Ms. Moore 2/11/13 WORMS AND MOLLUSKS What is a flatworm? Phylum: Platyhelminthes Flatworms are soft, flattened worms that have tissues and internal organ systems. They are the simplest animals to have 3 embryonic germ layers, bilateral symmetry, and cephalization. Acoelomates: without coelom (fluid-filled body cavity, lined with tissue from mesoderm Flatworms: Form and Function Feeding: Carnivores or Scavengers ; can be parasitic Digestive cavity with single opening (mouth) Pharynx: extends outside the mouth and pumps food into digestive cavity (gut) Food diffuses from the digestive cavity into all other body tissues Respiration, Circulation, and Excretion: Since their bodies are so flat and thin, many flatworms do not need a circulatory system to transport materials (use diffusion). No gills or respiratory organs; no heart, blood vessels, or blood. Flame cells: specialized cells that remove excess water from the body; filter and remove ammonia and urea using pores of the skin Response: Ganglia: groups of nerve cells that control the nervous system (no brain) Eyespot: group of cells that can detect changes in the amount of light in their environment Movement: Cilia on the epidermal cells help glide through the water. Muscles controlled by the nervous system help to twist and turn to react to environment. Reproduction: Hermaphrodite: both male and female reproductive organs Sexual: two worms join in a pair and they deliver sperm to each other Asexual: fissionorganism splits in two and each half grows new parts to become a complete organism Groups of Flatworms Turbellarians Flukes Tapeworms Turbellarians Free-living flatworms Most live in marine or fresh water Bottom dwellers: living in sand or mud Planarians: “cross-eyed” freshwater worms Flukes Class: Trematoda Parasitic flatworms that infect internal organs of their host; can also be external parasites. Tapeworms Class: Cestoda Long, flat, parasitic worms that are adapted to life inside the intestines of their hosts. Scolex: contains suckers or hooks; attaches Proglottids: segments that make up most of worm’s body; contain male and female reproductive organs Youngest proglottids are at the anterior end and the largest and most mature are at t he posterior. After eggs have been fertilized, proglottids break off and release zygotes that are passed out of the host in feces/ Testes: fertilize eggs of other tapeworms or of self What is a Roundworm? Phylum: Nematoda Roundworms are slender, unsegmented worms with tapering ends; Range in size from microscopic to a meter in length Pseudocoelom: false coelom (only partially lined with mesoderm Digestive tract with two openings—mouth and anus (posterior opening of digestive tract) “tube within a tube”: inner tube is digestive tract and outer tube is body wall Food moves in one direction Roundworms: Form and Function Feeding: Carnivorous: eat small animals by latching on to them with grasping mouth parts and spikes Scavengers: eat algae or decaying mater Consume bacteria and fungi The free living roundworms tend to be more complex than parasitic roundworms. Respiration, Circulation, and Excretion Diffusion through body walls Response Simple nervous systems with several ganglia; sense organs that detect chemicals given off by prey or host Movement Muscles extend length of body; function as hydrostatic skeleton Reproduction Sexually with male and female worms Internal fertilization Roundworms and Human Disease Trichinosis-Causing Worms Caused by Trichinella roundworm Worms burrow into intestine walls and females release larvae that travel through the bloodstream and live in organs and tissues of host’s body Filarial Worms Found in tropic regions of Asia; live in blood and lymph vessels of birds and mammals (humans) Transmitted host-to-host by biting insects like mosquitoes Large numbers could block lymph passageselephantiasis Ascarid Worms The cause of malnutrition of more than 1 billion people worldwide. Ascaris lumbricoides usually spread by eating vegetables that are not washed properly. Hookworms 25% of the world’s population is infected with these worms Eggs hatch outside the body and mature in the soil Use tooth-like plates to burrow into skin of an uncovered foot and live in bloodstream Suck blood and cause weakness and poor growth Research on C. elegans DNA sequence has been mapped out (97 million bp) Help us find out how eukaryotes become multicellular and how multicellular animals are similar and different What is an Annelid? Phylum: Annelida; “little ring” Septa: internal walls between each body segment Setae: bristles attached to each segment Annelids are worms with segmented bodies Have true coelom that is lined with tissue derived from mesoderm. Like roundworms, annelids have a tube within a tube system with an anus. Annelids: Form and Function Feeding and Digestion Range from filter feeders to predators Use a pharynx that hold two or more sharp jaws used to attack prey. Earthworms: pharynx pumps food into esophagus moves to crop (storage) through gizzard (ground into smaller pieces) intestine Circulation Closed Circulatory System: blood is contained