium, nitrogen, phosphorus, sulfur, iron. • 1900s- manganese, zinc, copper, chlorine, boron, molybdenum, and finally nickel. 17 Essential Elements Essential Elements Three criteria are used to judge essentiality 1) If it is needed for the plant to complete its life cycle. 2) If it is part of any molecule or constituent of the plant that is itself essential. a) Magnesium in chlorophyll molecule. b) Nitrogen in proteins. 3) If deficiency symptoms appear in the absence of the element. 2 Types of Essential Elements • Macronutrients- required in large amounts. – 1000 mg/kg or > of dry matter. • Micronutrients (trace elements)- required in very small amounts. – 100 mg/kg of dry matter. Certain plants contain unusually high and low amounts of specific elements. Silicon Macronutrient Functions • • • • Sulfur- amino acids and coenzyme A. Phosphorus- ATP, nucleic acids, phospholipids. Magnesium- chlorophyll molecule. Calcium- cell walls, cofactors, cellular membrane. • Potassium- osmosis and ionic balance, stomatal activity. • Nitrogen- amino acids, proteins, nucleotides, nucleic acids, chlorophylls. • Oxygen, Carbon, and Hydrogen- almost all chemical molecules, 96% of the plant. Micronutrient Functions • • • • • Molybdenum- nitrogen fixation and nitrate reduction. Nickel- enzyme functioning in nitrogen metabolism. Copper- involved in some REDOX reactions. Zinc- activator or component of many enzymes. Manganese- enzyme activator, membrane integrity, oxygen release in PSN. • Boron- Ca2+ utilization, nucleic acid synthesis, membrane integrity. • Iron- chlorophyll synthesis. • Chlorine- osmosis and ionic balance. Global Nitrogen and Phosphorus Use Nitrogen Losses ~ 1500 lbs/day or $500/day! Asynchrony Between N Supply and Demand Source: G. Philip Robertson IN Ecology in Agriculture, L.E. Jackson, ed. NUISANCE ALGAE BLOOMS Eutrophication – Lake Tahoe story Data courtesy of C.R. goldman and J.E. Reuter, Tahoe Reesrach Group, U. of California-Davis, http://www.news.ucdavis.edu/tahoetv/ Soil • Soil- primary nutrient (inorganic ions), water, suitable gaseous environment, and support medium for plants. – Provides a chemical and physical environment for plant growth. – Minerals- naturally occuring inorganic compounds that are usually composed of two or more elements. i.e.- Quartz (SiO2), Calcite (CaCO3). Soil Layers (Horizons) • Soils consists of at least 3 horizons (layers) A Horizon= topsoil. B Horizon= subsoil. C Horizon= soil base. Bedrock= below the horizon. Soils Are Composed of Solid Matter and Pore Space • Solid Matter- fragments of rock and minerals in the soil. Coarse Sand 200-2000 µm Fine Sand 20-200 µm Silt 2-20 µm Clay < 2 µm • Pore Space- the space around the soil particles. – Air – Water Plant Pathology- bacteria Plant Pathology- viruses Plant Pathology- fungi