Running head: MODERN HUMAN ORIGINS Modern Human Origins Jordan Woodbury English 1010 29 April 2015 1 MODERN HUMAN ORIGINS 2 Modern Human Origins Utilizing fossil evidence and what scientists have discovered about the behaviors and patterns displayed by Homo heidelbergensis, and later Neandertals and modern humans, they can make very interesting inferences about which group, if just one, modern day humans owe their life to. There are two popular theories about human origins and both have merit in their own rights. The regional continuity model, or multiregional model, follows the theory that modern humans evolved from separate populations of pre-modern humans all over the globe. The second theory, the replacement model, follows a group, or groups, of humans, recently evolved from African Homo heidelbergensis, that immigrated into Europe and Asia to later out-compete and replace Neandertals and Asian Homo heidelbergensis populations. Starting with the first theory, logical evidence supporting the regional continuity model and some of its roots will be given. The regional continuity model, highly influenced by Milford Wolpoff, claims that local populations in Asia, Europe and Africa evolved into modern humans almost independently. For this to be true, there are a couple of ideas that have to be taken as truth. First, the theory suggests that more fossils have yet to be found of modern humans in Europe and Asia. These new fossils will be found to be around two hundred thousand years old, proving that the oldest modern humans are not just found in Africa. Second, the theory implies large amounts of gene flow between the populations took place, ensuring that no speciation would separate these groups and each population would evolve, roughly, in the same direction. In order to answer the question of Neandertals being European ancestors, scientists utilized fossil evidence and nuclear DNA as well as simple evolutionary theory to prove their relation (Wolpoff, 2011). This explains that, while theory and fossil evidence are helpful, the final straw was the discovery of ancient nuclear DNA of Neandertals. Utilizing the newly MODERN HUMAN ORIGINS 3 sequenced genome of Neandertals and immediately comparing the results with modern humans, scientists found that European and Asian populations still have small amounts of Neandertal DNA (Wolpoff, 2011). “Neanderthals have contributed approximately 1% to 4% to the genomes of non-African modern humans” (What does it mean to be Human?, 2015). While the multiregional evolution model has merit and some evidence to back it up, so too does the replacement model. The replacement, or “out of Africa” model is a look at Homo sapiens' evolution in Africa and their developments afterward. This theory, outlined in (Human Origins Evolution and Diversity, 2013, pp. 284-287), suggests that Homo heidelbergensis living in Africa evolved into the first Homo sapiens. After the evolution, these humans left Africa as they started entering Europe and parts of Asia approximately 60,000 years ago. This theory explains why modern humans are so similar to one another and what happened to the Homo heidelbergensis groups living in the Eastern Europe region along with sections of Asia. It can also help to explain the absence of heidelbergensis features and characteristics in today's species (Jurmain, Kilgore, & Trevathan, 2013). With all of the newfound evidence, it has sparked countless arguments and counter arguments leading to a very big debate as to where the human origins truly lie. Whether it be the isolated evolution of separate species along the same timeline, or one species dominating the landscape and becoming absolute as they spread throughout Europe. The most suitable theory is a combination of the two. Recent evidence has given birth to another iteration of the replacement modern that is called the partial-replacement model, also called the assimilation model. As the newly evolved humans started immigrating into Europe, it is theorized that they exterminated Neandertals and other pre-modern hominids. This extermination was not necessarily of a violent nature, but rather a competition for resources and other necessities that MODERN HUMAN ORIGINS 4 humans won in the end. This does not exclude the definite possibility of violent killings between the groups, but since there is Neandertal DNA among modern European and Asian populations, it has to be accepted that some interbreeding was taking place between these new Homo sapiens and Neandertals (Wolpoff, 2011). The partial-replacement model, shown in (Human Origins Evolution and Diversity, 2013, pp. 297-299), acknowledges that no complete extermination of other hominid species happened, but more of an assimilation process. Homo sapiens leaving Africa experienced interbreeding with separate hominid species and because of their superior technology and tools, they were able to stand the test of time as a species. It's important to note that any breeding between hominid species and Homo sapiens occurred after they had left Africa and moved into western Asia and Europe. Very recent findings are shaking the roots of the “out of Africa” theory, claiming that humans are being found in areas like China and Europe from as late as 125,000 years ago (Roberts, 2010). If these new findings continue to compound, the total replacement model may have to adapt if it is to keep afloat in this time of constant discovery and evolution (Jurmain, Kilgore, & Trevathan, 2013). The partial-replacement model, or assimilation model, has the strongest theory and the most supporting evidence and groundwork. It is difficult to believe that multiple populations were able to evolve to become almost identical in a genetic sense without having originated in the same environment and circumstances. Even accounting for substantial gene flow, this wouldn’t be enough for natural selection to work on such a large scale and on multiple continents. The “out of Africa” model is not without fault. It has been quickly losing support due to the fossil and DNA evidence that continues to stack against it. “It is apparent that both the complete replacement and the regional continuity models have difficulty accounting for all of the MODERN HUMAN ORIGINS 5 fossil and genetic data” (O'Neil, 2013). Evidence points to modern humans first evolving in Africa. As they migrated into other European areas, rather than replacing pre-modern species, they interbred to create offspring with both sets of genetic DNA (Janette & H.J., 1999). This is why Neandertal and even other species within the Homo genus can be found in the DNA of today's humans. African populations likely made their migration in a very gradual process as they slowly migrated north and assimilated various populations along the way (Falsetti, Smith, & Donnelly, 2005). Brauer also admits that, while there is observable local continuity in European areas like the Neandertal facial characteristics, modern Eurasians were largely due to migrations of African populations into parts of the Old World. With real evidence working to disprove both the regional continuity model and replacement model, the best informed and most suitable contender of all of the theories of modern human origins is the assimilation model. Anthropologists will continue to tirelessly chip away at the edges, searching for new evidence to push any one of these theories to the next step. While waiting for more solid evidence, however, all that can be done is to work with is the evidence that is already unearthed and the theory that is ever changing. MODERN HUMAN ORIGINS 6 References Falsetti, A. B., Smith, F. H., & Donnelly, S. M. (2005, June 14). Modern Human Origins. American Journal of Physical Anthropology, 35-68. J. D., & H. D. (1999). Fossil Evidence for the Emergence of Modern Humans. Jurmain, R., Kilgore, L., & Trevathan, W. (2013). Human Origins Evolution and Diversity. CENGAGE Learning. O'Neil, D. (2013). Early Modern Homo Sapiens. Retrieved from Anthro.palomar.edu: http://anthro.palomar.edu/homo2/mod_homo_4.htm Roberts, A. (2010). The Incredible Human Journey. Bloomsbury. What does it mean to be Human? (2015, 4 11). Retrieved from Smithsonian National Museum of Natural History: http://humanorigins.si.edu/evidence/genetics/ancient-dna-andneanderthals/sequencing-neanderthal-dna Wolpoff, M. H. (2011). Evolution or Extinction of Neandertals: A Brief History. Journal of Cosmology, 14.