BOTANY General Overview - - - - - - 4 Main Groups of Land Plants: o Bryophytes Non-Vascular Mosses Flagellated Sperm Gametophyte is the dominant phase; the sporophyte is dependent on the gametophyte o Pteridophytes Vascular but Seedless Ferns Flagellated Sperm Sporophyte is the dominant phase; the gametophyte is FREE-LIVING o Gymnosperms Vascular with Seeds Conifers (ex. Pine Trees) Non-Flagellated Sperm Sporophyte is dominant and the gametophyte is retained in the sporophyte o Angiosperms Vascular with Seeds, Fruits, and Flowers Flowering Plants Non-Flagellated Sperm Sporophyte is dominant and the gametophyte is retained in the sporophyte Seeds multicellular; sporophyte embryo and food supply; can be dormant; prevents against desiccation; aids in dispersal for the plant; develops from the ovule Flowers Specialized for reproduction o Sepals o Petals o Stamens (male reproductive part – produces microspores, which develop into the male gametophyte) o Carpals/Pistils (female reproductive part – produces megaspores, which develops into the female gametophyte) Seed forms from ovule Fruit forms from the ovary Double Fertilization pollen contains two sperm cells; one fertilized the egg to make the diploid zygote, one fertilizes the two polar nuclei to form the endosperm (food source); the purpose of double fertilization is to synchronize the development of the food storage with the development of the embryo Monocots vs. Dicots Monocots Dicots EmbryosOne Cotyledon (seed leaf) Two cotyledons (seed leaves) Leaf veinsParallel Netlike Stemsvascular bundles – random vascular bundles – ring RootsFibrous Taproot Flowers Multiples of 3 Multiples of 4 or 5 EvolutionMonophyletic Polyphyletic Plant Parts: o Roots anchor plant, absorb water (root hairs!), store food o Shoots stems and leaves o Stem nodes and internodes; axillary buds and terminal buds (meristems!) Plant Tissues: o Dermal Epidermis; can secrete waxy cuticle (on leaves!) o Vascular Xylem and Phloem Xylem – transports water; uses tracheids and vessel elements; goes “up” from roots Phloem – transports “food” (sugars); uses sieve tube cells and companion cells; goes from source to sink; active transport (translocation) o Ground Neither dermal nor vascular; photosynthetic, storage, support; pith (center) and cortex (outside) in dicots - - - - - Plant Cells: o Parenchyma least specialized; thin, flexible, primary cell walls; all cells start as parenchyma o Collenchyma thicker, but uneven primary cell walls; supports young parts of plants; no secondary cell walls o Sclerenchyma thick secondary cell walls made of lignin; supports plants; very rigid Meristems: o Allows for lifelong growth o Two types: Apical meristems found in the tips of roots and buds of shoots; allows for primary growth (length) Protoderm = makes dermal tissue (epidermis) Procambium = makes the stele (vascular tissue – primary xylem and phloem) Ground = makes ground tissue Lateral meristems thickens the plant; found in woody plants Vascular cambium = makes secondary xylem (WOOD!) and secondary phloem Cork Cambium = makes the tough covering for stems which replaces the epidermis (ex. Bark) o Primary growth order: Root cap zone of cell division zone of elongation zone of maturation Water Potential determines the direction of movement in plants o ALWAYS goes from higher to lower o Adding solute = lowers water potential o Adding pressure = increases water potential Movement in Vascular Tissue: o Xylem Root Pressure (push) – water flows into the stele (many solutes) and therefore pushes the water upward, can result in “guttation” Transpiration (pull) – due to adhesion/ cohesion/ H-bonding; water is lost by transpiration (evaporation) through the stomata and is therefore pulled up from other cells by osmosis; this is the MAIN mechanisms for xylem flow o Phloem Source to Sink Uses energy (translocation); occurs in sieve tube cells and is helped by companion cells Life Cycle = “Alternation of Generations” Sporophyte (2n) Meiosis Spores (n) Microspore – male Megaspore - female Mitosis Gametophyte (n) Pollen – male Embryo Sac - Female Mitosis Mitosis Zygote (2n) Vocabulary Terms to Know: Stomata Root Hairs Companion Cells Fertilization Guard Cells Translocation (phloem) Sieve Tube Elements/Cells Gametes (n) Sperm – male Egg - female Charophyceans Tracheids Stele Guttation Vessel Elements Mesophyll Tissue Signal Transduction Greening (roots, shoots and leaves) Growth hormones and enzymes for photosynthesis Hormones Tropism growth response toward or away from a stimulus Auxin Cytokinins Gibberellins Abscisic Acid Ethylene Brassinosteroids Elongation and apical dominance Commercially for herbicide or fruit growth Induce growth, stimulate germination Can delay senescence (aging) Stimulate growth in leaves and stems Flower and fruit development including bolting Maintains dormancy (inhibits germination) Reduces drought stress by closing stomata Gas produced during stress (obstacle) and ripening (positive feedback) Role in apoptosis and leaf abscission Similar to steroid hormones in animals Promotes cell elongation and retards leaf abscission Light responses Photomorphogenesis (growth and development) blue and red (phytochromes) Circadian rhythym cycle with 24 hour frequency Photoperiodism relative length of day/night and physiological response (critical night length) Other responses Gravity (root – positive gravitropism; shoots – negative gravitropism) Stress Drought reduce transpiration (guard cells lose turgor, stomata close) Heat heat shock proteins help scaffold folding Cold change in membrane fluidity (increase unsaturated fatty acids) Plant Defense Herbivory : physical and chemical Pathogen: gene to gene (avirulent) hypersensitive (localized) SAR (systematic acquired resistance)