Chapter 7 Audition, the Body Senses, and the Chemical Senses This multimedia product and its contents are protected under copyright law. The following are prohibited by law: •any public performance or display, including transmission of any image over a network •preparation of any derivative work, including the extraction, in whole or in part, of any images •any rental, lease or lending of the program. 1 Copyright © 2008 Pearson Allyn & Bacon Inc. • Chapter 7 Outline • Audition • Vestibular System • Somatosenses • Gustation • Olfaction 2 Copyright © 2008 Pearson Allyn & Bacon Inc. • The Stimulus • Sound is produced by objects that ________and set molecules of air into __________. • Sound travels approximately 700 miles per hour. • Humans are sensitive to vibrations between ____and _________________ times per second. • Vibrations of air are perceived as sounds. 3 Copyright © 2008 Pearson Allyn & Bacon Inc. • The Stimulus • ___________ • A perceptual dimension of sound; corresponds to the _______________________of the stimulus. • _____________ • Cycles per second. • ________________ • A perceptual dimension of sound; corresponds to the ______________ of the stimulus. (__) • Timbre • A perceptual dimension of sound; corresponds to the _____________. 4 Copyright © 2008 Pearson Allyn & Bacon Inc. 5 Copyright © 2008 Pearson Allyn & Bacon Inc. 6 Copyright © 2008 Pearson Allyn & Bacon Inc. • Anatomy of the Ear • _______________membrane • The ___________; vibrates when stimulated by sound waves. • _____________ • The bones of the _____________. • The three ________________ in the body. 7 Copyright © 2008 Pearson Allyn & Bacon Inc. • Anatomy of the Ear • __________ • The first of the three ossicles; attached to the tympanic membrane. • __________ • Located between the malleus and the stapes. • _______________ • The third ossicle; attached to the ______________ of the cochlea. 8 Copyright © 2008 Pearson Allyn & Bacon Inc. • Anatomy of the ear • _______________ • The _________, fluid-filled, bony structure of the inner ear; contains the basilar membrane and the auditory receptor hair cells. • ____________window • An opening in the bone of the cochlea that reveals a membrane against which the baseplate of the stapes presses, transmitting sound vibrations into the fluid within the cochlea. • _____________window • An opening in the bone of the cochlea that permits vibrations to be transmitter, via the oval window, into the fluid of the cochlea. 9 Copyright © 2008 Pearson Allyn & Bacon Inc. • Anatomy of the ear • _______________ • The sensory organ on the basilar membrane that contains the auditory hair cell; considered the receptive organ of the auditory system. • _________________ • The sensory receptive cell of the auditory apparatus. • Deiter’s cell • A supporting cell found in the organ of Corti; sustains the auditory hair cells. 10 Copyright © 2008 Pearson Allyn & Bacon Inc. • Anatomy of the ear • _______________ membrane • A membrane in the cochlea of the inner ear; contains the organ of Corti. • ______________ membrane • A membrane located above the basilar membrane; serves as the shelf against which the cilia of the auditory hair cells move. 11 Copyright © 2008 Pearson Allyn & Bacon Inc. 12 Copyright © 2008 Pearson Allyn & Bacon Inc. 13 Copyright © 2008 Pearson Allyn & Bacon Inc. • Auditory Hair Cells and the Transduction of Auditory Information • ___________ • A _________ appendage of a cell involved in movement or transducing sensory information; found on the __________________________________________ __________________________________________. • Tip link • An elastic filament that attaches the tip of one cilium to the side of the adjacent cilium. • Insertional plaque • The point of attachment of a tip link to a cilium. • Site of origin of ___________________. 14 Copyright © 2008 Pearson Allyn & Bacon Inc. 15 Copyright © 2008 Pearson Allyn & Bacon Inc. 16 Copyright © 2008 Pearson Allyn & Bacon Inc. • The Auditory Pathway • Cochlear nerve • A branch of the ____________cranial nerve; the branch of the auditory nerve that transmits auditory information from the cochlea to the brain. • _____________________ • One of a group of nuclei in the medulla that receive auditory information from the cochlea. 