It is not the strongest of the species that survives, nor the most intelligent that survives. It is the one that is the most adaptable to change. Charles Darwin Background extinction rate Typical low rate of extinction Mass extinction Significant rise above background level Fig. 4-13, p. 90 Starting in 1958, Mao Zedong, leader of the Communist Party of China, initiated a series of policies to transform the country into a modern, industrialized, communist society. One of the first actions taken was known as the Four Pests Campaign. This campaign sought to eliminate rats, flies, mosquitoes, and sparrows. 5 Masses of people were mobilized to eradicate the Eurasian Tree Sparrow. Tactics included: Banging pots and pans, preventing the birds from landing, until they were exhausted. Tearing down nests. Shooting them from the sky using guns and sling shots. “Everyone come and fight sparrows.” 6 The sparrows were hunted because they ate grain seeds; reducing crop yields. By April of 1960, Chinese leaders came to realize that the sparrows also ate a large number of pest insects, including locusts. 7 The government made a series of other poor agricultural decisions at this time, including: Ordering farmers to increase the density of their planting by 6 times, believing that the same species of plant would compete with itself. Deeper plowing of the soil, which brought up sand and rocks instead of more topsoil. 8 The locust plague, overplanting, and overplowing combined with a severe drought. The number of victims is unknown, but estimated between 20-43 million. 9 Ecology is the study of the interactions between living organisms and their surroundings, including their physical environment and other forms of life. Ecology is only one part of the overall study of life, matter, and the universe. 10 An organism is an individual living thing. A single species of organism is one that is similar enough to breed and produce healthy, fertile offspring. A population includes all members of a species that live in the same area at the same time. The biological community is made of all populations living and interacting in one area. 11 An ecosystem includes the biological community and its surrounding physical environment. Abiotic (nonliving) factors like soil, precipitation, etc. The biosphere is the part of Earth that supports life - - all ecosystems are found and sustained here. 12 Each species plays a specific ecological role called its niche Any given species may play one or more of five important roles—native, nonnative (exotic), indicator, keystone, or foundation species—in a particular ecosystem Ecological niche Pattern of living, way of life or role a species plays in its ecosystem Generalist species Broad niche - can live in many different places and eat a variety of foods, and tolerate a broad range of environmental conditions Ex. Cockroaches, rats, mice, humans Specialist species Narrow niche – consumes just one or only a few types of food, and can live in only a narrow range of conditions Ex. Some shorebirds, Pandas Number of individuals Specialist Species and Generalist Species Niches Specialist species with a narrow niche Niche separation Generalist species with a broad niche Niche breadth Region of niche overlap Resource use Fig. 4-15, p. 92 Cockroaches Generalists galore High reproductive rates Can live in pretty much any conditions…your house, your bed, hot, cold can survive nuclear fallout, can hold breath for 10 minutes, and can live without their head for about a week. Native species Normally live in an ecosystem Nonnative species Not native to the ecosystem (introduced, alien or exotic species) – not all are villainous! Indicator species Are usually found almost everywhere and are affected quickly by environmental changes Provide early warning of damage to a community Can monitor environmental quality Examples: Lichens (sensitive to pollutants, like factory smoke), Macroinvertebrates (some are sensitive to water pollutants/low quality water), Amphibians (sensitive to pesticides, habitat fragmentation), Birds (pesticides/habitat fragmentation) Keystone species Have a large effect on the types and abundances of other species Can play critical roles in helping sustain ecosystems Ex. Bees and pollination (essential for plant reproduction, and therefore essential for LIFE) Top predators (keep populations of other species in check) Remember the Coyote removal activity? Biologists describe foundation species as “engineers of ecosystems.” The activities of foundation species physically modify the environment and produce and maintain habitats that benefit other organisms that use those habitats. Examples: Corals build coral reefs that many other species use. Beavers harvest trees and create dams, thus creating wetlands that other species utilize. American Alligators dig gator holes that become microcosm ecosystems Each organism has a particular type of environment where it can survive. This is its habitat. Habitats have a range of factors that influence the community that lives there – temperature, precipitation, etc. These factors can