Chapter 53: Population Ecology Essential Knowledge 2.a.1 – All living systems require constant input of free energy (53.3 & 53.4). 2.d.1 – All biological systems from cells and organisms to populations, communities, and ecosystems are affected by complex biotic and abiotic interactions involving exchange of matter and free energy (53.1 – 53.5). 4.a.5 – Communities are composed of populations of organisms that interact in complex ways (53.1-53.3, 53.5 & 53.6). Population Ecology Study of the factors that affect population size and composition. Population: Def: Individuals of a single species that occupy the same geographic area Ex: Humans living in Indianapolis, IN Important Characteristics 1. Density 2. Dispersion Density Number of individuals per unit area or volume. Ex: Diatoms - 5 million/m3 Trees - 5,000/km2 Deer - 4/km2 Dispersion Pattern of spacing among individuals. Types: 1. Clumped 2. Uniform/Even 3. Random Clumped Dispersion May result form a patchy environment. May increase chances for survival. Ex: Schooling behavior Flocks of birds Uniform Dispersion Often the result of antagonistic interactions between individuals. Known as even or regular dispersion Ex: Territories Spacing between desert plants Random Dispersion Often the result of the absence of strong attractions or repulsions between individuals. Not a common pattern. Demography The study of the vital statistics that affect population size. Ex: Birth and Death rates Factors of Demography: Age structure of population Birth and death rates Generation time Sex ratio and reproductive behavior Life Tables Mortality summary for a cohort of individuals. First developed from life insurance studies. What do they show us? Mortality rate per year Life span of the organism Fecundity (birth rate) Survivorship Curve Plot of the numbers of a cohort still alive over time. Curve Types: Type I Type II Type III Type I Low early deaths. High late deaths. Ex: Humans Other large mammals Type II Constant death rate. Ex: Annual plants Many invertebrates Type III High early deaths. Low late deaths. Ex: Trees Oysters Comment Curve type may change between young and adults. Ex: Nestlings - Type III Adult Birds- Type II Life History Strategies 1. "r" or Opportunistic species 2. "k" or Equilibrial species "r" Species Increase fitness by producing as many offspring as possible. Do this by: Early maturation Many reproductive events Many offspring Result Maximize reproduction so that at least a few offspring survive to the next generation. Most offspring die (Type III curve). "k" Species Increase fitness by having most offspring survive. Do this by: High parental care Late maturation Few reproduction events Few offspring. Result Maximize survivorship of each offspring. Few offspring, but most survive (Type I curve). What is the strategy? For a weed? For an endangered species? For Garden Pests? Population Growth DN/Dt = b - d Where: N= population size t = time b = birth rate d = death rate Rate of Increase r = difference between birth rate and death rate. r=b-d Equation with “r”: DN/Dt = rN N = population size t = time r = rate of increase From Calculus The equation DN/Dt = rN becomes: dN/dt = rmax N rmax = intrinsic rate of increase Exponential Growth dN/dt = rmax N Characteristic of "r" species. Produces a “J-shaped” growth curve. Only holds for ideal conditions and unlimited resources. Logistic Growth dN/dt = rmax N K-N K K = carrying capacity Result of logistic growth? “S-shaped” growth curve Characteristic of “k” species Common when resources are limited Comment K is not a constant value. Populations often oscillate around “K” as the environment changes. Additional Comments Populations often overshoot “K”, then drop back to or below “K”. AP Exam rarely asks you to work the equations, but you should be able to give them. Regulation of Population Size 1. Density- Dependent Factors 2. Density- Independent Factors Density-Dependent Affect is related to N As N increases, mortality increases Ex: Food, nesting space, disease Density-Independent Affect is not related to N Mortality not related to population size Ex: Weather and climate Population Cycles Cyclic changes in N over time Often seen in predator/prey cycles Ex: Snowshoe Hare – Lynx Causes? Density dependent factors Chemical cycles Saturation strategy to confuse predators Age Structure Diagrams Show the percent of a population in different age categories Method to get data similar to a Life Table, but at one point in time Importance Can be used to predict future population growth trends, especially for long lived species. Exponential Growth Produces age structures that are a triangle or pyramid shape Logistic Growth Produces age structures that have even sizes between most age categories Declining Populations Produce age structures with a narrow base and wider middles Summary Identify the difference between population density and dispersion. Recognize the types of dispersion patterns and the interactions that lead to them. Identify the types of survivorship curves. Recognize the characteristics of "r" and "k" life history strategies. Identify the types of population growth models. Identify factors that regulate population size. Recognize how age-structure diagrams relate to population growth.