Animals eat food for both the energy and the organic molecules used to assemble new molecules. An animals diet must supply chemical energy, organic molecules, and essential nutrients. Cells, tissues, organs and whole animals depend on chemical energy in the diet. This energy must be converted in ATP to power life processes Animals ingest and digest nutrients, such as carbohydrates, proteins, and lipids for use in cellular respiration and energy storage. Animal’s diet must allow provide raw materials needed for biosynthesis of complex molecules There are four classes of essential nutrients: Essential amino acids; about half of the twenty amino acids in proteins cannot be synthesized and must be obtained from food. Essential fatty acids that animals cannot make, example linoleic acid Vitamins, such as B and E vitamins Minerals, such as calcium and phosphorus Ingestion is the act of taking in food Suspension feeders sift small food particles from water Substrate feeders are animals that live in or on their food source. Fluid feeders suck nutrient rich fluids from a living host. Bulk feeders eat relatively large pieces of food and include most animals. Digestion is the breakdown of food into small molecules capable of being absorbed by the cells of the body. Enzymatic hydrolysis, the breaking of bonds with the addition of water, is the reaction type by which macromolecules are digested. Mechanical digestion precedes chemical digestion. Breaks food into smaller pieces, increasing the surface area for chemical processes. Chewing is an example of mechanical digestion. Chemical digestion Polysaccharides and dissacharides are split into simple sugars. Proteins are broken down into amino acids. Nucleic acids are broken down into nucleotides Fats and phospholipids are broken down into fatty acids and other components. In the third stage of food processing, absorption, the animals cells take up (absorb) small molecules, such as simple sugars and amino acids. Elimination completes the process as undigested material passes out of the digestive system. Digestive compartments prevent animals from digesting their own cells and tissues Food is processed with specialized compartments. Compartments may be intracellular, such as food vacuoles. May be extracellular, as in digestive organs or systems. Intracellular Digestion Cell engulfs solid food by phagocytosis of liquid food by pinocytosis Newly formed food vacuoles fuse with lysosomes. Fusion of food vacuoles and lysosomes brings food together with hydrolytic enzymes, allowing digestion to occur safely with a compartment enclosed by a protective membrane. Sponges digest food entirely by intracellular digestion Extracellular digestion is the breakdown of food in compartments that are continuous with the outside of the animal’s body. Enables an animal to devour much larger sources of food that can be ingested by phagocytosis. Gastrovascular Cavities Animals with a simple body plan have a digestive compartment with a single opening. Functions in digestion and distribution of nutrients. Cnidarians and many flatworms have gastrovascular cavities in which food and waste products of digestion enter and exit through same opening. Alimentary canals Most animals have a digestive tube that extends between two openings, a mouth and an anus. Food moves along the alimentary canal in one direction. Tube can be organized into specialized compartments that carry out digestion and absorption in steps. Can ingest food while digesting earlier meals. Earthworm: mouth sucks in food, passed through esophagus, stored and moistened in crop, crushed in gizzard, then digestion and absorption in intestine. Grasshopper: Digestive tube divided into foregut (esophagus and crop), midgut (gastric caeca and intestine) and hindgut. Gastric caecae are pouches where most digestion and absorption takes place Birds: Three separate chambers, crop, stomach, and gizzard. Crop and gizzard moisten and crush food. Chemical digestion and absorption takes place in the intestine. Mammalian Digestion Organs specialized for successive stages of food processing form the mammalian digestive system. Movement of food through the digestive system is controlled by peristalsis, the rhythmic contraction of the smooth muscle walls of the alimentary canal. Spincters, muscular ringlike valves regulate the passage of material between digestive compartments. Digestion begins in the Mouth Food in the mouth stimulates the secretion of saliva. Saliva lubricates food for swallowing. Saliva contains enzyme amylase, which hydrolyzes starch and smaller polysaccharides into the disaccharide maltose. Chemical digestion begins in mouth. Chewing is mechanical digestion. During chewing food is shaped into a ball called a Bolus During swallowing, the bolus enters the pharynx (the throat), a junction that opens to the esophagus and the trachea (windpipe). During swallowing a flap of cartilage – the epiglottis - covers the opening to the trachea and keeps food from going down the airway. Esophagus moves food to the stomach. The Stomach The stomach’s functions include storing food and secreting digestive fluid called gastric juice. Gastric juice is composed of Hydrochloric Acid: pH of 2; begins breakdown of meat and plant materials; also kills most bacteria Pepsin: enzyme that attack peptide bonds and breaks down proteins. Gastric Glands contain mucous cells that secrete mucus which lubricates and protects the cells lining the stomach, chief cells which secrete pepsinogen, and parietal cells that secrete HCL. The result of digestion in the stomach is a substance called acid chyme. Acid chyme is shunted from the stomach to the small intestine through the pyloric sphincter. Small Intestine The first section of the small intestine is the duodenum. Duodenum is the major site of chemical digestion. The intestine does not have mucus protection. The pancreas releases a bicarbonate fluid that buffers the acidic contents from the stomach. In the duodenum, acid chyme mixes with secretions from the pancreas and the liver. Bile Bile is made in the liver and stored in the gall bladder. Bile emulsifies fat – bile coats fat droplets, turning large fat droplets into small fat droplets, which are easier to digest. Bile enters the intestine at the duodenum. Chemical digestion in the duodenum Carbohydrates: pancreatic amylase breaks down starch, glycogen, and small polysaccharides into maltose. Maltose and other disaccharides are broken down by enzymes in the wall of the duodenum. Proteins: Trypsin and chymotrypsin break polypeptides into smaller chains. Dipeptidases, carboxypeptidase and aminopeptidase break polypeptides into amino acids. Duodenal digestion continued Nucleic acids: nuclei acids are hydrolyzed into nucleotides by nucleases that enter the duodenum from the pancreas. Fats: digestion of fats starts in the small intestine. Bile coats the fat droplets and keeps them from clumping. Lipase, produced in the pancreas hydrolyzes small fat droplets. Villi Lining of the small intestine has folds called villi, which are divided into microvilli. Together they greatly increase the surface area available for absorption. Each villus contains capillaries for absorption of monomers and a lymph vessel, called a lacteal which absorb fatty acids. Passive, facilitated diffusion, and active transport move monomers across the intestinal membrane into blood vessels. Hepatic portal vessel Capillaries and veins that carry nutrients away from the villi form the hepatic portal vessel. Hepatic portal vessel carries nutrients to the liver. Liver regulates the distribution of nutrients to the body. The remaining regions of the small intestine, the jejunum and the ileum function mainly in the absorption of water and nutrients. Large Intestine Large intestine, also called the colon is connected to the small intestine by a sphincter. The point of this connection is the site of the caecum, and the extension called the appendix. Functions of the large intestine are to compact waste and recover water. Harmless bacteria in the colon produce some vitamins. At the end of the colon is the rectum, where feces is stored until elimination. Evolutionary adaptations of vertebrate digestive systems correlate with diet. Mammals have specialized teeth that allow them to ingest their food. Herbivores have longer alimentary canals than carnivores. Many herbivores harbor large populations of symbiotic bacteria and protists than enable the digestion of cellulose.