A provisional checklist of Berkshire Coleoptera Dr.Jonty Denton 2013 1 REFERENCES Aalto, M.M. Coope, G.R., & Gibbard, P.L. 1984. Late Devensian river deposits beneath the floodplain terrace of the River Thames at Abingdon, Berkshire, England. Proceedings of the Geologists’ Association, 95: 65-79. Alexander, K.N.A., 2003. Is Malthodes brevicollis (Paykull) a British Beetle? Coleopterist,12: 35-39. Balfour-Browne, F. 1950. British water beetles. Vol. 2. London. Ray Society. Balfour-Browne, F. 1952. The aquatic Coleoptera of South Hampshire with some remarks on the composition of the fauna and its origin. Entomologist’s Gazette. 3 127-160. Balfour-Browne, F., 1958. British water beetles. Vol. 3. London. Ray Society. Barclay, M.V.L. 2003. Otiorhynchus (s. str.) armadillo (Rossi, 1792) and Otiorhynchus (s. str.) salicicola Heyden, 1908 (Curculionidae: Entiminae: Otiorhynchini), two southern European vine weevils established in the UK. Coleopterist, 12: 41-54 Bowestead, S. 1999. A revision of the Corylophidae (Coleoptera) of the West Palaearctic region. Geneva: Museum d'histoire Naturelle. pp. 203. Campbell, J.M. 19 . An atlas of Oxfordshire Coleoptera Chrysomelidae. Oxfordshire County Council Occasional paper No.13. Donisthorpe, H.St.J.K. 1927. The guests of British ants their habits and life histories. London, Routledge. Fowler, W.W., 1887.. The Coleoptera of The British Islands Vol. 1. London, Reeve. Fowler, W.W., 1888. The Coleoptera of The British Islands Vol. 2. London, Reeve. Fowler, W.W., 1889. The Coleoptera of The British Islands Vol. 3. London, Reeve. Fowler, W.W., 1890. The Coleoptera of The British Islands Vol. 4. London, Reeve. Fowler, W.W., 1891. The Coleoptera of The British Islands Vol. 5. London, Reeve. Fowler, W.W. & Donisthorpe, H.St.J.K. 1913.. The Coleoptera of The British Islands Vol. 6. London, Reeve. Hammond, P.M. 2002. UK tachyine beetls (Coleoptera: Carabidae) with special reference to the status of Tachys edmonsi Moore. English nature Research Reports. Number 465. English Nature, Peterborough. Hawkins, R D., 2000. Ladybirds of Surrey. Surrey Wildlife Trust, Pirbright. Hyman, P.S & Parsons, M.S., 1992. A review of the scarce and threatened Coleoptera of Great Britain. Part 1. JNCC, Peterborough Hyman, P.S & Parsons, M.S., 1994. A review of the scarce and threatened Coleoptera of Great Britain. Part 2. JNCC, Peterborough. Johnson, C. 1993. Provisional atlas of the Cryptophagidae-Atomariinae (Coleoptera) of Britain and Ireland. Huntingdon. Biological Records Centre. Johnson, C., 1997. Clambus simsoni Blackburn (Col., Clambidae) new to Britain, with notes on its wider distribution. Entomologist’s Monthly Magazine, 133: 196-99 Kirk-Spriggs, A.H., 1996. Handbooks for the Identification of British Insects Vol. 5. part 6a: Pollen Beetles. Coleoptera: Kateretidae and Nitidulidae: Meligethinae. Royal Entomological Society, London. 2 Levy, B. 2002. A revision of the British Species of Mordellistena (Col., Mordellidae) belonging to the parvula group and the subgenus Pseudomordellina Ermisch. British Journal of Entomology and Natural History, 15: 83-91. Luff, M.L., 1998. Provisional atlas of the ground beetles (Coleoptera, Carabidae) of Britain. Huntingdon. Biological Records Centre. Osborne, P. J. 1980. The insect fauna of the organic deposits at Sugworth and. its environmental and stratigraphic implications. Philosophical Transactions of the Royal Society of London, B289, 119-133. Robinson, M..A. 1979. The biological evidence. In G.Lambrick, & M.A.Robinson, Iron Age and Roman Riverside Settlements at Farmoor, Oxfordshire, 77-133. Council for British Archaelogy Research Report, 32, London. Robinson, M.A. 1981. The Iron Age to Early Saxon Environment of the Upper Thames Terraces. In M.Jones & G.Dimbleby (eds.) The Ewnvironment of man: the Iron Age to the Anglo-Saxon period, 251-286. British Archaeological Reports 87, Oxford. Robinson, M.A., Dickson, J.H. & Grieg, J.R.A. 1984.Waterlogged plant and invertebrate evidence. In D. Miles (ed.) Archaeology at Barton Court Farm, Abingdon, Oxon., microfiche chapters VIII & IX. Council for British Archaeology Research Report 50. Walker, J.J. 1906a. Preliminary list of Coleoptera observed in the neighbourhood of Oxford from 1819-1907. Proceedings of the Ashmolean Natural History Society. Walker, J.J. 1906b. First supplement to the Preliminary list of Coleoptera observed in the neighbourhood of Oxford from 1819-1907. Proceedings of the Ashmolean Natural History Society. Whitton, P. & Campbell, J.M. 1991. An atlas of Oxfordshire Coleoptera Carabidae. Oxfordshire County Council Occasional paper No.15. 3 CHECKLIST OF BERKSHIRE COLEOPTERA Carabidae (Ground Beetles) 247 species + 1 sub-fossil Cicindela campestris Linnaeus, 1758 Green tiger beetle Cicindela sylvatica Linneaus, 1758 Heath tiger beetle Cychrus caraboides (Linneaus, 1758) Carabus arvensis Herbst, 1784 Carabus auratus Linnaeus, 1761 SUB VCH 1907- Notes 2005 + ++ Local on heaths, scattered in woodlands and disturbed ground. * (Na) . On open dry heathland with short heather & open sandy areas. ++ ? + Widespread in wooded districts. A flightless snail predator. Very local on heathland and commons. Introduced. Well established in allotmenst in Reading (MVB etc) ++ ++ ++ ++ ++ Local in damp meadows, (Nb) Rare, declining? 1990 Widespread in open habitats. Local in dry areas especially on heaths and commons. Widespread and common occurring in woods, parks, gardens, and more open habitats. * ++ ++ ++ ++ (Na) Local in wooded country and heaths, Local in leaf litter etc. in woods, hedgerows, and disturbed ruderal sites. Local in damp shady places in leaf litter. 20th century colonist (first noted in the 1940s), now widespread especially in woods in litter under trees especially beech. Frequent in woods, gardens etc. Widespread and common. Local on heathland and open sandy ground.. Local in dry habitats especially heathland. Hitchcopse Frequent in all kinds of open situations including gardens. Rare in open dry areas. Frequent in leaf litter in woods etc. (Nb) Rare in open sites esp. on clay. Local in leaf litter in moist, often shady places, including gardens. Frequent in open dry country including gardens. Local at margins of water and in marshy ground. Local at margins of water and in marshy ground. (Nb) ? Extinct? Frequent in damp litter especially in woods, also in large gardens. Local on open bare ground in sandpits, heaths etc. Local on moist open ground, especially on heaths where it often shelters under mats of Zygogonium algae on bare sand. The Coppersmith Carabus granulatus Linneaus, 1758 Carabus monilis Fabricius, 1792 Carabus nemoralis Müller, 1764 Carabus problematicus Herbst, 1786 Carabus violaceus Linneaus, 1758 Violet ground beetle Calosoma inquisitor (Linneaus, 1758) Leistus ferrugineus (Linneaus, 1758) Leistus fulvibarbis Dejean, 1826 Leistus rufescens (Fabricius, 1775) Leistus rufomarginatus (Duftschmid, 1812) Leistus spinibarbis (Fabricius, 1775) Nebria brevicollis (Fabricius, 1792) Nebria salina Fairmaire & Laboulbène, 1854 Notiophilus aquaticus (Linneaus, 1758) Notiophilus biguttatus (Fabricius, 1779) Notiophilus germinyi Fauvel, 1863 Notiophilus palustris (Duftschmid, 1812) Notiophilus quadripunctatus Dejean, 1826 Notiophilus rufipes Curtis, 1829 Notiophilus substriatus Waterhouse, 1833 Elaphrus cupreus Duftschmid, 1812 Elaphrus riparius (Linneaus, 1758) Elaphrus uliginosus Fabricius, 1792 Loricera pilicornis (Fabricius, 1775) Dyschirius aeneus (Dejean, 1825) Dyschirius globosus (Herbst, 1784) + Rom + + Rom Rom Rom + + ++ ++ ++ ++ ++ + ++ + + ++ ++ ++ * ++ ++ ++ Leud ?? 4 Carabidae (Ground Beetles) 247 species + 1 sub-fossil Dyschirius politus (Dejean, 1825) Clivina collaris (Herbst, 1784) Clivina fossor (Linneaus, 1758) Broscus cephalotes (Linnaeus, 1758) Patrobus atrorufus (Ström, 1768) Trechus obtusus Erichson, 1837 Trehus quadristriatus (Schrank, 1781) Trechus rubens (Fabricius, 1792) Trechus secalis (Paykull, 1790) Lasiotrechus (=Trechus) discus (Fabricius, 1792) Trechoblemus (=Trechus) micros (Herbst, 1784) Asaphidion curtum (Heyden, 1870) Asaphidion flavipes sens str.