Evolution The Unifying Principle of Biology • http://www.pbs.org/wgbh/evolution/educato rs/teachstuds/svideos.html Background • Each branch of physical science has at its core one or more unifying principles. These principles provide predictions that can be (and have been extensively) tested and verified experimentally. – – – – – Those that study these sciences do not blindly assume that these are true – every experiment, every piece of data, provides a new test for the principle. Remember that hypotheses are grouped together into theories. “A theory” is a laundry list of hypotheses under a single umbrella. Successful theories make predictions that are experimentally verified through many lines of independent work. Data that refutes theories results in revisions of the theory. Through many revisions, a theory eventually becomes solid enough that additional experimention continually validates the underlying hypotheses again and again. Over the course of time, theories mature into principles, or laws. Physics General Relativity, Quantum Mechanics Chemistry Atomic Theory, Thermodynamics Geology Plate Tectonics Biology Cell Theory, Gene Theory, Evolution Theory Common misconceptions • Evolution Theory does not describe how life began. • Evolution theory says nothing about the origins of the universe, the big bang, or the laws of thermodynamics. • The only concept that evolution addresses is the diversity of life on earth. Not its origins, not its future. It simply explains why the family tree of life is so branched. Part I Historic Perspective Popular Beliefs up through the 1800s Commonly held beliefs were not based on looking at the world, or examining nature. They were based on stories handed down from generation to generation, in both oral and written forms. The Bible and Catholic dogma was the most influential force on thought. Therefore, when the world and nature was examined, anything that was observed was interpreted according to what was already believed. Beliefs based on these forces: The earth was young. Species were made and did not change. When observations were made that did not fit what was already believed, new theories were contrived to fit the data into the context of what was already believed. • • • • • Fixity of Species Lamark: The first cohesive theory Darwin: Natural Selection Mendel: factors of inheritance Modern Synthesis: Connected Darwin and Mendel Jean-Baptiste Lamarck (1744-1829) A major contributor to Cell Theory One of the first to use “biology” in its modern sense Started out as an “essentialist” who believed species did not change. But his own work on mollusks convinced him that change had to have occurred. Was the first to propose a cohesive theory of evolution (inheritance of acquired characteristics) George Cuvier (1769-1832): paleontologist, father of comparative anatomy He worked with fossils (he found and named pterodactyl in Bavaria). He thought Lamark’s ideas were ridiculous, and firmly believed in the fixity of a species. He came up with the principle of “correlation of parts” – The notion that comparative anatomy was so accurate that after inspecting a single bone, the class and sometimes genus of an animal could be inferred. This is because the number, direction, and shape of the bones composing each part of a body are always in a necessary relation to all parts such that a great deal can be inferred with only a few bones. That was in 1798. Modern Synthesis • The modern synthesis solved difficulties and confusions caused by the specialisation and poor communication between biologists in the early years of the twentieth century. Discoveries of early geneticists were difficult to reconcile with gradual evolution and the mechanism of natural selection. The synthesis reconciled the two schools of thought, while providing evidence that studies of populations in the field were crucial to evolutionary theory. It drew together ideas from several branches of biology that had become separated, particularly genetics, cytology, systematics, botany, morphology, ecology and paleontology. Part II Natural Selection • • • • • • • • • • • • • • • • 1. heritable variation functional, phsical, behavioral variations. IT IS COMPLETELY RANDOM 2. more offspring are produced than the environment can support. Organisms struggle to exist. The work of Malthus: humans reproduction is held in check by death and famine due to the fact that populations tend to increase faster than their food supply does. Darwin applied this concept to ALL of life. reproductive potential of elephants: assuming 100 year lifespan with breeding of 30-90 years, a single female would bear ~6 young. If all of these survive and reproduce at the same rate, after only 750 years, a single pair of elephants would produce 19 million. 3. some individuals survive better than others as a result of their variations the most fit capture a disproportionate amount of resources. This is converted to a disproportionately larger number of offspring that also survive. What determines fitness differs for different populations (it depends on the environment) Humans carry out artificial selection (breeding) THERE ARE OTHER MECHANISMS BESIDES NATURAL SELECTIION: gene flow, nonrandom mating, genetic drift ONLY natural selection leads to adaptation to the environment. the others drive diversity. Part III Sources of Genetic Variation Part IV Lines of Evidence http://en.wikipedia.org/wiki/Evidenc e_of_common_descent Part V Controversies Controversies Short answer: there aren’t any. Scientists argue about the mechanism, not underlying idea. Of all Western Countries, Americans believe in evolution the least. The next country is Turkey. Why? It’s not just religion Politicizing Evolution In America, those who say they are pro-life, pro death penalty, and anti- gay marriage also do not believe in evolution. In Europe, there is no correlation between these and evolution belief. • Christian fundamentalists have made evolution a political issue. Less people in America believe in evolution today than they did a few years ago. The Exception of Evolution Why is evolution so contested even today, when the last century of work has done nothing but deepen our confirmations of the underlying principles? It is the one theory that directly impacts our personal identity. The very thing that makes us human causes strife. Evolution leads many to ponder the core questions of our very existence – where did we come from, where are we going, why are we here – what is our purpose? However, evolution theory does not address these questions, contrary to what popular belief states. Controversies • “Young Earth” Creationists • “Expelled” the movie – http://www.discovery.org/expelled/ – http://www.rottentomatoes.com/m/expelled_no_inte lligence_allowed/ • The Creationist MUSEUM – http://www.creationmuseum.org/ – http://www.discovery.org/ – http://friendlyatheist.com/2007/09/18/video-of-mycreation-museum-speech/