17 Copyright © 2008 Pearson Allyn & Bacon Inc. • Shearing force (left-right bending of the cilia) • Opens and closes __________________. • Hair cells _________________________as the cilia bend back and forth. ______________ are receptors. Outer hair cells ___________________________________________ ___________________________________________ ___________________________________________ ___________________________________________ • • 18 Copyright © 2008 Pearson Allyn & Bacon Inc. • The Central Auditory System • ______________________ • A group of nuclei in the medulla; involved with auditory functions, including ____________ of the sound source. • Lateral lemniscus • A band of fibers running rostrally through the medulla and pons; carries fibers of the auditory system. Subthalamic auditory system is very complex. The ______________ processing is largely devoted to sound localization. 19 Copyright © 2008 Pearson Allyn & Bacon Inc. 20 Copyright © 2008 Pearson Allyn & Bacon Inc. • The Central Auditory System • _______________________ • A topographical organized mapping of different frequencies of sound that are represented in a particular region of the brain. 21 Copyright © 2008 Pearson Allyn & Bacon Inc. • Perception of Pitch • ________________ • The system by which information about different frequencies is coded by different locations on the basilar membrane. • ____________________ • An electrical device surgically implanted in the inner ear that can enable a deaf person to hear. • The implants simulate place coding and improve speech perception. 22 Copyright © 2008 Pearson Allyn & Bacon Inc. 23 Copyright © 2008 Pearson Allyn & Bacon Inc. 24 Copyright © 2008 Pearson Allyn & Bacon Inc. • Rate Coding • Rate code (________________) • The system by which information about different frequencies is coded by the rate of firing of neurons in the auditory system. 25 Copyright © 2008 Pearson Allyn & Bacon Inc. • Perception of Timbre • _____________________ • The _____________, and usually most intense, frequency of a complex sound; most often perceived as the sound’s basic __________. • Overtone • The frequency of complex tones that occurs at ______________ of the fundamental frequency. 26 Copyright © 2008 Pearson Allyn & Bacon Inc. • Perception Spatial Location • Humans can determine the location of a sound because auditory neurons respond selectively to _____________________________________________ ____________________________________________. • _____________ difference • The difference in arrival times of sound waves at each of the eardrums. 27 Copyright © 2008 Pearson Allyn & Bacon Inc. 28 Copyright © 2008 Pearson Allyn & Bacon Inc. 29 Copyright © 2008 Pearson Allyn & Bacon Inc. Elevation specific transfer functions produced by head and pinna geometry. 30 Copyright © 2008 Pearson Allyn & Bacon Inc. • Perception of Environmental Sounds • Transfer functions reveal the elevation of the signal. • ___________ • Loss or impairment of musical abilities, produced by hereditary factors or brain damage. Inability to recognize ___________, but the emotional quality of the music may be identified (happy/sad). 31 Copyright © 2008 Pearson Allyn & Bacon Inc. Blue = dorsal pathway, location. Orange = ventral pathway, pattern recognition . 32 Copyright © 2008 Pearson Allyn & Bacon Inc. Recognized (sound played forward), unrecognized (sound played backwards). 33 Copyright © 2008 Pearson Allyn & Bacon Inc. • Vestibular System • __________________ • One of a set of two receptor organs in each inner ear that detects changes in the ____ of the head (orientation). • ______________ • One of the three ring-like structures of the vestibular apparatus that detect changes in head rotation (angular _______________). • Match the three ____________ of the head: sagittal, transverse, & horizontal. 