( Linneaus, 1761) Asaphidion pallipes (Duftschmid, 1812) Asaphidion stierlini (Heyden, 1880) Bembidion nigricorne Gyllenhal, 1827 Bembidion lampros (Herbst, 1784) Bembidion properans Stephens, 1828 Bembidion punctulatum Drapiez, 1821 Bembidion dentellum (Thunberg, 1787) Bembidion obliquum Sturm, 1825 Bembidion varium (Olivier, 1795) Bembidion decorum (Zenker in Panzer, 1800) Bembidion femoratum Sturm, 1825 Bembidion maritimum Stephens, 1835 Bembidion nitidulum (Marsham, 1802) Bembidion stephensi Crotch, 1871 Bembidion testaceum (Duftschmid, 1812) Bembidion tetracolum Say, 1823 Bembidion quadrimaculatum (Linneaus, 1761) Bembidion quadripustulatum Serville, 1821 Bembidion gilvipes Sturm, 1825 Bembidion assimile Gyllenhal, 1810 Bembidion clarki Dawson, 1849 Bembidion minimum (Fabricius, 1792) Bembidion genei Küester, 1847 Bembidion doris (Panzer, 1796) Bembidion articulatum (Panzer, 1795) Bembidion obtusum Serville, 1821 Bembidion aeneum Germar, 1824 Bembidion biguttatum (Fabricius, 1779) Bembidion guttula (Fabricius, 1792) SUB VCH 19072005 + Rom * Rom ++ ++ ++ Rom ++ Rom ++ * + + Rom + + + * ++ + Rom ++ Rom ++ + Br/Sax ++ + + + ++ ? + Rom Br/Sax Br/Sax Rom Br/Sax Rom Rom + + + + ++ ++ ? + + ++ + ++ ++ + Notes Rare on bare sand near water. Rare, partly subterranean in gardens, and especially on sandy banks of rivers s Local, partly subterranean in wetlands, sandy river banks etc. HitchCopse Pit (DJM) Rare Wet woods, Local in open habitats including gardens, and cultivated land. Frequent in open country including gardens. (Nb) ?, Rare in wet grassland. (Nb) Rare in runs of mammals. [Wetland margins] Rare? [In mammal runs or burrows] 1988 Local on bare usually clayey soils in woods, rides and gardens. Rare in a variety of habitats primarily in vicinity of water. M.Moorcroft1994 doubtful Local. Recently split species, widespread especially on chalky and clay soils, and also in ruderal sites. (Nb) Rare. In heather litter on open heathland. Widespread on various open habitats including fields. Local in open habitats. ? [At margins of running water]. Widespread in marshy places near ponds, also on bare drawdown zones of drying water bodies. (Nb) Rare in marshy places near ponds and rivers on exposed silt. Rare on margins of ponds and marshy places. . Local in open sandy places, including sandpits and disturbed areas on heathlands. Extinct? [Exposed sediments in estuaries, sea shores] Rare near water [on exposed sediments]. , Rare [Shady bare ground]. (Nb) [At margins of rivers]. Widespread on bare ground near water, often on open shingle, and sand by rivers and streams. Local in open sparsely vegetated areas on a variety of soils. (Nb) Rare at margins of water on bare silt and sand. (Nb) Rare in marshes. Rare in marshes, litter at margins of ponds.. (Nb) Scarce. Shaded wetland/carr.. Mainly coastal, a recent colonist inland, on bare margins of standing water. Local on open barish ground in pits and quarries. Rare in marshes, wet woodland. Local on damp bare ground near water. Local on open dry, often cultivated land, including arable land. Local Local in marshland. Local in wetlands, margins of rivers and large ponds, lakes, also in waterlogged pasture and arable land. 5 Carabidae (Ground Beetles) SUB VCH 1907- Notes 247 species + 1 sub-fossil 2005 Bembidion lunatum (Duftschmid, 1812) *? (Nb) ? [On banks of rivers and streams]. Bembidion lunulatum (Fourcroy, 1785) Rom ++ Rare in wetlands. Bembidion mannerheimi Sahlberg, 1827 + Local in woodlands especially in damp litter and in bogs, and river margins. Ocys (Bembidion) harpaloides Serville, 1821 + Widespread in woods on trees, under bark and in suspended debris including old nests. Ocys (Bembidion) quinquestriatum Gyllenhal, 1810 + Rare? On old walls especially under ivy (Hedera helix). Kingston 1980 (DAP),, Horley 2001 (RDH). Tachys bistriatus (Duftschmid, 1812) + + (Nb) Rare on open ground at margins of water. Windsor Great Park (PMH), Elaphropus (=Tachys) parvulus (Dejean, 1831) + (Nb) Rare ? Increasing? Open ground including gardens, Stomis pumicatus (Panzer, 1795) + Local in grasslands, gardens [on rich soils?]. (Nb) Rare on recently burnt areas on heath/pinewoods. Pterostichus angustatus (Duftschmid, 1812) Pterostichus anthracinus (Panzer, 1795) ++ (Nb) Rare, in carr woodland. Pterostichus cupreus (Linneaus, 1758) Rom ++ Widespread in open habitats including ruderal areas. Pterostichus diligens (Sturm, 1824) ++ Local. Most frequent in wet heath/boggy areas, and other wetlands. (Nb) Extinct? [Marsh and carr] r Pterostichus gracilis (Dejean, 1828) Rom * (RDB1) Extinct? Last recorded at in 1913. [Heathland] Pterostichus kugelanni (Panzer, 1797) (Nb) Rare pre 1935 recs Pterostichus lepidus (Leske, 1785) * (Nb) 1970s Pterostichus longicollis (Duftschmid, 1812) * Pterostichus macer (Marsham, 1802) + Rare, though partly subterranean. Pterostichus madidus (Fabricius, 1775) The ++ Ubiquitous in open habitats including gardens, one of the commonest beetles in Black-clock Sax Pterostichus melanarius (Illiger, 1798) ++ Local in a wide variety of damp habitats from woodland, marshes, bogs etc. Pterostichus minor (Gyllenhal, 1827) Rom ++ Local in marsh and carr. Pterostichus niger (Schaller, 1783) Rom ++ Widespread in parkland and other partly wooded habitats. Pterostichus nigrita (Paykull, 1790) ++ Local in marsh and wet woodlands. Pterostichus oblongopunctatus (fabricius, 1787) ESax ++ (Nb) Pterostichus rhaeticus Heer, 1837 ++ Local in wetlands especially carr woodland and wooded heaths. Pterostichus strenuus (Panzer, 1796) ++ Local in woods and shaded places. Pterostichus vernalis (Panzer, 1795) ++ Local in open habitats on heaths and in marshes. Pterostichus versicolor (Sturm, 1824) ++ Widespread in open habitats including ruderal areas. Abax parallelepipedus (Piller & Mitterpacher 1783) Rom ++ Widespread and common especially in wooded areas, under logs etc. Calathus ambiguus (Paykull, 1790) ++ (Nb) Rare in open dry sandy and chalky soils. Frilford etc Calathus cinctus Motschulsky, 1850 ++ Rare in open sandy places Calathus erratus (Sahlberg, 1827) ++ Local on dry sandy ground especially on heaths. Calathus fuscipes (Goeze, 1777) Rom ++ Widespread and common in open areas including farmland and gardens. Calathus melanocephalus s. str. (Linneaus, 1758) Rom ++ Widespread in dry areas including ruderal sites. Calathus rotundicollis Dejean, 1828 ++ Widespread especially in wooded areas, under logs etc. Also occurs in gardens. =piceus (Marsham, 1802) Sphodrus leucophthalmus (Linneaus, 1758) * Extinct? [Synanthropic in stables, cellars etc. declining]. Laemostenus terricola (Herbst, 1784) ++ Local. Most records come from near rabbit warrens, also occurs under bark. Platyderus ruficollis (Marsham, 1802) ++ (Nb) Local in gardens and waste ground. R/ESax Synuchus vivalis (Illiger, 1798) = nivalis (Panzer, ++ Local in open dry areas including disturbed sites. 1797) Rom Olisthopus rotundatus (Paykull, 1790) ++ Widespread in open habitats including ruderal areas. 6 Carabidae (Ground Beetles) 247 species + 1 sub-fossil Sericoda (=Agonum) quadripunctata (Degeer, 1774) Anchomenus (=Agonum) dorsale (Pontoppidan, 1763) Paranchus (=Agonum) albipes (Fabricius, 1796) Oxypselaphus (Agonum) obscurum (Herbst, 1784) Agonum fuliginosum (Panzer, 1809) Agonum gracile Sturm, 1824 Agonum livens (Gyllenhal, 1810) Agonum marginatum (Linneaus, 1758) Agonum micans Nicolai, 1822 Agonum moestum (Duftschmid, 1812) Agonum muelleri (Herbst, 1784) Agonum nigrum Dejean, 1828 Agonum piceum (Linneaus, 1758) Agonum scitulum Dejean, 1828 Agonum sexpunctatum (Linneaus, 1758) Agonum thoreyi Dejean, 1828 Agonum versutum Sturm, 1824 Agonum viduum (Panzer, 1796) Platynus (=Agonum) assimile (Paykull, 1790) Perigona nigriceps (Dejean, 1831) Amara aenea (Degeer, 1774) Amara anthobia Villa, 1833 Amara apricaria (Paykull, 1790) Amara bifrons (Gyllenhal, 1810) Amara communis (Panzer, 1797) Amara consularis (Duftschmid, 1812) Amara convexior Stephens, 1828 Amara curta Dejean, 1828 Amara equestris (Duftschmid, 1812) Amara eurynota (Panzer, 1796) Amara famelica Zimmerman, 1832 Amara familiaris (Duftschmid, 1812) Amara fulva (Müller, 1776) Amara lucida (Duftschmid, 1812) Amara lunicollis Schiødte, 1837 Amara montivaga Sturm, 1825 Amara ovata (Fabricius, 1792) Amara plebeja (Gyllenhal, 1810) Amara praetermissa (Sahlberg, 1827) Amara similata (Gyllenhal, 1810) SUB VCH 1907- Notes 2005 ++ (RDB1) ? Established immigrant . Heaths, and woods on sand or peat. Windsor Forest 1980 (JAO) Rom ++ Widespread in grassland, woods etc. Rom ++ ++ ++ + * ++ > ++ ++ * * ? * ++ + ++ ++ ++ ++ ++ + + ++ ++ ++ + Widespread in wetlands, especially carr areas and woodland pools. Local in damp litter in wet woodland and marsh. Local in marshland in litter and tussocks. Rare in marshes and richer bogs. (Nb) Rare by flowing water, in carr and on river banks Local at margins of standing water in bare areas under stones etc. Rare [Wetlands]. Local in marshy areas. Local in dry areas including cultivated land. (Nb) Rare in marshes. ? [In marshes]. (Na) Rare in marshes. (Na) Local, on open wet heathland also in open woodland rides ( Common). Local in marshes and boggy areas especially on heathy commons. (Nb) Rare in marshy wetlands. Rare in marshy areas. Widespread in woods etc. under bark in moss. Recent immigrant. Increasing in compost/manure heaps. Southcote 2003 (JSD) Widespread in open habitats including ruderal areas. Local on open dry soils. Local in disturbed areas including ruderal sites and gardens. Also can be abundant at light. Local in open dry sites. Rare on and commons. (Nb) Rare on heathland, [downland] Local on gravelly sites, & downland. (Nb) Rare in open sandy places. (Nb) Rare on heaths and downland. Tubney 1899 Hardwick VCH Local in open disturbed areas with ruderal vegetation. (RDB3) Rare open heathy places. Widespread in open habitats including ruderal areas (Nb) Rare on disturbed sandy ground including cultivated sites. Cothill 1986 (PW) (Nb) dry grassland, on chalk sand and gravel]. Wytham Wood 1951 (WS) Local in open areas including heathland and gardens. Local in dry areas often in disturbed