within a network of blood vessels Earthworm: blood circulates through two major blood vessels Dorsal runs to head; ventral runs to tail Dorsal functions as a heart due to contractions = pump blood Respiration Aquatic annelids breathe through gills Land-dwelling annelids take in oxygen and give off carbon dioxide through moist skin; mucus Excretion Digestive waste passes through the anus Cellular waste eliminated by nephridia (excretory organs that filter fluid in the coelom) Response Brain and nerve cords Marine annelids are more developed: sensory tentacles, chemical receptors, statocyts, two or more pairs of eyes Movement Hydrostatic skeleton Longitudinal muscles: front to rear; muscles contract to make worm shorter and fatter Circular muscles: contract to make worm longer and thinner Marine annelids: paddle like appendages (parapodia) Reproduction Sexually: external fertilization Hermaphrodites: two worms exchange sperm and store them in special sacs Clitellum: band of thickened, specialized segments, secretes a mucous ring into which sperm and egg are released; rings slips off and forms a cocoon; worms hatch weeks later Groups of Annelids Class: Oligochaetes Annelids that typically have streamlined bodies and relatively few setae; soil or freshwater Castings: mixture of sand, clay and undigested food that an earthworm expels from its anus. Class: Hirudinea Leeches External parasites that suck the blood and body fluids of their host ¼ are carnivorous that that feed on soft-bodies invertebrates Suckers at both ends of body Proboscia: muscular extension that can be forced into tissue of host Can release a substance that anesthetizes wound and prevents blood from clotting. Class: Polychaeta Polychaetes Marine annelids that have paired , paddle like appendages tipped with setae Ecology of Annelids Earthworms (and other annelids) burrow through soil, aerating it and mixing it to depths of two meters or more Mine minerals from deeper soil layers Diets of many birds and other vertebrates What is a Mollusk? Phylum: Mollusca Mollusks are soft-bodied animals that usually have an internal or external shell. Ex: snails, slugs, clams, squids, and octopi. Trochophore: free-swimming larval stage Characteristic of Annelida = related 550 m.y.a. Form and Function: Mollusks True coeloms and organ systems Body Plan (4 basic parts): Foot: muscular structure used for crawling, burrowing, and capturing prey Mantle: thin layer of tissue that covers the body (cloak) Shell: glands in mantle secrete calcium carbonate; reduced or lost in slugs and other mollusks Visceral Mass: beneath mantle; consists of internal organs (Figure 27-21) Feeding: Can be herbivores, carnivores, filter feeders, detritivores, or parasites Radula: flexible tongue-shaped structure used by snails and slugs; 100s of teeth are attached Octopi use sharp jaws and tentacles to feed Clams, oysters, and scallops filter feed with their gills and mucus Siphon: tube like structure where water enters and leaves the body Respiration: Use gills inside their mantle cavity Land species have no gills, but they do have thin blood vessels in mantle that stay moist for oxygen passage. Circulation: open or closed Open circulatory system: blood is pumped through blood vessels by a simple heart; the blood makes way through body to the gills, then back to the heart; found in slow moving mollusks Closed circulatory system: used in fast-moving mollusks bc it moves blood through body faster Excretion: Cells release wastes into blood and nephridia remove it from the body Response: Slugs use simple ganglia and octopi use complex brain Complex brain allow them to remember things and be trained for reward or avoid punishment Movement: Slugs secrete mucus and use their foot to glide Octopi use jet propulsion Reproduction: Sexually by internal or external fertilization Can be hermaphrodites, but do not fertilize own eggs Groups of Mollusks Gastropods Class: Gastropoda Gastropods are shell-less or single-shelled mollusks that move by using a muscular foot located on the ventral side EX: snails, land slugs, sea butterflies, and sea horses Protection: some can retract back into their shell or use ink to make a smoke screen; some produce chemicals that make them taste bad; nudibranchs can recycle nematocysts and use them Bivalves Class: Bivalvia Bivalves have 2 shells held together by one powerful muscle EX: clams, oysters, mussels, and scallops Most stay in one spot for long periods of time, except scallops (move to run from predators) Eat through filter feeding or sifting through the mud. Cephalopods Class: Cephalopoda Cephalopods have a head that is attached to their foot which is divided into tentacles or arms (8+). EX: octopus, squids, cuttlefishes, nautiluses Nautiluses are the only ones with external shells; they have over 90 tentacles; control water depth by amount of gases in their mantle Octopi have lost shells completely Cuttlefishes have small shells inside their body Ecology of Mollusks Mollusks can be used to detect water quality Filter algae out of the water Live symbiotically with other organisms Major food source for humans