Origin and Evolution of Vertebrates The clades to which the Dinosauria belong, and the shared, derived characters associated with each are listed below. 1. Chordata bilateral symmetry, segmented muscles, notocords & nerve cord, gill slits fossil chordates are known from the Cambrian (520 Ma) 2. Vertebrata Big head with brain, bone around head, more developed senses 3. Gnathostoma true jaws 4. Tetrapoda Four weight-bearing limbs, neck, loss of gills, lungs 5. Amniota Amniotic egg, thick skin, distinctive skulls Skulls: anapsid, synapsid, and diapsid patterns (euryapsid = derived diapsid) Holes also called fenestrae Why holes? To lighten skulls and provide muscle attachment. 6. Synapsida One opening in skull; lower: anapsid, synapsid, and diapsid patterns 7. Reptilia Distinctive neck vertebrae and skull roof and braincase bone arrangement. 8. Diapsida Two openings behind the eye, in the temporal region 9. Archosauria Teeth in sockets, antorbital and mandibular fenestrae, rotating ankles 10. Ornithodira Erect stance Advanced mesotarsal ankle: astragalus much larger than calcaneum; both rigidly attached to one another and to the tibia; AM ankle has single hinge between the ankle and the rest of the foot; allows little twisting of the ankle when walking; swings backwards and forwards only; enables speedy movement 11. Pterosauria Elongate fourth finger (for wing support) Many other flight specializations 12. Dinosauria 3+ sacral vertebrae Perforate acetabulum Asymmetric hand Etc..