ruderal sites. Widespread in open disturbed areas. Local in damp clayey areas. (Nb) Local primarily on heathland, but also on downland. Widespread in dry open areas with crucifers. Rom Rom Rom ESax LRom LRom * ++ * ++ ++ ? ++ ++ ++ ++ + ++ 7 Carabidae (Ground Beetles) SUB 247 species + 1 sub-fossil Amara tibialis (Paykull, 1798) LRom Curtonotus (=Amara) aulica (Panzer, 1796) LRom Curtonotus (=Amara) convexiuscula (Marsham, 1802) Zabrus tenebrioides (Goeze, 1777) Harpalus rufipes (Degeer, 1774) Rom Harpalus affinis (Schrank, 1781) Rom Harpalus anxius (Duftschmid, 1812) Harpalus attenuatus Stephens, 1828 Harpalus dimidiatus (Rossi, 1790) Harpalus honestus (Duftschmid, 1812) Harpalus latus (Linneaus, 1758) Harpalus rubripes (Duftschmid, 1812) Harpalus rufitarsis (Duftschmid, 1812) Harpalus servus (Duftschmid, 1812) Harpalus smaragdinus (Duftschmid, 1812) Harpalus tardus (Panzer, 1796) Harpalus tenebrosus Dejean, 1829 Harpalus vernalis (Duftschmid, 1812) Ophonus (=Harpalus) ardosiacus Lutschnik, 1922 Ophonus (=Harpalus) azureus (Fabricius, 1775) Rom Ophonus (=Harpalus) melleti Heer, 1837 Ophonus (=Harpalus) punctatulus (Duftschmid, 1812) Ophonus (=Harpalus) puncticeps (Stephens, 1828) Ophonus (=Harpalus) puncticollis (Paykull, 1798) Rom Ophonus (=Harpalus) rufibarbis (Fabricius, 1792) Ophonus (=Harpalus) rupicola Sturm, 1818 ??? Ophonus (=Harpalus) schaubergerianus Puel, 1937 Anisodactylus binotatus (Fabricius, 1787) Anisodactylus nemorivagus (Duftschmid 1812) Scybalicus oblongiusculus (Dejean, 1829) Trichocellus placidus (Gyllenhal, 1827) Bradycellus distinctus (Dejean, 1829) Bradycellus harpalinus (Serville, 1821) Bradycellus ruficollis (Stephens, 1828) Bradycellus sharpi Joy, 1912 Bradycellus verbasci (Duftschmid, 1812) Stenolophus mixtus (Herbst, 1784) Stenolophus teutonus (Schrank, 1781) VCH 19072005 ++ ++ + ++ ++ ++ + + + ++ ++ ++ * ++ ++ Notes Widespread in open habitats including ruderal areas. Widespread especially in disturbed areas with abundant composites. Local on open disturbed dry sites especially ruderal sites. Also comes to light. (Na) Open ground on light soils, esp. field margins. Phytophagous. Widespread and common, frequent at MV light traps. Widespread and common. Rare? Tubney 1986 (PW) Primarily coastal. Tubney 1907 (HSD) (Nb) Rare on chalk downland. Hardwick VCH (RDB1) Streatley Downs 1905 Widespread on open ground of many kinds. Local on dry open soils especially on sandy heaths. Local though abundant on some sandy heaths. (Nb) Rare on disturbed sandy soils. . Local on dry sand and gravely soils, frequent on heathland. (Na) [Coastal cliffs on dry grassland]. * ++ + + + (Nb) Local, increasing? Grassland, ruderal sites, gardens etc, frequent at MV light traps. (Nb) Rare on open chalk downland. ++? (Na) Rare on dry sandy and chalk grasslands.. Little Wittenham NR 1985 (M.Hirst) (Na) [Open chalky and sandy sites] ++ + + Local in open dry sites, including ruderal areas, also frequent at MV light traps. (RDB3) [Calcareous grassland]. Local in a variety of habitats especially on sandy soils. (Nb) Calcareous grassland]. , ++ (Nb) Rare in open dry places. + * + ++ * + ++ ++ ++ ++ ++ Rare in marshy areas (Na) Rare on dry sandy heathland. Import? Rare? In marshy areas. (Na) Local on sandy soils especially on heathland often under heather. Rare on sandy soils especially on heathland. Shotover 1990 (PW) Local in damp shady places. Local on sandy soils. Also frequent at light traps. Local in wetlands, pond margins in litter and under stones. (Nb) Rare in wetlands, pond margins, also in old gravel pits. Heron’s Nest 2003 (JSD) 8 Carabidae (Ground Beetles) 247 species + 1 sub-fossil Acupalpus brunnipes (Sturm, 1825) Acupalpus consputus (Duftschmid, 1812) Acupalpus dorsalis (Fabricius, 1787) Acupalpus dubius Schilsky, 1888 Acupalpus exiguus Dejean, 1829 Acupalpus flavicollis (Sturm, 1825) Acupalpus meridianus (Linneaus, 1761) Licinus depressus (Paykull, 1790) Licinus punctatulus (Fabricius, 1792) Badister bullatus (Schrank, 1798) Badister dilatatus Chaudoir, 1837 Badister peltatus (Panzer, 1797) Badister sodalis (Duftschmid, 1812) Badister unipustulatus Bonelli, 1813 Panagaeus bipustulatus (Fabricius, 1775) Panagaeus cruxmajor (Linnaeus, 1758) Chlaenius nigricornis (Fabricius, 1787) Chlaenius vestitus (Paykull, 1790) Callistus lunatus (Fabricius, 1775) Oodes helopioides (Fabricius, 1792) Odacantha melanura (Linnaeus, 1767) Lebia chlorocephala (Hoffmannsegg, 1803) Lebia cruxminor (Linneaus, 1758) Lebia cyanocephala (Linneaus, 1758) Demetrias atricapillus (Linneaus, 1758) Demetrias imperialis (Germar, 1824) Dromius agilis (Fabricius, 1787) Dromius angustus Brullé, 1834 Dromius linearis (Olivier, 1795) Dromius melanocephalus Dejean, 1825 Dromius meridionalis Dejean, 1825 Dromius notatus Stephens, 1827 Dromius quadrimaculatus (Linneaus, 1758) Dromius quadrinotatus (Zenker, 1800) Dromius quadrisignatus Dejean, 1825 Microlestes maurus (Sturm, 1827) Microlestes minutulus (Goeze, 1777) * Metabletus foveatus (Fourcroy, 1785) Metabletus obscuroguttatus (Duftschmid, 1812) Metabletus truncatellus (Linneaus, 1761) Polistichus connexus (Fourcroy, 1785) Brachinus crepitans (Linneaus, 1758) SUB VCH 19072005 * Rom > + ++ Br/Sax > * ++ + * Rom ++ ? * ++ * ++ * ++ ++ * ++ E.Ho ++ * * * ++ ++ ++ + LRom + ++ + * LBr ++ ++ * LRom ++ ++ LRom ++ ++ ++ ++ LRom * Notes (Na) [Marsh, saltmarsh]. (Nb) Rare in marshy areas, also comes to light. Local in boggy wetlands. Local in marsh and wet heath areas. (Nb) Rare in marshes. (Na) Rare in boggy areas. , . Rare? [Open areas on clay and sand including cultivated ground]. (Nb) Rare on open chalk downland. Dry Sandford 1976 (PW) (Na) ? Extinct [open chalky areas]. , Widespread and common including waste ground and large gardens. (Nb) Rare in wetlands. Dinton ? 1997 (Na) [In marshes]. V.local in wet woodland, and marshy areas. Pingewood 2003 (JSD) (Nb) Rare in [Marsh, carr and fen]. (Nb) Rare in open dry places in grasslands including ruderal sites. (RDB2) Tubney area (Nb) Rare wetlands, pond margins. Local at margins of water in old pits under stones etc. avoiding well vegetated ground. (RDB1) Extinct, (Nb) Rare in wetlands. Dinton Pastures 1997 ((DIBH) (Nb) Subfossil only [wetlands] Early E.Holocene (Robinson, 1993) (Nb) Rare. Grassland with Hypericum. Berrys lane 2003 (JSD) (RDB1) (RDB1) Rare. Formerly on chalk downland. Widespread amongst litter in rough grassy areas. (Nb) Very local in wetlands amongst tall emergent vegetation. Widespread on trees, including conifers. Sulhamstead 2003 (JSD) Rare on pines. First recorded at in 1908 . Widespread in grass tussocks etc. Widespread and common on the ground or in tussocks. Local. Arboreal on hardwoods and conifers. ? [Sparsely vegetated dry areas]. Widespread on deciduous trees and pine esp. on dead and moribund boughs and branches, often under bark. Widespread on deciduous trees esp. on dead and moribund boughs and branches, often under bark. (RDB1) ? On deciduous trees. Local on the ground in dry open areas, including open downland. Rare. Recent immigrant Widespread in open dry areas on heaths and disturbed sandy areas. Local in damp areas. Calcott 2003 (JSD) Local in open sparsely vegetated areas. (RDB2) Migrant? Margins of ponds, attracted to light traps. (Nb) Rare on chalky soils, no post 1980 records, possibly extinct? 9 Carabidae (Ground Beetles) 247 species + 1 sub-fossil Bombardier Beetle SUB VCH 1907- Notes 2005 Haliplidae (Crawling Water Beetles) 16 species Brychius elevatus (Panzer, 1793) Peltodytes caesus (Duftschmid, 1805) Haliplus confinis Stephens, 1828 Haliplus flavicollis Sturm, 1834 Haliplus fluviatilis Aubé, 1836 Haliplus fulvus (Fabricius, 1801) Haliplus heydeni Wehncke, 1875 Haliplus immaculatus Gerhardt, 1877 Haliplus laminatus (Schaller, 1783) Haliplus lineatocollis (Marsham, 1802) Haliplus lineolatus Mannerheim, 1844 Haliplus obliquus (Fabricius, 1787) Haliplus ruficollis (Degeer, 1774) Haliplus sibiricus Motschulsky,1860 = wehnckei Gerhardt, 1877 SUB VCH 19072005 + ++ ++ ++ ++ ++ * ++ ++ ++ ++ ++ ++ ++ Hygrobiidae 1 species Hygrobia hermanni (Fabricius, 1775) Screech Beetle SUB VCH 1907- Notes 2005 ++ Widespread but local in ponds especially with silty substrates. > 20 post 1980 records. Noteridae 2 species Noterus clavicornis (Degeer, 1774) Noterus crassicornis (Müller, 1776) SUB VCH 1907- Notes 2005 ++ Widespread and common in ponds, canals, and lakes. > 30 post 1980 records. (Nb) Extinct? * Dytiscidae (Diving Beetles) 81 species + 1 sub-fossil Laccophilus hyalinus (Degeer, 1774) Laccophilus minutus (Linneaus, 1758) Hyphydrus ovatus (Linneaus, 1761) Hydroglyphus pusillus (Fabricius, 1781) Hygrotus confluens (Fabricius, 1787) Hygrotus decoratus (Gyllenhal, 1810) Hygrotus inaequalis (Fabricius, 1777) Hygrotus impressopunctatus (Schaller,1783) Hygrotus nigrolineatus (von Steven, 1808) Hygrotus versicolor (Schaller, 1783) SUB VCH 19072005 ++ ++ ++ ++ ++ > ++ ++ ++ ++ Notes Very Local in flowing water especially in submerged gravel and near weirs. (Nb) Local but increasing? Weedy ponds, Green Park 2001 (JSD) Local in ponds, lakes and canals with Charophytes Very Local in large