LIMIT the growth, abundance, or distribution of an organism or a population of organisms in an ecosystem, and are therefore called limiting factors Some organisms have a single critical factor that plays the greatest role in determining its range. Critical factors have a “Goldilocks Effect”, meaning there can be too much or too little of it. 22 When the right level of an environmental factor is present, population levels will be growing or at their peak. This is the optimal range for that factor. At the zone of physiologic stress, levels of the factor are too high or too low. The population barely survives. At the zone of intolerance, the population dies out. 23 Why do coniferous trees have such a wider range (3,5,6,7,8) than broadleaf deciduous trees (8)? Why are broadleaf trees more prevalent than coniferous trees in areas where they can both survive? 24 The range of tolerance for a species is largely defined by the presence of physical, behavioral, or physiologic adaptations. Physical adaptations are structural differences in coloration, body shape, musculature, etc. Behavioral adaptations include migration, or marking a territory. Physiologic adaptations, such as skin tanning, occur at the cell or tissue level in an organism. The gorilla is adapted for living and feeding on the ground, while chimpanzees gather food from trees. 25 All of the unique adaptations found in different organisms are the result of evolution -- small, advantageous mutations that have accumulated over countless generations. Natural Selection describes the process where individuals with better genes survive and reproduce more successfully, while those with weaker genes do not. The primary source of this genetic variety is random mutations. These are small DNA changes. Usually these changes produce no effects or bad effects, but occasionally, they can be quite beneficial. 26 Evolutionists will cite three categories of evidence in support of the theory. Physical Similarities Most animals have similar bones in their limbs (fins, arms, wings). These similarities can also be found between living species and fossils. Comparing DNA Chimpanzees, bonobos, and humans share about 99% of the same DNA. Vestigial Structures Still exist in the body but are no longer needed Ex: Appendix, wisdom teeth, brains (that’s a joke) 27 An example of natural selection in the human race can be seen in sickle-cell anemia. Normal red blood cells have a biconcave –disc shape. Sickle-shaped red blood cells are the result of a single mutated gene. Sickle cells can get stuck in small blood vessels, causing tissues to become oxygen-deprived. 28 Individuals with sickle-cell anemia or carriers are also highly resistant to malaria. The mutation is a disadvantage in climates where malaria is absent due to the long-term health issues. The mutation is an advantage in climates where malaria is present due to the resistance it provides. Sickle cell gene frequency in Africa. Source: University of Oxford study. Distribution of malaria transmission in Africa. Source: American Journal of Tropic Medicine and Hygiene Study. 29 Evolution is an very slow process. Current life on Earth represents billions of years of small mutations and natural selection (due to selective pressures). Over a long enough period of time, enough mutations occur that a new species emerges. When a population splits and become two different species, it is called divergent speciation. 32 Geographic Isolation Can Lead to Reproductive Isolation – results in Speciation Adapted to cold through heavier fur, short ears, short legs, and short nose. White fur matches snow for camouflage. Arctic Fox Northern population Early fox population Different environmental conditions lead to different selective pressures and evolution into two different species. Spreads northward and southward and separates Gray Fox Southern population Adapted to heat through lightweight fur and long ears, legs, and nose, which give off more heat. Fig. 4-12, p. 88 Chimpanzees and bonobos are very closely related primates, with only a few differences between the species. They likely diverged during a severe drought millions of years ago. 34 PBS Evolution – Chimps vs. Bonobos 35 Four ecological factors (called “selective pressures”) will “encourage” natural selection to favor certain individuals in a population. Speciation is more likely to occur in these situations. Physiological stress, inappropriate levels of a critical environmental factor. Moisture, Light, pH Predation, when one organism is hunted and killed by another. Competition, the result of other organisms attempting to use same resources. Sexual Selection occurs when the female (usually) responds to specific behaviors or physical traits. 36 PBS Evolution – Peacock Experiment 37 Convergent speciation can occur when natural selection favors unrelated organisms to evolve to look similar. This is also called Convergent Evolution. Mantis Fly Preying Mantis 38 The selective breeding of organisms by humans for characteristics desirable to them. 39 Evolution has produced a tremendous amount of biodiversity on Earth. Biodiversity is defined as the number of different species within an ecosystem or area. The number of species is unknown. About 1.5 million have been identified. A recent study estimates 8.7 million exist currently. 