Classification AP BIOLOGY 11 13. The Protists • The protists are by far the most difficult group of organisms to classify. • It is the most artificial and disagreed upon by biologists. • We will not focus so much on the taxonomy as the diversity within this group. • We will divide them into three groups based upon their nutrition method. Classification 1. Ingestive Protists • These are animal like protists that bring food into cells (ingest) and digest it. • Examples include the Protozoa. A Paramecium Classification 3. Photosynthetic Protists • Plant like protists that do photosynthesis. • Include most of the unicellular, colonial, and multicellular algae. Classification 2. Absorptive Protists • Fungus like protists that use extracellular digestion, ie. They secrete enzymes onto a food source, digest it, then absorb the digested products. • Ex: Water and slime molds. A plasmodial slime mold Evolution • Protists represent the first eukaryotes. • Probably evolved from prokaryotes that didn’t have a cell wall (therefore flexible). • Maintained shape with their cytoskeleton. • This could also “pull” is some plasma membrane to surround their nuclear material forming a nucleus (by process like phagocytosis). 1 Evolution • This started cytoplasmic compartmentalization which allows for internal specialization. • Cytoskeleton also allowed for new mode of cell division - mitosis. • This removed the restriction of one circular chromosome, allowing many linear chromosomes - this produces more genetic information and more complexity. Ingestive Protists 1. Archeozoa • “ancient protists” • Lack mitochondria (primitive feature), often flagellated. • Ex. Giardia sp. (Beaver Fever) • This also allowed for true sexual reproduction to occur with meiosis and fertilization - increasing variations! • The evolution of mitochondria and chloroplasts probably involved the evolution of a symbiotic relationship between a prokaryote and the evolving eukaryote. • Protist evolution is still quite obscure. Ingestive Protists 2. The Flagellates • Move by flagella. • Contain both photosynthetic and heterotrophic varieties. • Also contains parasitic forms. • Three main subgroups: Ingestive Protists A. Dinoflagellates • Photosynthetic. • Major component of phytoplankton in the ocean. • Cause “Red Tides” which can cause paralytic poisoning when eating shellfish. Evolution Ingestive Protists B. Sporozoans • Form spores at one stage of life cycle, other stage moves by flagella. • Ex. Plasmodium vivax parasite that causes malaria. • Learn its life cycle and vector. P. vivax in human blood smear 2 Ingestive Protists C. Euglenophyta • Contains Euglena and related organisms. • Combine auto and heterotroph character. • Shows why protists are so difficult to classify. • Moves by flagellum. Ingestive Protists 3. Ciliates • Among most complex of protozoans - show high degree of intracellular specialization. • Have body “systems” like animals. • Reproduce sexually by conjugation. Paramecia Ingestive Protists Ingestive Protists Ingestive Protists Ingestive Protists 4. Rhizopoids • Move by pseudopods with cytoplasmic streaming demonstrates “sol-gel” phase reversal. • Includes largest eukaryotic cells - the amoeba. 5. Radiolaria • Secrete a “glass” skeleton. • Major component of fresh and salt water plankton. • Basis of aquatic food chains. 6. Forminifera • Have CaCO3 skeleton (chalk) form large deposits on ocean floors. • Mostly marine, have pseudopods. • Major source of chalk (white Cliffs of Dover). 3 Absorptive Protists Slime Molds • Fungus like protist that also has stage that resembles ameba. • Most common stage resembles a mass of gooey fungus called the “Plasmodium”. Absorptive Protists • The plasmodium forms an upright sporangium that produces and releases haploid spores. • These germinate to form the haploid ameba like and flagellated stages. Absorptive Protists • These synapse to form a zygote which grows back to the plasmodium. • The zygotes all merge together to form the plasmodium which is the feeding stage. • The asexual (spore) stage is environmentally resistant to survive winters. Plant Like Protists: Algae A) The Diatoms • Classified as “Golden - Brown Algae”. • Produce glass-like (silica based) cell walls in “box and lid”. • Compose a large part of fresh and salt water plankton. Plant Like Protists: Algae • • • • Most diverse group. Not all agree should be classified as protists All are photosynthetic. May be unicellular, colonial, or multicellular (some classify these in the plants). Plant Like Protists: Algae B) Brown Algae • Phylum Phaeophyta. • Contains brown pigment “fucoxanthin” as accessory light pigment (masks chlorophyll color). • Mostly marine and temperate. Kelp 4 Plant Like Protists: Algae • Kelp such as Macrocystis sp. shows basic adaptation to living near land and was necessary precursor to evolution of plants to land. • Show specialized regions: blades for photosynthesis and reproduction, stipe to hold and suspend blades, air bladders to float plant, and... Plant Like Protists: Algae C) Red Algae • Phylum Rhodophyta. • Red due to red accessory pigment (phycoerythricin). • Most common in warmer tropical waters. • In temperate zones, found in deeper water. Plant Like Protists: Algae 1. Unicellular Green Algae • Very common in fresh water as part of phytoplankton. • Ex. Chlamydomonas sp. Plant Like Protists: Algae • Holdfasts to enable plant to stay anchored in nutrient rich continental shelf area. • Kelp beds provide foundation of a major marine ecosystem. • Show “Alternation of Generations” Holdfast of Macrocytis sp. From Bamfield BC Plant Like Protists: Algae D) Green Algae • Phylum Chlorophyta. • Very diverse group. • Include unicellular, colonial, and multicellular forms. • Live in all environments: fresh and salt water, soil. Ulva sp. Sea lettuce Plant Like Protists: Algae 2. Colonial Green Algae • Live in long filaments or as colonial spheres. Filamentous green algae Volvox, spherical colonial 5 Plant Like Protists: Algae 3. Multicellular Green Algae • Mostly marine. • Likely ancestors of land plants. • Show equal Alternation of Generations (a pre-adaption to land) with equal Sporophyte and Gametophyte. • (end show) 6