ponds, canals. Widespread in ponds, lakes, canals etc. Very Local in large ponds, (Nb) Very Local in small detritus ponds. Widespread in weedy ponds. (Nb) Weedy streams and ponds. Sheffield Bottom 2003 (JSD) Widespread in streams, ditches and weedy ponds. Local in canals, ponds, flooded gravel pits. Local, but can be a rapid coloniser of new ponds with Chara Widpespread in detritus ponds. Local in standing water. Notes Local, in slow flowing water Widespread and common in ponds, canals, and lakes. Widespread and common in ponds, canals, and lakes. (Nb) Widespread, a rapid coloniser of new ponds with silty substrates. Widespread a rapid coloniser of new ponds with silty substrates. (Nb) Rare in well vegetated marshy wetland and pond margins. Widespread and common in ponds, canals, and lakes. Widespread in ponds, especially on heathland. (Na) Recent immigrant increasing in new ponds. Green Park 2002 (JSD) (Nb) Local Sheffield Bottom 2003 (JSD) 10 Dytiscidae (Diving Beetles) SUB VCH 1907- Notes 81 species + 1 sub-fossil 2005 Hydroporus angustatus Sturm, 1835 ++ Widespread, and common in ponds, canals, and lakes, avoiding acidic heath ponds. Hydroporus discretus Fairmaire, 1859 ++ Very Local in spring fed puddles and pools. Hydroporus erythrocephalus (Linnaeus, 1758) ++ Widespread and common in ponds and lakes, especially in acidic heath ponds. Hydroporus ferrugineus Stephens, 1829 + (Nb) Rare. Confined to small boggy springhead areas on the slopes of Leith Hill. Hydroporus gyllenhali Schiødte, 1841 ++ Widespread and common in ponds and lakes, especially in acidic heath ponds. Hydroporus incognitus Sharp, 1869 ++ Local in shaded pools and marshy pools beside streams. Hydroporus longulus Mulsant, 1860 ++ (Nb) Rare in spring fed streams. Snelsmore Common 2003 (JSD). + Hydroporus marginatus (Duftschmid, 1805) Hydroporus melanarius Sturm, 1835 ++ Local in small pools on heathland. Hydroporus memnonius Nicolai, 1822 ++ Widespread in woodland pools in flooded leaf litter. Hydroporus neglectus Schaum, 1845 ++ (Nb) Very Local in shaded acid pools. 6 post 1980 records. Hydroporus nigrita (Fabricius, 1792) ++ Local in a variety of small water bodies including small seasonal pools. Hydroporus obscurus Sturm, 1835 + Rare in flooded Sphagnum on heaths and in bogs. Hydroporus palustris (Linnaeus, 1761) ++ Widespread in detritus ponds and marshy areas by lakes and flowing water. Hydroporus planus (Fabricius, 1781) ++ Widespread rapid coloniser of new and recently cleared ponds also in water troughs. Hydroporus pubescens (Gyllenhal, 1808) ++ Widespread in acid standing water. (RDB2) Extinct? [Old ponds and wetlands, oxbows etc.]. . Hydroporus rufifrons (Müller, 1776) Hydroporus striola (Gyllenhal, 1827) ++ Local in standing water. Hydroporus tessellatus Drapiez, 1819 ++ Widespread. Rapid coloniser of new and recently cleared ponds also in water troughs, isolated puddles. Hydroporus tristis (Paykull, 1798) ++ Local in small pools especially Sphagnum pools on heathland. Hydroporus umbrosus (Gyllenhal, 1808) + Local in small pools especially Sphagnum pools on heathland. Suphrodytes dorsalis (Fabricius, 1784) ++ Local in shaded detritus ponds. Deronectes latus (Stephens, 1829) * (Nb) Extinct. Stictonectes lepidus (Olivier, 1795) * (RDB2) Extinct? . Graptodytes flavipes (Olivier, 1795) * Graptodytes granularis (Linnaeus, 1767) ++ (Nb) Rare. Richly weeded ponds. Hutchins Copse 1998 (JSD). Graptodytes pictus (Fabricius, 1787) * Rare. No modern records. [Ponds and ditches] last record Porhydrus lineatus (Fabricius, 1775) * Rare. No modern records. [Ponds, ditches etc.]. Nebrioporus assimilis (Paykull, 1798) * Extinct. [Ponds, lakes and drains] Nebrioporus depressus (Fabricius,1775) ssp. ++ Local in rivers and permanent streams. Also occasionally in deep ponds in pits etc. elegans (Panzer, 1794) Stictotarsus duodecimpustulatus (Fabricius, 1792) ++ ILocal in rivers and streams especially in bare sections under submerged ledges etc. Oreodytes sanmarki (Sahlberg, 1826) ++ Rare. Gravelly streams in root tangles, shade tolerant. Copelatus haemorrhoidalis (Fabricius, 1787) ++ Local. In weedy detritus ponds and bog pools. Scarodytes halensis (Fabricius, 1787) + Platambus maculatus (Linnaeus, 1758) ++ Local in flowing water, streams, and rivers. Tolerant of shade. Agabus affinis (Paykull, 1798) ++ Very Local in acid ponds, typically in flooded Sphagnum. Agabus bipustulatus (Linaneus, 1767) Ro/ESa ++ Ubiquitous in stagnant water including horse troughs etc., a strong flyer. x Agabus congener (Thunberg, 1794) * Sunningdale Agabus didymus (Olivier, 1795) ++ Local in flowing water amongst marginal vegetation. Agabus guttatus (Paykull, 1798) ++ Local in spring fed streams and well-heads, also outfalls of man-made drains. 11 Dytiscidae (Diving Beetles) 81 species + 1 sub-fossil Agabus labiatus (Brahm, 1790) Agabus melanarius Aubé, 1837 Agabus nebulosus (Forster, 1771) Agabus paludosus (Fabricius, 1801) Agabus sturmii (Gyllenhal, 1808) Agabus uliginosus (Linnaeus, 1761) Agabus unguicularis (Thomson, 1867) Ilybius aenescens Thomson, 1870 Ilybius ater (Degeer, 1774) The Mud-dweller Ilybius chalconatus (Panzer, 1796) Ilybius fenestratus (Fabricius, 1781) Ilybius fuliginosus (Fabricius, 1792) Ilybius guttiger (Gyllenhal, 1808) Ilybius montanus Zimmermann, 1915 Ilybius quadriguttatus (Lacordaire, 1835) Rhantus exsoletus (Forster, 1771) Rhantus grapii (Gyllenhal, 1808) Rhantus suturalis (MacLeay, 1825) Rhantus suturellus (Harris, 1828) Colymbetes fuscus (Linnaeus, 1758) Hydaticus seminiger (Degeer, 1774) Acilius sulcatus (Linnaeus, 1758) Dytiscus circumcinctus Ahrens, 1811 Dytiscus circumflexus Fabricius, 1801 Dytiscus marginalis Linnaeus, 1758 Great Diving Beetle Dytiscus semisulcatus Müller, 1776 Gyrinidae (Whirlygigs) 6 species Gyrinus aeratus Stephens, 1832 Gyrinus marinus Gyllenhal, 1808 Gyrinus substriatus Stephens, 1828 SUB Rom ESax Hydraenidae 16 species + 3 sub-fossils Ochthebius aeneus Stephens, 1835 ++ ++ * + ++ ++ ++ ++ + ++ ++ ++ + ++ + ++ ++ ++ * ++ ++ + SUB Rom Gyrinus suffriani Scriba, 1855 Gyrinus urinator Illiger, 1807 The Artist Orectochilus villosus (Müller, 1776) Hairy whirlygig VCH 19072005 + ++ ++ ++ LBrA SUB Notes (Nb) Rare. In shallow heath pools. (Nb) Rare. Seepages in woodlands, amongst flooded Sphagnum and leaf litter. Snelsmore Common 2003 (JSD) Widespread. In silt ponds esecially. New or recently cleared sites, a strong flier also frequent in troughs etc. Local, in weedy margins of flowing water, also around small seepages and springheads in flooded litter and moss. Widespread in detritus ponds. (Nb) Rare in shallow fens Hutchin’s Copse 1998 (JSD) Rare in mesotrophic pools. (Nb) Rare. In acid ponds typically in flooded Sphagnum. Widespread in all kinds of standing waters. Also comes to light. Local. In ponds and rut pools in wooded areas, (Nb) Very Local. In large ponds, lakes especially in heathland areas. Widespread in ponds and weedy margins of flowing water. Also comes to light. (Nb) Very Local. In pools on heathland. Local in acid ponds especially on heathland. Local in detritus ponds. Local in detritus ponds and marshy areas (Nb) Local in richly vegetated (usually old) ponds and in Sphagnum in richer bog pools. (Nb) Widespread good coloniser of new and recently cleared silt pools. Local. In peat pools on heathland. Frequent species of ponds of all kinds. (Nb) Local, but increasing. In mossy margins of pools especially on heaths. Tolerant of shading. Local in a wide variety of still waters, but intolerant of fish stocking. (Nb) Rare. Burton (EJP) (Nb) Scarce, increasing? In ponds with open silty substrates. Green Park, Pingewood 2001 (JSD) Ubiquitous in standing water of all kinds, especially where amphibians are breeding, as larvae specialise in tadpoles. Local, most records from stagnant water in heath ponds and peat pools. VCH 1907- Notes 2005 (Nb) ? Canals and slow flowing river margins. ++ Local, on open water especially on large water bodies such as lakes and canal margins. ++ Widespread and often abundant on ponds, canals, ditches and lakes. ++ Cothill (DB) ++ (Nb) Local. Virginia Water 1985 (RBA). ++ Local, on streams often in deep shade. Largely nocturnal and easily overlooked. VCH 1907- Notes 2005 (RDB1) Extinct in UK? Formerly found on heath pools in Surrey, last seen in UK in 1913. 