EARTH AND ENVIRONMENT THROUGH TIME LABORATORY - EES 1005 LABORATORY SEVEN PALEOZOIC MARINE FOSSILS Introduction Very few fossils have been found that date before the Paleozoic. During the PreCambrian there were no skeletal animals. Nearly all of the fossils that have been found from the PreCambrian are trace fossils, and a few soft body impressions. One of the most important fossils from the PreCambrian is called a stromatolite. Stromatolites are trace fossils of a type of blue-green aglage. This algae secrets a mucus around its strands, and in clam water sediments can fall into the mucus and become stuck. As the algae grows on top, more and more sediments become bound to the mass, forming a cabbage-like mound. Blue-green algae, which produces oxygen (O2) as a waste product during photosynthesis, was one of the first organisms to evolve, approximately 3,500 million years ago, and had a few predators until the Cambrian, approximately 570 million years ago. During that interim, enough free oxygen was produced and accumulated in the atmosphere, that new life forms were able to evolve. Stromatolites become increasingly rare throughout the Phanerozoic, because organisms evolved which consume the mucus on the algae before the sediments accumulate. The beginning of the Paleozoic marks the first appearance of organisms with hard pars. This lab will introduce some of the important fossils in the geological fossil record. You will be expected to identify the italicized fossils for the next quiz, and know the phylum for each. Some of these are index fossils which are important tools. Index fossils are fossils which are very useful in correlating and dating rock beds. In order for a fossil to be a good index fossil, it must be: 1) geologically short lived, 2) abundant during its existence, and 3) widespread in its occurrence. Paleozoic Fossils This week we begin working on the Paleozoic fossil shelf for your notebook. Remember to include the age ranges (pgs. 32-33) when requested. PHYLUM ARTHROPODA Cl. Trilobita (Cambrian - Permian) Trilobites had a hard segmented body with an outer shell made of chitin, which preserves fairly easily. As the trilobites grew, they shed this exoskeleton and formed a new one. Their body was divided into three lobes: cephalon (head), thorax (body), and pygidium (tail). Their fossil remains often consist of disarticulated heads, tails, or body segments. They can also be found rolled up into a ball. Trilobites were most abundant during the Cambrian and Ordovician. PHYLUM BRACHIOPODA G. Brachiopods (Cambrian - Recent, common in Paleozoic) Brachiopods were extremely common in the Paleozoic. They are easily mistaken for bivalves because they are enclosed by two hinged valves, but their soft parts are fundamentally different. They show bilateral symmetry like the bivalves, however they symmetry is perpendicular to the plane of separation, whereas in bivalves the plane of symmetry is the same plane as the plane of separation. In most species, one valve has an opening at its apex, through which a stalk-like organ protrudes, anchoring the animal to stable surfaces. Articulate brachiopods have a mechanical "tooth & socket" hinge which can keep the shells together even after they die. Inarticulate brachiopods are held together by muscle, so the shells usually fall apart from each other when they die. PHYLUM BRYOZOA G. "Branching" Bryozoans (Ordovician Recent) Bryozoans are microscopic organisms which live in colonies. They look like corals, however they have a complex anatomy. A colony consists of individual zooids which are asexually budded clones of the original. The branching bryozoans were common in the Paleozoic. The Archimedes bryozoan was a particular genus common in the Mississippian, and makes an excellent index fossil: it was abundant, widespread, and had a very recognizable shape. It had a lacy colony attached to a spiral-shaped support structure. Paleozoic Fossils PHYLUM CNIDARIA Corals are simple animals with no organs. They have just a mouth surrounded by tentacles, a gut cavity, simple muscles, and a nerve network. Specialized stinging cells called cnematocysts aid in capturing prey, defense, and aggression. True corals have aragonite or calcite skeletons and are also commonly fossilized. Jellyfish are also in this phylum, but are rarely fossilized. G. Rugose Corals (Ordovician - Permian) Rugose corals contained a cup-like depression in which the polyp