40 The majority of known species are insects, followed by fungi and bacteria. 41 The biodiversity found in genes, species, ecosystems, and ecosystem processes is vital to sustaining life on the earth Functional Diversity The biological and chemical processes such as energy flow and matter recycling needed for the survival of species, communities, and ecosystems. Genetic Diversity The variety of genetic material within a species or a population. Ecological Diversity The variety of terrestrial and aquatic ecosystems found in an area or on the earth. Species Diversity The number and abundance of species present in different communities. Fig. 4-2, p. 79 The Number and variety of species in a given area is called species diversity (combo of species richness and evenness) The number of different species in a given area is called species richness The comparative (abundance) number of individuals of each species present is called species evenness For example, if a scientist found 20 different plant species in a 1 m2 area, but only ONE of each species, it could be said that there is a high species evenness, and a high species richness. Habitat fragmentation is the process by which habitat loss results in the division of large, continuous habitats into smaller, more isolated remnants. The more habitat is fragmented, the more biodiversity suffers There is a theory that larger habitat “islands” are more holistically healthy than smaller islands. This theory is called island biogeography. In bigger habitat areas, all aspects of biodiversity remain more intact for that reason, scientists have tried to build corridors between habitat fragments, connecting them with more habitat 45 With the tremendous diversity of living organisms, a system was devised to organize them based on their interrelatedness. Domain Kingdom Phylum Class Order Family Genus Species Taxon Reason Domain Eukarya Cells have nuclei and organelles (eukaryote). Kingdom Animalia Multicellular. Unable to produce own food (heterotrophic). Phylum Chordata Have a nerve cord along the back. Class Mammalia Warm-blooded, gives birth to live young, has hair. Order Primates Forward-facing eyes, enlarged brains, vertical posture. Family Hominidae Capacity for language, culture, empathy. The final two taxa, genus and species, are used to define the species’ scientific name. Homo sapiens Italicized or underlined Genus capitalized, species lower-case. 47 The scientific naming system is important, as many species have multiple common names. The cougar holds the Guinness world record for number of common names, with 40 in English alone! Cougar, catamount, painter, panther, ghost cat, puma, shadow cat, mountain lion, deer tiger, devil cat, sneak cat, plain lion, fire cat, mountain screamer, Florida panther, silver lion…… Puma concolor 48 Two animals with many classification levels in common are considered very closely related – they diverged recently. Wolves and dogs share the same domain, kingdom, phylum, class, order, family, and genus. Canis lupus Gray Wolf Canis familiaris Domesticated dog 49 As their ecosystem and communities change, populations of organisms undergo different growth patterns. The most common growth pattern is called logistic growth, which takes the shape of an “S”. 50 Logistic growth starts off fast, but gradually slows down as the population encounters environmental resistance (factors that act to limit the growth of a population) . Included in env resistance are limiting factors, chemical and physical factors that determine the number of organisms in a population. Environmental resistance can come in two forms: Density-dependent factors, such as disease, which affect dense populations more. Density-independent factors, such as natural disasters or climate change, which similarly affects all populations. 51 The density-dependent growth-limiting factors will cause the population’s growth to slow and eventually stabilize. The point at which it stabilizes is the carrying capacity, or the maximum population size that can be sustained by the ecosystem. 52 China’s great famine in 1958 was partially due to the removal of a density-dependent growth limiting factor for locusts – the predatory Eurasian tree sparrow. 53 Locusts were able to grow exponentially, creating swarms with millions of individuals. 54 Exponential growth, which takes the shape of a “J”, does not experience growth-limiting factors. The population will continue to grow, with the growth rate increasing over time, eventually exceeding the carrying capacity. This is called an overshoot. When a population overshoots the carrying capacity, it experiences a dieback, often in the form of mass-starvation. Exponential growth is unusual, and does not typically occur under normal conditions. 