12 Ochthebius bicolon Germar, 1824 Ochthebius dilatatus Stephens, 1829 Ochthebius minimus (Fabricius, 1792) Ochthebius pusillus Stephens, 1835 Ochthebius viridis Peyron, 1858 Enicocerus exsculptus (Germar, 1824) Hydraena britteni Joy, 1907 Hydraena flavipes Sturm, 1836 Hydraena gracilis Germar, 1824 Hydraena minutissima Stephens Hydraena nigrita Germar, 1824 Hydraena pulchella Germar, 1824 Hydraena riparia Kugelann, 1794 Hydraena rufipes Curtis, 1830 Hydraena testacea Curtis, 1830 Limnebius aluta Bedel, 1881 Limnebius nitidus (Marsham, 1802) Limnebius papposus Mulsant, 1844 ESax ++ + (Nb) Very Local at margins of streams and trickles. Very Local in muddy trickles and pond margins. Frequent at margins of ponds in litter, and moss. (RDB3) Rare in silty margins of newly cleared ponds. (Nb) [Coastal pools] (Nb) Extinct? Rare. I (Nb) Extinct? Rare in gravel in streams. ( ) Burghfield (Robinson, 1992) (Nb) Local in flowing water especially near springheads and in small streams. (RDB3) Extinct? Late Bronze Age Burghfield (Robinson, 1992) Frequent in ponds and wetlands. (Nb) Rare in (Nb) Local in flooded leaf litter in woodland pools and puddles. (RDB3) [Fens] (Nb) Mud and litter at margins of streams and ponds. Burghfield (Robinson, 1992) (Nb) In flooded fen and marsh litter ++ Widespread at margins of ponds and streams. ++ ++ ++ + + + lBrA + LBr LBr LBrA LR/ESa x Limnebius truncatellus (Thunberg, 1794) ++ + ++ Helophoridae 18 species Helophorus aequalis Thomson, 1868 Helophorus arvernicus Mulsant, 1846 Helophorus brevipalpis Bedel, 1881 Helophorus dorsalis (Marsham, 1802) Helophorus flavipes (Fabricius, 1792) Helophorus grandis Illiger, 1798 Helophorus granularis (Linnaeus1761) Helophorus griseus Herbst, 1793 Helophorus laticollis Thomson, 1853 Helophorus longitarsis Wollaston, 1864 Helophorus minutus Fabricius, 1775 Helophorus nanus Sturm, 1836 Helophorus nubilus Fabricius, 1777 Helophorus obscurus Mulsant, 1844 SUB VCH 19072005 Rom ++ + R/ESax ++ + + R/ESax ++ + ++ Helophorus porculus Bedel 1881 Helophorus rufipes (Bosc d'Antic, 1791) Helophorus strigifrons Thomson, 1868 LRom LRom R/ESax Rom ++ ++ + ++ ++ ++ + Notes Common in ponds, pools and puddles. A strong flier and rapid coloniser of new and ephemeral pools. (Nb) Rare. In mud and silt at margins of rivers. Common in ponds and pools. Strong flier often landing on cars remote from water. (Nb) Rare in muddy woodland ponds and rut pools. Local. Frequent in acid pools and peat pools on heathland. Common in ponds, pools and puddles. A strong flier. (Nb) Rare, but under-recorded as winter active in shallow pools in woods, and commons. (Nb) Very Local in new ponds with little vegetation. (RDB2) Extinct? Formerly in shallow heath pools around . (RDB3) Rare in new ponds with little vegetation. Also at light. Green Park 2002 (JSD) Common in ponds, pools and puddles. A strong flier. (Nb) Rare in marshy pools [and fen areas] Open, often cultivated ground, at roots of plants]. Corallian sands Tubney etc Widespread in pools and ponds, especially common in small leaf-filled puddles in woodland, avoiding very acid water. Extinct? M ? [On dry ground at roots of plants] (Nb) Rare in marshy pools. 13 Sphercheidae Sphercheus emarginatus (Schaller, 1783) Georissidae (Mud-coater Beetles) 1 species Georissus crenulatus (Rossi, 1794) SUB LBrA VCH 1907- Notes 2005 (Nb) On exposed sandy river banks. Burghfield (Robinson, 1992) Hydrochidae 2 species Hydrochus angustatus Germar, 1824 Hydrochus elongatus (Schaller, 1783) SUB VCH 1907- Notes 2005 + (Nb) Local in weedy detritus ponds especially on heaths. + (RDB3) Rare in weedy detritus ponds. Hydrophilidae (Water Scavenger Beetles) 58 species Berosus affinis Brullé, 1835 Berosus luridus (Linnaeus, 1761) Berosus signaticollis (Charpentier, 1825) Chaetarthria seminulum (Herbst, 1797) Paracymus scutellaris (Rosenhauer, 1856) Anacaena bipustulata (Marsham, 1802) Anacaena globulus (Paykull, 1798) Anacaena limbata s. str. (Fabricius, 1792) Anacaena lutescens (Stephens, 1829) Laccobius atratus (Rottenberg, 1874) Laccobius biguttatus Gerhardt, 1877 Laccobius bipunctatus (Fabricius, 1775) Laccobius minutus (Linnaeus,1758) Laccobius sinuatus Motschulsky, 1849 Laccobius striatulus (Fabricius, 1801) Laccobius ytenensis Sharp 1910 Helochares lividus (Forster, 1771) Helochares punctatus Sharp, 1869 SUB VCH 19072005 ++ + + + + ++ ++ ++ ++ + > ++ ++ ++ ++ + ++ ++ Enochrus affinis (Thunberg, 1794) Enochrus coarctatus (Gredler, 1863) Enochrus nigritus Sharp, 1872 = isotae Hebauer, 1981 Enochrus melanocephalus (Olivier, 1792) Enochrus ochropterus (Marsham, 1802) Enochrus testaceus (Fabricius, 1801) Cymbiodyta marginellus (Fabricius, 1792) Hydrobius fuscipes (Linnaeus, 1758) Coelostoma orbiculare (Fabricius, 1775) R/ESax LBr Notes + ++ + (Nb) Local, increasing in new ponds with little vegetation, especially in sand, gravel and clay pits. (Nb) Rare in marshy pools especially on heathland. (Nb) Local increasing in weedy ponds especially on heaths. (Nb) Local in mud and moss in richer bogs and fen areas. (Nb) Seepage fed pools on heathland. (Nb) Local in mud and litter at margins of ponds, lakes, and canals. Common in ponds and pools, larvae in leaf litter especially in woods. Widespread in ponds and pools. Common in standing acid waters. (Nb) Very Local in flowing water in bogs and seepages. Rare in ponds. Frequent in a variety of ponds and weedy flowing water. Local and scarce in Surrey. (Nb) Local in new ponds etc. Local in silt at margins of ponds and flowing water. (Nb) Very Local in flowing water in bogs and seepages. Widespread and increasing in new ponds with little vegetation. (Nb) Local in heath pools, occasionally in mossy margins of richer ponds on the clays. Padworth Common 2004 (JSD) (Nb) Local in Sphagnum and other mossy litter in acid pools on heaths. Local in weedy margins of detritus rich ponds. (RDB3) Rare in weedy mesotrophic and base rich-fen areas and weedy ponds. ++ + ++ ++ ++ ++ (Nb) Very Local but increasingly frequent in new silty ponds. (Nb) Very Local in weedy mesotrophic areas of bogs and marshy ponds avoiding very acidic conditions. Local in detritus ponds. Local in detritus ponds and marshy lake margins. Common in all kinds of standing waters and weedy river margins. Also frequent at light. Local in detritus at margins of ponds and canals. 14 Hydrophilidae (Water Scavenger Beetles) 58 species Sphaeridium bipustulatum Fabricius, 1781 Sphaeridium lunatum Fabricius, 1792 Sphaeridium marginatum Fabricius, 1787 Sphaeridium scarabaeoides (Linnaeus, 1758) Cercyon analis (Paykull, 1798) Cercyon atomarius (Fabricius, 1775) Cercyon atricapillus (Marsham, 1802) Cercyon bifenestratus Küster, 1851 Cercyon convexiusculus Stephens, 1829 Cercyon granarius Erichson, 1837 Cercyon haemorrhoidalis (Fabricius, 1775) Cercyon laminatus Sharp, 1873 Cercyon lateralis (Marsham, 1802) Cercyon lugubris (Olivier, 1790) Cercyon marinus Thomson, 1853 Cercyon melanocephalus (Linnaeus, 1758) Cercyon pygmaeus (Illiger, 1801) Cercyon quisquilius (Linnaeus, 1761) Cercyon sternalis Sharp, 1918 Cercyon terminatus (Marsham, 1802) Cercyon tristis (Illiger, 1801) Cercyon unipunctatus (Linnaeus, 1758) Cercyon ustulatus (Preyssler, 1790) Megasternum obscurum (Marsham, 1802) Cryptopleurum crenatum (Kugelann, 1794) Crytopleurum minutum (Fabricius, 1775) Cryptopleurum subtile Sharp, 1884 Histeridae (Hister Beetles) 35 species + 1 sub-fossil Teretrius fabricii Mazur, 1972 Plegaderus dissectus Erichson, 1839 Plegaderus vulneratus (Panzer, 1796) Abraeus globosus (Hoffmann, 1803) Abraeus granulum Erichson, 1839 Aeletes atomarius (Aubé, 1842) Acritus homoeopathicus Wollaston, 1857 Acritus nigricornis (Hoffmann, 1803) Gnathoncus buyssoni Auzat, 1917 Gnathoncus nannetensis (Marseul, 1862) Gnathoncus nanus (Scriba, 1790) SUB LRom LBrA LBrA LRom LBrA LBrA R/Sax Rom SUB Rom LRom VCH 19072005 ++ ++ ++ ++ ++ ++ + ++ ++ ++ ++ ++ ++ ++ ++ ++ ++ ++ ++ + ++ ++ ++ ++ + + + VCH 19072005 * ++ > ++ ++ ++ + + > + > Notes Local in herbivore dung especially that of cattle. Local in herbivore dung especially that of cattle. Local in herbivore dung especially that of cattle. Local in herbivore dung especially that of cattle. Widespread in dung, compost heaps, and other rotting vegetation. Widespread in herbivore dung. (Nb) Rare? In dung. (Na) Rare on bare margisn of open water . Green Park 2002 (JSD0 Holybrook on shingle bar 2004 (JSD) Local in detritus at margins of all kinds of water bodies. (RDB3) Rare In tussocks on Thames floodplain. Widespread in dung. Naturalised, probably widespread in compost heaps etc., comes readily to light. Kennet Floodplin Southcote 2003 (JSD) Widespread in dung, and carrion. Widespread in dung, and carrion. Local in wet litter, dung, carrion etc. often near water. Widespread in herbivore dung. Widespread in herbivore dung. Widespread in dung. (Nb) Very Local in marsh litter at margins of ponds. Burghfield (Robinson, 1992) (Nb) Rare? In dung and compost heaps. (Nb) Local in marsh litter around ponds avoiding acid heath pools. Widespread in herbivore dung especially cow, and occasionally in carrion. (Nb) Local in detritus at margins of standing water. Also in sewage filter beds. Widespread and common in all kinds of decaying vegetation, dung, and fungi. (N) Rare [in dung, and other decaying organic matter]. Widespread in dung and compost. Rare in dung and rotting fungi. Notes (RDB1) ? [Predatory on Lyctus in hardwood especially oak including fresh oak palings]. (ECB). (Nb) Local under bark especially beech. Naturalised. Rare, under pine bark. Local under bark and fungoid wood especially beech. (Na) Rare in ancient wood/parkland on rotting hardwoods. . (RDB3) Rare. In frass of wood borings of [Lucanids] and longhorns. (RDB3) Rare on burnt ground on the fungus Pyronema confluen Local in rotting vegetation, haystacks etc. (Na) Rare in hollow trees and old bird nests in woodland. Rare in hollow trees and old bird nests in woodland. , . Rare in bird’s nests especially pigeon, granaries [also in carrion]. 