34 Copyright © 2008 Pearson Allyn & Bacon Inc. • Vestibular System • Anatomy of the vestibular apparatus • Utricle • One of the vestibular sacs • Saccule • One of the vestibular sacs. 35 Copyright © 2008 Pearson Allyn & Bacon Inc. • Vestibular System • Anatomy of the vestibular apparatus • ______________ • An enlargement in a semicircular canal; contains the cupula and crista. • _______________ • A ________________ mass found in the ampulla of the semicircular canals; moves in response to the flow of the fluid in the canals. 36 Copyright © 2008 Pearson Allyn & Bacon Inc. • Vestibular System • The receptor cells • Hair cells • Similar to the hair cells found in the cochlea; method of transduction is also similar to hair cells of the cochlea. • Vestibular ganglion • A nodule on the vestibular nerve that contains the cell bodies of the bipolar neurons that convey vestibular information to the brain. 37 Copyright © 2008 Pearson Allyn & Bacon Inc. 38 Copyright © 2008 Pearson Allyn & Bacon Inc. 39 Copyright © 2008 Pearson Allyn & Bacon Inc. • Somatosenses • The stimuli • The cutaneous senses respond to several different types of stimuli: pressure, vibration, heating, cooling, and events that cause tissue damage (and hence pain). • Some receptors report changes in _______________ to the brain; providing our sense of ______________. • Additional receptors provide information about the internal organs such as the linings of muscles and the gastrointestinal system. 40 Copyright © 2008 Pearson Allyn & Bacon Inc. • Somatosenses • Cutaneous sense • One of the somatosenses; includes sensitivity to stimuli that involve the skin. • Kinesthesia • Perception of the body’s ___________________. • Organic sense • A sense modality that arises from receptors located within the inner organs of the body. 41 Copyright © 2008 Pearson Allyn & Bacon Inc. • Somatosenses • Anatomy of the skin and its receptive organs • __________________ • Skin that does not contain ____; found on the palms and soles of the feet. • Ruffini corpuscle • Detects _____________ of the skin, located in hairy skin. • Pacinian corpuscles • A specialized, encapsulated somatosensory nerve ending that detects mechanical stimuli, especially _________________________. 42 Copyright © 2008 Pearson Allyn & Bacon Inc. •Somatosenses • Anatomy of the skin and its receptive organs • ____________________ •The touch-sensitive end organs located in the papillae, small elevations of the dermis that project up into the epidermis. Detect low frequency vibration or ________________. • ___________________ •The touch-sensitive end organs found at the base of the epidermis, adjacent to sweat ducts. Detect ___________________ of the skin. 43 Copyright © 2008 Pearson Allyn & Bacon Inc. 44 Copyright © 2008 Pearson Allyn & Bacon Inc. •Somatosenses •Perception of cutaneous stimulation •___________________ •Sensations that appear to originate in a limb that has been amputated. 45 Copyright © 2008 Pearson Allyn & Bacon Inc. •Somatosenses •Perception of cutaneous stimulation Touch •Fast conducting myelinated pathway •Small diameter unmyelinated pathway underlies ___________________________________________ ____________________________________ •Temperature (free nerve endings) •There are at least 6 classes of TRP family receptor sites that detect ____________________________. •Some receptors respond to warmth •Some receptors respond to coolness (menthal and mint bind with the TRPM8 receptor (transcient receptor potential) in the cutaneous fiber pathway. 46 Copyright © 2008 Pearson Allyn & Bacon Inc. •Somatosenses • The somatosensory pathways • Trigeminal nerve (5th cranial nerve) [Head & Face] • Dorsal columns fine touch • Medial lemniscus • Ventral posterior nuclei of the thalamus • Spinothalamic tract poor localization pain & temperature 47 Copyright © 2008 Pearson Allyn & Bacon Inc. • Pain • • Free nerve ending are the ________________. Three categories of receptors: • High threshold mechanoreceptors – respond to intense pressure. • Extreme heat and acids (Nerve endings contain TRPV1 receptors. • Free nerve fibers responsive to ATP – released from ______________ muscles, blood vessel blockage, and rapidly growing tumors. 