55 r-Selected species, opportunists K-selected species, competitors R strategists usually create an abundance of offspring in the hopes that a few will make it. These species usually have a very short maturation time, often breed at a very young age, have a short lifespan, produce many offspring very quickly, have young with high mortality rates and invest relatively little in parental care. The parents do not focus on passing down memes, units of cultural information, to their young. Instead the behavior of the young is determined by their genes. The young are precocial, meaning that they often can make it on their own without any instruction from their parents. Examples of r-selected species include bacteria, insects, and fish. K strategists are very different in that they attempt to ensure the survival of their offspring by investing time in them, instead of investing in lots of them. It is a reproductive strategy that focuses on quality over quantity. K strategist have relatively few offspring and make an effort at being good parents. Their young are altricial meaning that they cannot survive on their own until they reach adulthood. This extended period of maturation is used for memetic transference- the parents teach the young so that they can go on to reproduce themselves. K strategists are known to have a relatively long life span, produce relatively few offspring, the offspring have low mortality rates and the parents provide extensive parental care. The offspring are also relatively intelligent so that they can internalize the lessons from their parents. K-selected species include elephants, apes and whales. Humans are perhaps the most K-selected because their young are truly helpless- they necessitate a full two decades of parental care and tutelage and the parents usually only produce one offspring at a time. Five types of species interactions—competition, predation, parasitism, mutualism, and commensalism—affect the resource use and population sizes of the species in an ecosystem How quickly populations grow and the maximum size they attain are often the result of interactions with other populations of organisms. One of the most basic interactions is predator-prey (predation), where one organism consumes the other. 63 Competition, where organisms and populations compete for resources, is another common interaction. Water, food, territory, mates. When the competition occurs within members of the same species, it is called intraspecific. When the competition occurs between different species, it is called interspecific. 64 Another significant cause of the great famine was planting too densely. This caused the rice plants to compete with each other directly for water and soil nutrients. 65 Today, textbooks in China omit discussion of the true causes of the famine, instead citing “three years of difficulty” caused by “bad weather.” Mao Zedong’s portrait still hangs at the center of Tiananmen Square. 66 Some organisms have developed adaptations to avoid directly competing with their own species. Plants will send their seeds far away to avoid sharing the same soil. Wolves mark and occupy a territory so they have enough hunting area. Adult monarch butterflies and their caterpillars each utilize a different part of plants (this is called resource partitioning). 67 The closest relationships of all are symbiosis. These involve a lot of close contact between two species. When the relationship benefits both organisms, it is called mutualism. When one organism benefits while the other is unaffected, it is called commensalism. When one organism benefits at the expense of the other, it is called parasitism. When one organism benefits at the expense of another but ultimately ends up killing or consuming the host, it is called parasitoidism. 70 Barnacles create homes by attaching themselves to whales. The whales are unaffected. 71 Clownfish have a mucus coating that allows them to live in sea anemones. Their presence attracts other fish for the anemone to eat. 72 As bison walk through grass, insects are disturbed and fly away. They are eaten by cowbirds. 73 Ostriches and gazelles feed next to each other. Ostriches have excellent eyesight, while gazelles have stronger senses of hearing and smell. 74 Mistletoe extracts water and nutrients from the spruce tree directly. 75 76 Primary succession occurs when a new ecosystem develops where there was none before. A combination of wind, water, and pioneer species such as lichens break down rock into soil. Once the soil has enough organic matter, small plants and shrubs can be supported. Over time, trees spout and become dominant. 77 Secondary succession occurs following the disruption of an existing ecosystem. Fire, flood, volcanic eruption, clear-cutting, etc. This form of ecological succession does not take as long. Soil is already in place, and pioneer species appear within days or weeks. 78 Biomes are dynamic – they change as the Earth changes. This process is called succession. Organisms that thrive during the early stages of succession are called pioneer species. Those only found in later stages are called climax species. Ecosystem succession takes two forms, depending on the starting point. 79