15 Histeridae (Hister Beetles) SUB VCH 1907- Notes 35 species + 1 sub-fossil 2005 Gnathoncus schmidti Reitter, 1894 > Rare in bird’s nests. Saprinus aeneus (Fabricius, 1775) + Widespread in dung and carrion. Saprinus semistriatus (Scriba, 1790) ++ Local on dung and carrion. (RDBK) [Parasitic on Phaedon especially on water-cress Nasturtium. Increasing?]. Saprinus virescens (Paykull, 1798) * Myrmetes piceus (Paykull, 1809) + (Nb) Local in nests of Formica rufa on heaths, woods and conifer plantations. Dendrophilus punctatus (Herbst, 1792) ++ Local in hollow trees in old nests etc. Also with ant Lasius fuliginosus. Dendrophilus pygmaeus (Linnaeus, 1758) ++ Local in nests of Formica rufa. LRom Kissister minimus (Aubé, 1850) ++ Local at roots of plants in dry grassland. Often under Rumex acetosella. Carcinops pumilio (Erichson, 1834) + Local. Synanthropic in gardens etc. around compost heaps. Often enters hot kitchens! Paromalus flavicornis (Herbst, 1792) ++ Widespread under bark and in decaying timber especially beech. Onthophilus punctatus (Müller, 1776) ++ (RDBK) [In mole nests] JC Frilford R/ESax Onthophilus striatus (Forster, 1771) ++ Local on dung especially horse. LRom (Nb) [In cow dung] Sub-fossil Appleford (Robinson ) Hister bissexstriatus Fabricius, 1801 ? Hister impressus Fabricius, 1798 + Rare? Hister merdarius Hoffmann, 1803 > Rare? In rotting vegetation, and bird’s nests. Bradfield (NHJ) Hister striola Sahlberg, 1819 ++ Widespread in herbivore dung and rotting fungi, carrion. Hister unicolor Linnaeus, 1758 ++ Widespread in herbivore dung and rotting fungi, carrion. Margarinotus (Paralister) neglectus (Germar, ? [In carrion, moss]. (Fowler, 1889) 1813) Margarinotus (Paralister) purpurascens (Herbst, LRom + Rare on ground especially on sandy soils. 1792) Margarinotus (Paralister) ventralis (Marsuel, 1854) Rom ++ Widespread in dung, rotting vegetation and carrion. =carbonarius (Hoffmann, 1803) Margarinotus marginatus (Erichson, 1834) + (Nb) Rare? In subterranean mammal nests (mole, rabbit). Bradfield (NHJ) Atholus duodecimstriatus (Schrank, 1781) LRom ++ Widespread on dung. Peranus bimaculatus (Linnaeus, 1758) ++ Local, in dung and compost heaps. Ptiliidae 46 species Nossidium pilosellum (Marsham, 1802) Ptenidium formicetorum Kraatz, 1851 Ptenidium fuscicorne Erichson, 1845 Ptenidium gressneri Erichson, 1845 Ptenidium intermedium Wankowicz, 1869 Ptenidium laevigatum Erichson, 1845 Ptenidium nitidum (Heer, 1841) Ptenidium punctatum (Gyllenhal, 1827) Ptenidium pusillum (Gyllenhal, 1808) Ptenidium turgidum Thomson, C.G., 1855 Ptilium exaratum (Allibert, 1844) Ptilium horioni Rosskothen, 1934 Ptilium myrmecophilum (Allibert, 1844) SUB VCH 1907- Rare? In rotting wood, litter, and wood ant nests. 2005 ++ (N) Local in fungi especially dryad’s saddle Polyporus squamosus, + ? [In grass heaps] + (N) Local in frass and debris in old trees especially beech. + ? In marshy areas. + ? Local in litter and with ants. . Also in gardens ? . ? . + Local in litter and vegetable refuse. (RDBK) Old woodland in hollow trees, log piles etc. ( + ? + ? + Rare in wood ant Formica rufa nests. 16 Ptiliidae 46 species Oligella foveolata (Allibert, 1844) Oligella intermedia Besuchet, 1971 Micridium halidaii (Matthews, 1868) Ptiliola kunzei (Heer, 1841) Ptiliolum fuscum (Erichson, 1845) Ptiliolum marginatum (Aubé, 1850) Ptiliolum spencei (Allibert, 1844) Ptinella simsoni (Matthews, 1878) Ptinella aptera (Guérin-Méneville, 1839) SUB VCH 19072005 + + + + + + + + * ++ + + Ptinella britannica Matthews, 1855 Ptinella cavelli (Broun, 1893) Ptinella denticollis (Fairmaire, 1858) Ptinella errabunda Johnson, 1975 Ptinella limbata (Heer, 1841). Smicrus filicornis (Fairmaire & Laboulbéné, 1855) Microptilium pulchellum (Allibert, 1844) Baeocrara variolosa (Mulsant & Rey, 1867) Nephanes titan (Newman, 1834) Pteryx suturalis (Heer, 1841) Acrotrichis arnoldi Rosskothen, 1935 Acrotrichis atomaria (Degeer, 1774) Acrotrichis brevipennis (Erichson, 1845) Acrotrichis chevrolati (Allibert, 1844) Acrotrichis cognata (Matthews, 1877) Acrotrichis danica Sundt, 1958 Acrotrichis dispar (Matthews, 1865) Acrotrichis fascicularis (Herbst, 1793) Acrotrichis henrici (Matthews, 1872) Acrotrichis grandicollis (Mannerheim, 1844) Acrotrichis insularis (Mäklin, 1852) Acrotrichis intermedia (Gillmeister, 1845) Acrotrichis montandoni (Allibert, 1844) Acrotrichis rosskotheni Sundt, 1971 Acrotrichis rugulosa Rosskothen, 1935 Acrotrichis sericans (Heer, 1841) Acrotrichis sitkaensis (Motschulsky, 1845) Acrotrichis thoracica (Waltl, 1838) Rare? In rotting wood, litter, and wood ant nests. (RDBK) ?). (RDBK) (RDBK) Rare in dead wood. . ? In grass heaps. ? In litter (RDBK) [Old woodlands in litter, dung, and fungi] ? In litter in garden ? Local? [In compost etc.], . Local under bark of dead trees, especially beech. Bradfield (NHJ) Local, under bark of dead trees. (N) Under bark of dead trees especially Oak., Bradfield (NHJ) . Local, under bark of dead trees. * Bradfield (NHJ) * * + + > + Bradfield (NHJ) ? Bradfield (NHJ) Local, under bark and in frass and debris in hollow trees. (RDBK) 1965 . + + + + + + + + ? + + + Local? In litter etc. >5 post 1980 sites. (N) ? [In litter, and decaying vegetable matter in a wide variety of habitats] (RDBK) ? [In dung and compost heaps]. ,. ? , (N) [Woodlands in dung and fungi] Local in litter etc. 4 ? In litter in woods and parks. ? In litter etc. ? In litter. ? [In litter etc.] Local in wood ant Formica rufa nests, and in litter, etc. ? [In litter] ? ? In compost, litter etc. . ? [In grass-heaps, litter] ? in litter 17 Leiodidae 77 species Triarthron maerkeli Märkel, 1840 Hydnobius latifrons (Curtis, 1840) Hydnobius punctatus (Sturm, 1807) Leiodes badia (Sturm, 1807) Leiodes cinnamomea (Panzer, 1793) Leiodes ferruginea (Fabricius, 1787) Leiodes flavescens (Schmidt, 1841) Leiodes gallica (Reitter, 1884) Leiodes gyllenhali (Stephens, 1829) Leiodes litura Stephens, 1835 Leiodes longipes (Schmidt, 1841) Leiodes lucens (Fairmaire, 1855) Leiodes lunicollis (Rye, 1872) Leiodes macropus (Rye, 1873) Leiodes nigrita (Schmidt, 1841) Leiodes obesa (Schmidt, 1841) Leiodes oblonga (Erichson, 1845) Leiodes polita (Marsham, 1802) = calcarata Erichson, 1845 Leiodes rufipennis (Paykull, 1798) Leiodes rugosa Stephens, 1829 Leiodes strigipenne Daffner 1983 Leiodes triepkii (Schmidt, 1841) Liocyrtusa vittata (Curtis, 1840) Colenis immunda (Sturm, 1807) Agaricophagus cephalotes Schmidt, 1841 Anisotoma humeralis (Fabricius, 1792) Anisotoma orbicularis (Herbst, 1792) Amphicyllis globus (Fabricius, 1792) Agathidium atrum (Paykull, 1798) Agathidium confusum Brisout, 1863 Agathidium convexum Sharp, 1866 Agathidium laevigatum Erichson, 1845 Agathidium marginatum Sturm, 1807 Agathidium nigrinum Sturm, 1807 Agathidium nigripenne (Fabricius1792) Agathidium rotundatum Gyllenhal, 1827 Agathidium seminulum (Linnaeus, 1758) Agathidium varians Beck, 1817 Ptomaphagus medius Rey, 1889 SUB VCH 1907- Notes 2005 (N) [In coniferous woodlands and parkland]. (RDBK) . Wytham (JJW) (N) [Woods and downland] Tubney , Wytham (JJW) * ? Rare on downland. (N) [Beech woods on chalk] Wytham 1906 (JJW), * * ? Wytham (JJW) * (RDBI) Tubney 1928 (JJW) (RDBK) [Chalky and sandy places]. * (RDBK) ? [Woodlands?]. * + Woods on the chalk? Tubney (JJW) (RDB1) ? Tubney 1907 (JJW) Not recorded in UK since 1936. * ++ (RDBK) ? [Old woods and sandy places] Silwood Park 1998-9 (RGB) in FIT (RDBK) ? [Sandy soils?]. * (RDBK) ? [Woods and open areas] * (RDBK) ? [Woodland]. Wytham (JJW) * + Rare? In woods and open areas. + (N) [Woodland and pasture woodland]. ++ The most frequently encountered Leiodes, frequent in flight interception and malaise traps, but hard to find by other means. + Local in sandy and chalky areas. Tubney Wytham (JJW) (N) [Downland, and sandy soils]. Tubney 1906, Wytham 1906 (JJW) * = L. flavicornis (Brisout, 1863 * (RDBK) ? Sandy, common under pines and firs. Tubney (JJW) ++ Local, in many habitats. Comes to light. Wytham (JJW) Burghfield Mill 2003 (JSD) + ? [Downland] JJW general (RDBK) Downland? * ++ Local in woodlands in fungi and fungoid wood. + Local especially in woodlands. Usually present in malaise/interception trap material, but hard to find by other means. + ? In rotten wood. + Local in woods and open habitats in moss and tussocks. Tubney (JJW) ++ (RDB1) Rare, [on fungi]. Silwood Park 1998-9 (RGB) in FIT * ? [In fungoid wood, moss, esp,. in woodland]. Shotover (JJW) * ? [In litter, log piles etc.]. Tubney (JJW) (N) ? p + Local under bark and in fungi. + Rare in sappy bark of various trees. . * Local in rotten hardwoods. Tubney (JJW) + Local in rotting wood. . + Local in rotting wood and damp leaf litter. + Local. 