Lecture 12: Sensory Receptors and Special Senses I. General Terms A. B. C. D. E. F. G. H. sensation - awareness of external/internal conditions perception - conscious registration of conditions stimulus - change that can initiate nerve impulse receptor (sense organ) - converts stimulus to impulse transduction - changing stimulus signal into nerve signal generator potential - electrical impulse in receptor receptor potential - receptor releases neurotransmitter adaptation - decreased sensitivity with repeat stimuli a. rapidly adapting - pressure, touch, smell b. slowly adapting - pain, position, blood chemicals I. afterimage - sensation even after stimulus is gone J. modality - distinct property of each sensation II. General Classification of Receptors A. Location 1. exteroreceptors - respond to external environment 2. enteroreceptors - respond to internal environment 3. proprioreceptors - respond to body position/motion B. Type of Stimulus Detected 1. mechanoreceptors - any mechanical deviation a. touch, pressure, vibration, proprioception etc. 2. thermoreceptors - changes in temperature 3. nocireceptors - pain; physical or chemical damage 4. photoreceptors - light; rods & cones of the eye 5. chemoreceptors - shapes of different molecules a. taste, smell, chemicals of blood C. Simplicity or Complexity of the Receptor Structure 1. simple receptors - associated with general senses a. touch, pressure, temperature, vibration, pain 2. complex receptors - associated with SPECIAL senses a. smell, taste, sight, hearing, equilibrium III. Classification of General Senses A. Cutaneous (Skin) Senses 1. tactile (touch) a. light touch - location not perceived b. discriminative touch - location perceived c. Merkel's discs - discriminative touch d. Meissner's corpuscle - discriminative touch e. organs of Ruffini - deep, continuous touch 2. pressure a. felt over a large area than touch, deeper 1 b. Pacinian corpuscle - lower layer of dermis 3. vibration a. detect high and low frequency vibrations 4. thermosensation a. respond to hot/cold; may be free nerve endings 5. pain a. b. c. (nociception) acute pain - very quick, not felt in deep areas chronic pain - longer lasting, gradual increase somatic pain - skin, muscles, joints i. superficial -skin ii. deep - muscle, joint, tendon, fascia d. visceral pain - from receptors in organs e. referred pain - projected to skin above organ B. Proprioceptive (kinesthetic) Sense 1. function - position of limbs/body and equilibrium 2. muscle spindles a. intrafusal fibers - inner muscle fibers i. type Ia sensory fiber - in center ii. type II sensory fiber - at ends iii. gamma motor neurons - from ventral horn b. extrafusal fibers - outer muscle fibers i. alpha motor neurons - form ventral horn 3. Golgi (tendon) organs a. at junction of tendon and muscle 4. Joint Kinesthetic receptors a. within/around synovial joints IV. Classification of Special Senses A. Olfaction (smell) 1. 2. 3. 4. 5. olfactory olfactory olfactory olfactory olfactory cells - bipolar neurons in epithelium glands - secrete mucus to clean epithelium nerve (I) - axons of olfactory cells bulbs - brain region where (I) synapses tract - axons from bulbs to cortex B. Gustation (taste) 1. 2. 3. 4. 5. 6. gustatory cells - neuron with hairlike extension taste buds - location of gustatory cells facial nerve (VII) - anterior 2/3 of tongue glossopharyngeal nerve (IX) - posterior 1/3 vagus nerve (X) - throat and epiglottis -> medulla -> thalamus -> cortex C. Vision 1. Accessory Structures of the Eye 2 a. eyebrows b. eyelids (palpebrae) i. levator palpebrae superioris muscle ii. palpebral fissure iii. lateral commissure iv. medial commissure v. lacrimal caruncle (lacrimal gland) crying c. tarsal plate - inner wall of eyelid d. tarsal glands - secrete oil e. conjunctiva - mucous membrane of eyelid f. eyelashes g. lacrimal gland - for tear secretion i. lacrimal ducts ii. lacrimal puncta iii. lacrimal sac iv. nasolacrimal duct 2. The Structure of the Eyeball a. fibrous tunic - outer coat of the eyeball i. sclera - posterior portion ii. cornea - anterior portion b. vascular tunic (uvea) - middle layer i. choroid - posterior, pigment/vasculature ii. ciliary body - muscle shapes lens iii. iris - colored part, with pupil (hole) c. nervous tunic (retina) - posterior surface i. photoreceptors (rods & cones) ii. bipolar cells iii. ganglion cells d. lens - just behind pupil and iris 3. Pathway of Light to the Brain a. b. c. d. photoreceptors pick up the light ganglion cells converge signals -> optic nerve optic nerve -> lateral geniculate of thalamus lateral geniculate -> occipital cortex D. Hearing & Equilibrium 1. external ear a. b. c. d. e. f. pinna (auricle) - Ross Perot helix - rim of the pinna lobule - your mate's favorite part external auditory canal ceruminous glands - love that earwax! tympanic membrane (eardrum) 2. middle ear a. tympanic antrum - chamber to air cells b. auditory (Eustachian) tube - to nasopharynx c. auditory ossicles - bones of middle ear i. malleus - attached to tympanic membrane 3 ii. incus - intermediate bone iii. stapes - stirrup d. tensor tympani muscles - to malleus (protect) e. stapedius muscle - to stapes (protect) f. oval and round windows - to inner ear 3. inner ear (labyrinth) a. bony labyrinth - has fluid called perilymph i. vestibule ii. cochlea iii. semicircular canals b. membranous labyrinth - has endolymph c. vestibule - oval central portion body labyr. i. utricle & saccule - two sacs d. cochlea - sound organ, sounds are sensed here e. semicircular canals - equilibrium in 3-D 4. Neural Pathway for Sound/Equilibrium Sensation a. cochlea/vestibular -> vestibulocochlear (VIII) 4