lived. Its gut cavity was radially divided by septa.The septa can be seen by looking at the end of the fossilized skeleton. Rugose corals are found as "horn corals" - the remains of a solitary organism, or as a group of individuals cemented together forming a colony. G. Tabulate Corals (Ordovician Permian) Tabulate corals were similar to rugose corals, but did not have divided chambers and were exclusively colonial. PHYLUM ECHINODERMATA Echinoderms are easily recognized by their unique five-fold symmetry. They feature a body cavity typically enclosed by a calyx or similar structure of plates which cleave like calcite. All echinoderms have a circulatory system in which the fluid salt content is balanced with that of sea water by means of suckers on their tube feet. Cl. Crinoidea (Ordovician - Recent, common in Mississippian) Crinoids live on the shallow ocean bottom attached by a stem and superficially resemble sea lilies. A calyx of five-fold symmetry encloses their globular body. Their food gathering arms have tube feet and resemble feathers. The arms and stems consist of disc-like plates called columnals. Crinoids were so so common during the Mississippian that some limestones of this period are composed almost entirely of broken up crinoid stems. These limestones get their own name: encrinite. Cl. Blastoidea (Ordovician - Permian, very common in the Mississippian) Blastoids looked similar to crinoids except the plates on their calyx were somewhat flatter and had a more distinct five-fold pattern. Their arms were located along the margins of these plates. They were also extremely common during the Mississippian. G. Starfish (Ordovician - Recent) Starfish are composed of soft body parts surrounded by smaller spiny plates which break apart upon their death. They feed using their tube feet. Most starfish have five arms. Paleozoic Fossils PHYLUM HEMICHORDATA Cl. Graptolita (Cambrian - Mississippian, most common in the Ordovician) Graptolites were small colonial organisms that floated in the ocean. They are typically preserved by carbonization, appearing as black saw-tooth, pencil-like lines. They are good index fossils for the Ordovician and Silurian. PHYLUM MOLLUSCA Cl. Bivalvia (Ordovician - Recent) Bivalves have two shells that are mirror images of each other. They have a muscular foot that may be used for movement. Clams and oysters are bivalves. Cl. Gastropoda (Cambrian - Recent) Gastropods are the most widespread, diverse, and abundant molluscan class. Snails and slugs are in this class. Shelled gastropods have a clockwise rotation or twisting of an otherwise bilaterally symmetrical body. Their shells can be coiled (planar or helical) but there are no septa. Cl. Cephalopoda (Cambrian - Recent) Cephalopods are predatory invertebrates with bilateral symmetry. Shelled cephalopods live in the outer portion of a shell which is divided into chambers by a series of septa. The contact between the septa and the shell interior is called the suture. The unoccupied chambers are filled with gas and liquid. These chambers are connected by a tissuelined tube (siphuncle) and cephalopods can control their position in the water by regulating the distribution of the liquid. G. Nautiloid (Cambrian - Recent) Nautiloids have a straight, simple suture pattern on the outside of their shell, marking where the inside is divided into chambers. They may be straight (conar) or coiled (planar). G. Ammonoid (Devonian - Cretaceous) Ammonoids evolved from Nautiloids, developing a complex suture pattern that gave more support to the shell, allowing them to feed in deeper waters. They are usually coiled, but may be straight (cone-shaped). Paleozoic Fossils PHYLUM PORIFERA G. Stromatoporoid (Cambrian - Recent, most common in the Ordovician and Devonian) Stromatoporoids were a colonial group of simple multicellular animals that secreted a calcareous laminated skeleton. They were most abundant during the Ordovician and Devonian. These sponges had a hard skeleton and were reef formers. PHYLUM PROTOZOA Cl. Foraminifera (Ordovician - Recent) Forams are microscopic single celled organisms made up of calcium carbonate. G. Fusilinids (Pennsylvanian - Permian) These are the most common Paleozoic forams, and make good index fossils. Cl. Radiolaria (PreCambrian - Recent) Radiolarians are microscopic, single celled, free floating organisms which drop plates of silica (SiO2) into the ocean, forming oceanic deposits of chert.