18 Leiodidae 77 species Ptomaphagus subvillosus (Goeze, 1777) Ptomaphagus varicornis (Rosenhauer, 1847) Nemadus colonoides (Kraatz, 1851) Nargus anisotomoides (Spence, 1813) Nargus velox (Spence, 1813) Nargus wilkini (Spence, 1813) Choleva agilis (Illiger, 1798) Choleva angustata (Fabricius, 1781) Choleva fagniezi Jeannel, 1922 Choleva glauca Britten, 1918 Choleva jeanneli Britten, 1922 Choleva oblonga Latreille, 1807 Choleva spadicea (Sturm, 1839) Sciodrepoides fumatus (Spence, 1813) Sciodrepoides watsoni (Spence, 1813) Catops chrysomeloides (Panzer, 1798) Catops coracinus Kellner, 1846 Catops fuliginosus Erichson, 1837 Catops fuscus (Panzer, 1794) Catops grandicollis Erichson, 1837 Catops kirbii (Spence, 1813) Catops longulus Kellner, 1846 Catops morio (Fabricius, 1787) Catops nigricans (Spence, 1813) Catops nigrita Erichson, 1837 Catops tristis (Panzer, 1793) Catopidius depressus (Murray, 1856) Parabathyscia wollastoni (Janson, 1857) Colon angulare Erichson, 1837 Colon appendiculatum (Sahlberg, 1822) Colon brunneum (Latreille, 1807) Colon dentipes (Sahlberg, 1822) Colon latum Kraatz, 1850 Colon rufescens Kraatz, 1850 Colon serripes (Sahlberg, 1822) Colon viennense Herbst, 1797 Colon zebei Kraatz, 1854 Leptinus testaceus Müller, 1817 SUB Rom VCH 19072005 ++ >? + * ++ + ++ ++ ++ + + + + ++ + + + > + + + + ++ + * * + * * + Notes Widespread especially in open grassy areas on sandy soils. (RDBK) [Plant litter and moss on chalky soils]. Local in hollow trees and rot holes, and roots of old trees Rare? In litter. Tubney (JJW) Local in litter, carrion etc. ? In litter and moss. . Tubney (JJW) ? [In litter, fungi etc.]. Local in litter in woods and grassy places. . ? [In litter, fungi, etc.] (N) [Mole nests]. Silwood Park 1998 (RGB) in FIT Rare? [In carrion]. . ? [In litter, fungi, etc.] Rare [On ground in dry sand places?] Local in carrion [and rotting fungi]. . Local in carrion. . Local in carrion. Local in carrion Widespread in carrion, and in nests of small mammals etc. Local in litter in woods, etc. H Local in carrion. Local in woods and open habitats Widespread in carrion, and nests and runs of small mammals. Local in carrion. Widespread in carrion. ? In woodlands. . Silwood Park 1998 (RGB) in FIT Rare? Subterranean in gardens especially on the chalk,. (RDBK) ? [Woods and ] (RDBK) ? [Grassy hillsides and woodlands] Local in litter etc. . (RDBK) [Grassy hillsides and woodlands] (N) [Woodland?]. (RDB1) [Chalk downland]. [In litter, fungi, etc (RDBK) ? [Grassy hillsides and woodlands] Tubney , Wytham (JJW) (RDBK) Rare on downland. Local. In nests of mammals especially moles, also in nest boxes used by wood mice. Formerly placed in Leptinidae. 19 Silphidae (Sexton & Burrowing Beetles) 17 species Nicrophorus humator (Gleditsch, 1767) Black sexton Nicrophorus interruptus Stephens, 1830 Nicrophorus investigator Zetterstedt, 1824 Nicrophorus vespillo (Linnaeus, 1758) Nicrophorus vespilloides Herbst, 1783 Necrodes littoralis (Linnaeus, 1758) Thanatophilus rugosus (Linnaeus, 1758) Thanatophilus sinuatus (Fabricius, 1775) Oiceoptoma thoracicum (Linnaeus, 1758) Dendroxena quadrimaculata (Scopoli, 1772) Silpha atrata Linnaeus, 1758 Silpha laevigata Fabricius, 1775 Silpha obscura Linnaeus, 1758 Silpha tristis Illiger, 1798 Scydmaenidae 24 species Eutheia formicetorum Reitter, 1881 Eutheia linearis Mulsant. Eutheia plicata (Gyllenhal, 1813) Eutheia schaumi Kiesenwetter, 1858 Eutheia scydmaenoides Stephens, 1830 Cephennium gallicum Ganglbauer, 1899 Neuraphes angulatus (Müeller & Kunze,1822) Neuraphes carinatus (Mulsant, 1861) Neuraphes elongatulus (Müller & Kunze, 1822) Neuraphes plicicollis Reitter, 1879 Neuraphes praeteritus Rye, 1872 Neuraphes talparum Lokay, 1921 Scydmoraphes helvolus (Schaum, 1844) Stenichnus bicolor (Denny, 1825) Stenichnus collaris (Müller & Kunze, 1822) Stenichnus godarti (Latreille, 1806). Stenichnus poweri (Fowler, 1884) Stenichnus pusillus (Müller & Kunze, 1822) Stenichnus scutellaris (Müller & Kunze, 1822 Microscydmus minimus (Chaudoir, 1845) Microscydmus nanus (Schaum, 1844) SUB LRom Rom VCH 1907- Notes 2005 ++ Widespread on carrion, also frequent at MV light. ++ ++ ++ ++ ++ + + + * ++ ++ Rom ++ SUB VCH 19072005 + * * + > + + > ++ + ++ + + + ++ * + * (Nb) Local on carrion, also at mv light Local in carrion, Local on carrion. Local on carrion, also at mv light. Rare, increasing ? Comes to light. Local in carrion. Local in carrion. Local, declining? In carrion. (Na) Rare. On oak trees. Widespread and frequent in woodlands sheltering under bark. Local in grassland especially on the chalk, but also on corallian sands. (RDB2) Sub-fossil only Local but increasing inland? In rank grassland with snails. Notes (RDB1) Rare in rotten wood esp, beech Windsor Forest 1989 (JAO) (RDB1) Windsor Forest (RDBK) Bradfield (NHJ) ?? (N) ? [Eurytop (N) Rare ? Eurytopic, B Local in woods, often on moss clad tree trunks. Rare in woodlands, Tubney (JJW). (RDBIn) Rare in moss and litter especially on N Local in moss and litter in grassland and woods. . (N) Under bark, in moss and rotten timb (N) ? [Woodland] ? Woodlands? Silwood Park 1998 (RGB) in FIT (N) ? [In litter and plant debris] Local under bark and on hardwoods. Widespread in woods and parkland in litter, tussocks etc. (RDB3) Windsor Silwood Park 1998-9 (RGB) in FIT (RDBK) [In litter, ant nests etc.] (N) ? Fowler (1889) l ? [Open grassland]. (RDB3) Rare in ancient woods in hollow trees etc. Windsor Great park (N) Woodland. 20 Euconnus denticornis (Müller & Kunze, 1822) Euconnus duboisi Mequignon, 1929 Euconnus fimetarius (Chaudoir, 1845) Euconnus hirticollis (Illiger, 1798) Euconnus maeklini (Mannerheim, 1844) * ++ + * + Euconnus pragensis (Machulka, 1923) Euthiconus conicollis (Fairmaire, 1855) Scydmaenus rufus Müller & Kunze, 1822 + Scydmaenus tarsatus Müller & Kunze, 1822 + Staphylinidae (Rove Beetles) species SUB Omaliinae Philorinum sordidum (Stephens, 1834) Orochares angustatus (Erichson, 1840) Anthobium atrocephalum (Gyllenhal, 1827) Anthobium unicolor (Marsham, 1802) Olophrum piceum (Gyllenhal, 1810) Acidota cruentata Mannerheim, 1830 Rom Acidota crenata (Fabricius, 1792) Lesteva hanseni Lohse, 1953 Lesteva heeri Fauvel, 1872 Lesteva longoelytrata (Goeze, 1777) BrAge Lesteva pubescens Mannerheim, 1830 Lesteva punctata Erichson, 1839 Eusphalerum luteum (Marsham, 1802) Eusphalerum minutum (Fabricius, 1792) Eusphalerum primulae (Stephens, 1834) Eusphalerum sorbi (Gyllenhal, 1810) Eusphalerum torquatum (Marsham, 1802) Phyllodrepa floralis (Paykull, 1789) Phyllodrepa nigra (Gravenhorst, 1806) Dropephylla devillei Bernhauer, 1902 = grandiloqua Luze, 1910 Dropephylla gracilicornis (Fairmaire & Laboulbène, 1856) Dropephylla ioptera (Stephens, 1834) Dropephylla vilis (Erichson, 1840) Hapalaraea pygmaea (Paykull, 1800) Hypopycna rufula (Erichson, 1840) Acrolocha minuta (Olivier, 1795) Acrolocha sulcula (Stephens, 1834) (RDBIn) [In moss and litter on Downland]. Wytham (JJW) Synanthropic in stable debris etc. Silwood Park 1998 (RGB) in FIT ?F Very Local in marsh litter. (RDBI) Windsor (RDB1) Windsor Great park (PMH?/JAO) (RDB2) Local but increasing? A……lane 1998 (JSD) Under bark including pine, also wintering in manure heaps, Local in compost/manure heaps. VCH 1907- Notes 2005 * ++ ++ ++ + ++ ++ ++ ++ + > ++ * + ? On broom and gorse flowers. (RDBIn ) Bradfield 1903 (NHJ) Widespread in litter and carrion. Widespread in litter and carrion. Local in moss in woods and on heathy commons. (Nb) ? [In moss and under conifer bark etc]. Rare in Sphagnum moss etc. Rare on mossy banks of flowing water Widespread in wetlands especially seepages and mossy areas including bogs. Widespread in wetlands and damp grassland. Local in moss on banks of streams especially in wooded areas. Rare? Widespread on flowers especially umbellifers in woods. ? [In mires]. Local in flowers of primrose Primula in spring. ? [In woods at blossoms etc.] Rare. Widespread on blossom especially hawthorn. Windsor Recent immigrant to southern England, primarily Scottish. Rare under bark. (N) [Under bark and in rotten wood]. ++ ++ + + + + Frequent under bark of hardwoods, and on blossoms especially hawthorn. Saproxylic. Local under bark of hardwoods. Rare under bark in fungi. , (RDBK) ? Subterranean in small mammal runs etc. . (N) ?. ? In decaying vegetation. . 21 Staphylinidae (Rove Beetles) species Omalium allardi Fairmaire & Brisout, 1859 Omalium caesum Gravenhorst, 1806 Omalium excavatum Stephens, 1834 Omalium exiguum Gyllenhal, 1810 Omalium italicum Bernhauer, 1902 Omalium oxyacanthae Gravenhorst, 1806 Omalium rivulare (Paykull, 1789) Omalium rugatum Mulsant & Rey, 1880 Omalium septentrionis Thomson, 1856 Xylostiba bosnica (Bernhauer ) Phloeonomus punctipennis Thomson, 1867 Phloeonomus pusillus (Gravenhorst, 1806) Phloeostiba plana (Paykull, 1792) Xylodromus concinnus (Marsham, 1802) Xylodromus depressus (Gravenhorst, 1802) Coryphium angusticolle Stephens, 1834 Proteininae Metopsia retusa (Stephens, 1834) Megarthrus affinis Miller, 1852 Megarthrus denticollis (Beck, 1817) Megarthrus depressus (Paykull, 1789) Megarthrus hemipterus (Illiger, 1794) Megarthrus sinuatocollis (Boisduval & Lacordaire, 1835) Proteinus atomarius Erichson, 1840 Proteinus brachypterus (Fabricius,1792) Proteinus macropterus (Gravenhorst, 1806) Proteinus ovalis Stephens, 1834 Micropeplinae Micropeplus fulvus Erichson, 1840 Micropeplus porcatus (Paykull, 1789) Micropeplus staphylinoides (Marsham, 1802) Micropeplus tesserula Curtis, 1828 Pselaphinae Bibloporus bicolor (Denny, 1825) Bibloporus minutus Raffray, 1914 Bibloplectus ambiguus (Reichenbach,1816) Bibloplectus delhermi Guillebeau, 1888 Euplectus bonvouloiri rosae Raffray, 1910 Euplectus duponti Aubé, 1833 Euplectus fauveli Guillebeau, 1888 SUB BrAge VCH 1907- Notes 2005 (N) Synanthropic. In houses, hospitals etc. + Local in leaf litter and tussocks in a variety of habitats including gardens. + Rare? [In litter] . (N) ? [In decaying vegetatble matter, fungi in grasslands and gardens] . (Fowler, 1888) + Rare? In litter. + ? Local in rotting vegetation. ++ Widespread in fungi, carrion and rotting vegetation in many habitats including gardens. + (N) Rare? In decaying vegetation including compost heaps. . + , recently at Under bark Cothill 14.xii.2003, Windsor orest + Widespread under hardwood bark especially oak and beech. + Local under conifer bark, especially pine. + Rare under bark and at sap. + Rare on fungi. ? Park. + Rare under bark. ++ + + + * + Widespread in moss and haystack debris etc. Rare? ? In litter. ?, (Na) Rare in rotting fungi especially Beefsteak Fistulina hepatica. 