Introduction to the Amphibian Body Characteristics of Amphibians • Amphibians consists of frogs, salamanders, newts, and caecilians • Amphibians share five key characteristics – – – – – 1. Legs 2. Lungs 3. Double-loop Circulation 4. A partially divided heart 5. Cutaneous respiration • These characteristics allow amphibians to thrive on land Characteristics of Amphibians • Amphibian eggs are not watertight, and therefore amphibians need to reproduce in a wet area • Because amphibians need a wet area to reproduce many amphibians must live in a moist environment Movement and Response • Many challenges that amphibians face involve having to move on land • Not all of amphibian’s sense organs work as well in air as in water – Example: larval amphibians have a lateral line that they use to detect movement of the water, but they usually lose their lateral line by adulthood Amphibian Skeleton • Terrestrial vertebrates must rely on the support of their strong internal skeleton • Strong limbs of the skeleton help support the body’s weight and allows movement • Frog skeletons have several specializations for jumping and landing – The bones of the lower limbs are fused into a single, thick bone – The hips also have thick bones with a sturdy structure that absorbs the impact of landing Sense Organs • The sense of sight and hearing are well developed in most amphibians • The primary sense organs of amphibians are the eyes and ears Amphibian Eyes • The eyes of amphibians are covered by transparent, movable membranes called a nictitating membrane Amphibian Ears • The inner ear detects sound • Sounds are transmitted to the inner ear by the tympanic membrane, or eardrum, and a small bone that extends between the tympanic membrane and the inner ear • Sound first strikes the tympanic membrane – The tympanic membrane is located on the side of the head – Vibrations cause small movements that are transmitted to the fluid-filled inner ear – In the inner ear, sensitive hair cells change sound vibrations to nervous impulses – These impulses are then transmitted to the brain Respiration • One of the biggest challenges faced by amphibians when they leave the water is gathering oxygen from the air • In amphibians the following structures work together to ensure that sufficient oxygen reaches the body tissues: • • • • Skin Lungs Double-loop circulation Partially divided heart Amphibian Lungs • Adult amphibians breathe with lungs – A lung is an internal, baglike organ that allows oxygen and carbon dioxide to be exchanged between the air and bloodstream • The amount of oxygen a lung can absorb depends on the internal surface – The greater the surface area is, the greater amount of oxygen that can be absorbed How amphibians breathe • Amphibians breathe by changing the volume and pressure of air in its mouth while either opening or closing its nostrils • With each breath oxygen is drawn into the lungs • In the lungs oxygen mixes with a small amount of air that has already given up most of its oxygen – Due to the mixing the efficiency of lungs is much less than that of gills – This is okay though because air contains 20x as much oxygen as sea water Amphibian Skin • Amphibians can obtain oxygen through their thin, moist skin = cutaneous respiration • In cutaneous respiration, gases and water pass directly through the skin – Skin must be moist for cutaneous respiration to work – To keep skin moist amphibians have mucous glands Amphibian Circulation • The tissues of land animals require greater amounts of oxygen • The structure of the amphibian circulatory system, which includes a partially divided heart and double-loop circulation, allows oxygen to be delivered to the body in an efficient manner Amphibian Partially Divided Heart • The top chambers of the amphibian heart are divided into left and right sides by the wall called the septum • The heart’s bottom chamber is not divided – This allows a mixture of oxygen-rich and oxygenpoor blood to be delivered to the amphibian’s tissues Blood Flow in the Amphibian Heart • 1. Oxygen-poor blood from the body enters the right atrium • 2. The pulmonary veins carry oxygen-rich blood from the lungs to the left atrium • 3. A mixture of oxygen-rich and oxygen-poor blood enters the ventricle • 4. The ventricle pumps blood to the lungs and body tissues The Spiral Valve • Some amphibians have a spiral valve that divides the conus arteriosus • The spiral valve helps keep the two streams of blood separate as they leave the heart Double-Loop Circulation • Amphibians evolved to have a second circulatory loop • The evolution of the second circulatory loop allows more oxygen to be delivered to their bodies • In the circulatory loop that connects an amphibian’s heart to its lungs there are blood vessels called pulmonary veins – Pulmonary veins are the vessels that carry the oxygen-rich blood from the amphibian’s lungs directly back to its heart Double-Loop Circulation • The amphibian’s second circulatory loop carries the oxygen-rich blood from the heart to the body • The advantage: Amphibians can pump oxygen-rich blood to the amphibian’s tissues at a much higher pressure and speed than that of fishes Review Questions: • Describe the key characteristics shared by all amphibians • Identify the function of the tympanic membrane • Summarize how amphibians take in oxygen • Contrast the single-loop circulation o fishes with the double-loop circulation of amphibians