48 Copyright © 2008 Pearson Allyn & Bacon Inc. • Pain Perception • Three components: (see Figure 7.19) • Sensory – intensity of the pain • Mediated by ventral posterior thalamic pathway • Emotional consequence – the level of unpleasantness. • Mediated by pathways reaching the insular cortex, and cingular cortex. (Reduced by ___________________ • Long-term implications of chronic pain • Mediated by pathways reaching the prefrontal cortex. Patients with prefrontal damage are _____________ with the implications of chronic pain for healthfullness 49 Copyright © 2008 Pearson Allyn & Bacon Inc. Increased activity produced by a placebo that reduced pain. 50 Copyright © 2008 Pearson Allyn & Bacon Inc. 51 Copyright © 2008 Pearson Allyn & Bacon Inc. • Gustation • The stimuli • Gustation is related to eating and is our first chemical sense. • This sense modality helps us to determine the nature of things we put in our mouths. • The tongue, palate, pharynx, and larynx contain about 10,000 sensory receptor taste buds. • There are only _____qualities of taste: __________________________________________ _________________________________________. 52 Copyright © 2008 Pearson Allyn & Bacon Inc. • Gustation • The stimuli • The are only five qualities of taste • • • • • Bitterness Sourness Sweetness Saltiness Umami • Umami • A Japanese word for “good taste, refers to the taste of monosodium glutamate (MSG). This receptor detects the presence of glutamate, an amino acid found in ___________. 53 Copyright © 2008 Pearson Allyn & Bacon Inc. • Most vertebrates detect all 5 taste qualities • Cats are unable to detect sweetness, it is usually not a part of their diet. Most omnivores like ripe fruit associated with sweetness.. • Saltiness – sodium chloride, necessary to maintain electrolytes. • Umami – a protein detector. • Sour – associated with acidity associated with bacterial activity. Causes an avoidance reaction. • Bitter – associated with poisonous plant alkaloids. 54 Copyright © 2008 Pearson Allyn & Bacon Inc. 55 Copyright © 2008 Pearson Allyn & Bacon Inc. • Gustation • Taste buds consist of groups of 20-50 receptors cells. • The receptor cells have a life span of 10 days. There is a high turn over rate for these receptors. • The gustatory pathway • Nucleus of the solitary tract • A nucleus of the medulla that receives information from visceral organs and from the gustatory system. 56 Copyright © 2008 Pearson Allyn & Bacon Inc. 57 Copyright © 2008 Pearson Allyn & Bacon Inc. 58 Copyright © 2008 Pearson Allyn & Bacon Inc. • Olfaction • The stimulus • Olfaction is the second chemical sense. Used to help us avoid _______________. • For humans, olfaction is the most enigmatic of the modalities. ___________________. • The stimulus for odor (known as odorants) consists of volatile substances having a molecular weight in the range of approximately 15 to 300. • Almost all odorous compounds are ______________ and of organic origin. 59 Copyright © 2008 Pearson Allyn & Bacon Inc. • Olfaction • Anatomy of the olfactory apparatus • Olfactory epithelium • The epithelial tissue of the nasal sinus that covers the _____________plate; contains the _____of the olfactory ____________. • Olfactory bulb • The protrusion at the end of the olfactory tract; receives information from the olfactory receptors. 60 Copyright © 2008 Pearson Allyn & Bacon Inc. 61 Copyright © 2008 Pearson Allyn & Bacon Inc. • Olfaction • Anatomy of the olfactory apparatus • Mitral cell • A neuron located in the olfactory bulb that receives information from olfactory receptors; axons of mitral cells bring information to the rest of the brain. • Olfactory glomerulus • A bundle of dendrites of mitral cells and associated terminal buttons of the axons of olfactory receptors. 62 Copyright © 2008 Pearson Allyn & Bacon Inc. • Olfaction • The sense of __________ is a blend of taste and olfaction, the orbitofrontal cortex receives gustatory and taste projections and this may be where flavor is perceived. • Humans are thought to possess _____different olfactory receptors, and the pattern of stimulation gives rise to over ________________ odors. • Odors are not easily put into words. 63 Copyright © 2008 Pearson Allyn & Bacon Inc. 64 Copyright © 2008 Pearson Allyn & Bacon Inc. Lock & Key Concept 65 Copyright © 2008 Pearson Allyn & Bacon Inc.