1975 . ? > ++ ++ + Rare? Widespread in rotting fungi. Rare? In fungi. Widespread in decaying vegetation and carrion in grassland and gardens. + + + + Local in rotting vegetation including compost heaps. Rare? In rotting vegetation. Radley (Rob Widespread in manure and compost heaps Local in rotting vegetation. + + + + ++ ++ > Local under bark and in dead wood. (Nb) Rare under bark and in dead wood in old woods and parklands. Local in moss and litter in wetlands (RDBK) Rare in wetlands (N) Rare in dead wood. Silwood Park 1998 (RGB) in FIT (N) [litter in woods] , .Bradfield (NHJ) Silwood Park 1998 (RGB) in FIT (N) Rare in old nests, under bark etc. 22 Staphylinidae (Rove Beetles) species Euplectus infirmus Raffray, 1910 Euplectus karsteni (Reichenbach, 1816) Euplectus kirbyi Denny, 1825 Euplectus nanus (Reichenbach, 1816) Euplectus piceus Motschulsky, 1835 Euplectus sanguineus Denny, 1825 Euplectus signatus (Reichenbach, 1816) Euplectus tholini Guillebeau, 1888 Plectophloeus nitidus (Fairmaire, 1857) Trimium brevicorne (Reichenbach, 1816) Batrisodes adnexus (Hampe, 1863) Batrisodes delaporti (Aubé, 1833) Batrisodes venustus (Reichenbach, 1816) Tychobythinus glabratus (Rye, 1870) Bythinus burrelli Denny, 1825 Bythinus macropalpus Aubé, 1833 Bryaxis bulbifer (Reichenbach, 1816) Bryaxis curtisi (Leach, 1817) Bryaxis puncticollis (Denny, 1825) Trichonyx sulcicollis (Reichenbach, 1816) Amauronyx maerkeli (Aubé, 1844) Tychus niger (Paykull, 1800) Rybaxis longicornis (Leach, 1817) Brachygluta fossulata (Reichenbach, 1816) Brachygluta haematica (Reichenbach, 1816) Brachygluta waterhousei (Rye, 1876) Reichenbachia juncorum (Leach, 1817) Trissemus impressus (Panzer, 1803) Pselaphaulax dresdensis (Herbst, 1792) Pselaphus heisei Herbst, 1792 Claviger longicornis Müller, 1818 Claviger testaceus Preyssler, 1790 Phloeocharinae Phloeocharis subtilissima Mannerheim, 1830 Tachyporinae Sepedophilus bipunctatus (Gravenhorst, 1802) Sepedophilus constans (Fowler, 1888) Sepedophilus immaculatus (Stephens, 1832) Sepedophilus littoreus (Linnaeus, 1758) Sepedophilus lusitanicus Hammond, 1973 Sepedophilus marshami (Stephens, 1832) SUB VCH 19072005 ++ ++ ++ ++ ++ + + + Notes In rotten wood and under bark especially willow. Silwood Park 1998 (RGB) in FIT Local in litter and moss. Silwood Park 1998 (RGB) in FIT (N) Rare in rotting hardwoods. Silwood Park 1998 (RGB) in FIT (RDBIn) Rare in old woodland in rotting wood. Silwood Park 1998 (RGB) in FIT Local in red rotten oak and under bark. ? Downland. Silwood Park 1998 (RGB) in FIT Local in moss, litter under bark etc. ? [In manure heaps and vegetable refuse] (RDB3) In dead wood especially oak, under bark etc. . + (RDBK) ? (RDB1) In dead wood with the ant Lasius brunneus . Windsor Park (RDB1) In dead wood with the ant Lasius brunneus . Windsor Park 1985 (PSH) (Na) 2001 . Cothill (Collins) (RDBIn) Rare with ants (Ponera coarctata, and Lasius flavus?) on downland. ? [In moss in woodlands, also in grass tussocks] Bradfield (NHJ) Bradfield (NHJ) Local in wetlands including carr areas. Rare?. ? (RDB2) Rare in dead hardwoods. Silwood Park 1998 (RGB) in FIT (RDB3) Rare, with ants (Lasius fuliginosus, L. flavus, and Myrmica spp.) Local in compost heaps, grass tussocks etc. Local in wetlands. Local in litter and moss in wetlands including bogs and marshes. ? [In damp litter and wood]. (N) [In litter, mainly coastal] Local in litter and moss in wetlands including bogs and marshes. Widespread in marsh litter and moss by water. (N) [In moss and litter in wetlands]. Kennet Valley (NHJ) Local in moss in wetlands. (RDB1) ? In subterranean nests of the ants Lasius umbratus and L. mixtus. One to rediscover!! (N) Rare in nests of Lasius flavus. etc. + Widespread under bark and in bracket fungi especially Daedaleopsis on Salix. + + ++ ++ ++ ++ (Nb) Local in damp rotten wood including pine. (N) In fungoid wood and litter. Local in decaying vegetation and litter occasionally in dead wood. Local in dead and decaying wood. Rare in woodlands [especially in pine]. Widespread in litter in many habitats. ++ ++ ++ ++ * * ++ + + ++ + + ++ ++ + ++ + > + 23 Staphylinidae (Rove Beetles) species Sepedophilus nigripennis (Stephens, 1832) Sepedophilus pedicularius (Gravenhorst, 1802) Sepedophilus testaceus (Fabricius, 1792) Tachyporus atriceps Stephens, 1832 Tachyporus chrysomelinus (Linnaeus, 1758) Tachyporus dispar (Paykull, 1789) Tachyporus formosus Matthews, 1838 Tachyporus hypnorum (Fabricius, 1775) Tachyporus nitidulus (Fabricius, 1781) Tachyporus obtusus (Linnaeus, 1767) Tachyporus pallidus Sharp, 1871 Tachyporus pusillus Gravenhorst, 1806 Tachyporus scitulus Erichson, 1839 Tachyporus solutus Erichson, 1839 Tachyporus tersus Erichson, 1839 Tachyporus transversalis Gravenhorst, 1806 Lamprinodes saginatus (Gravenhorst, 1806) Tachinus bipustulatus (Fabricius, 1792) Tachinus elongatus Gyllenhal, 1810 Tachinus humeralis Gravenhorst, 1802 Tachinus laticollis Gravenhorst, 1802 Tachinus lignorum (Linnaeus, 1758) Tachinus marginellus (Fabricius,, 1781) Tachinus pallipes (Gravenhorst, 1806) Tachinus proximus Kraatz, 1855 Tachinus scapularis Stephens, 1832 Tachinus signatus Gravenhorst, 1802 Tachinus subterraneus (Linnaeus, 1758) Coproporus sp. Cilea siphoides (Linnaeus, 1767) Mycetoporus angularis Mulsant & Rey, 1853 Mycetoporus bimaculatus Lacordaire, 1835 Mycetoporus clavicornis (Stephens, 1832) Mycetoporus despectus Strand, 1969 Mycetoporus erichsonanus Fagel, 1965 Mycetoporus lepidus (Gravenhorst, 1802) Mycetoporus longulus Mannerheim, 1830 Mycetoporus nigricollis Stephens, 1835 Mycetoporus piceolus Rey, 1882 Mycetoporus punctus (Gravenhorst, 1806) Mycetoporus rufescens (Stephens, 1832) SUB VCH 19072005 ++ ++ + ++ ++ ++ +? ++ ++ ++ ++ ++ ++ > + + ++ ++ ++ + * ++ + + + + + + > + + Notes Local in litter etc. in a wide variety of habitats. (N) Rare, in wetland litter and flood debris. Kennet Floodplain 2003 (JSD) (N) Local in in rotting timber, especially beech. Local in moss and litter. Widespread and often abundant in many habitats including gardens. Widespread but confused with above species until recently. (Na) ? [Wetlands]. Widespread and abundant in most habitats. Widespread and abundant in most habitats. Widespread in most habitats especially damp areas. Local in litter in wetlands, river flood refuse etc. Widespread in moss and litter. (RDBK) ? [Habits unknown] . Local in litter and moss in a variety of open habitats. ? [Moss and litter in wetlands]. Rare in boggy wetlands Rare, myrmecophilous. (RDB1) [At sap on old trees ? [In moss and litter] Local in dung and rotting fungi etc. ? [In rotting grass, fungi, dung, ]. (N) Rare in dung [and sap and in fungi] Local [In decaying grass, dung and carrion]. (RDBIn) [In decaying vegetable matter, and carrion]. Common species in most habitats especially grasslands in dung and rotting vegetation and fungi. Widespread in leaf litter and other debris, also in fungi. Naturalized Windsor 19 (PMH) Local, associated with fermenting compost, and dung heaps. ? In litter and moss in woods? (RDBK) ? Woodland on chalk. ? Local in leaf litter etc. (N) ? [Woodlands] (N) [Moss in moorland and other open habitats]. In SUB as nanus. Widespread in litter in woods and grassland. ? Local in leaf litter etc. Rare? In moss and litter in woods. (N) [Dry grassland on chalky and sandy soils]. (N) [Eurytopic]. Rare? . 24 Staphylinidae (Rove Beetles) species Ischnosoma (Mycetoporus) longicornis Mäklin, 1847 Ischnosoma (Mycetoporus) splendidum (Gravenhorst, 1806) Lordithon exoletus (Erichson, 1839) Lordithon lunulatus (Linnaeus1761) Lordithon thoracicus (Fabricius, 1777) Lordithon trinotatus (Erichson, 1839) Bolitobius analis (Fabricius, 1787) Bolitobius cingulatus (Mannerheim, 1830) Bolitobius formosus (Gravenhorst, 1806) Parabolitobius inclinans (Gravenhorst, 1806) Trichophyinae Trichophya pilicornis (Gyllenhal, 1810) Habrocerinae Habrocerus capillaricornis (Gravenhorst, 1806) Aleocharinae Gymnusa brevicollis (Paykull, 1800) Deinopsis erosa (Stephens, 1832) Myllaena brevicornis (Matthews, A.H., 1838) Myllaena dubia (Gravenhorst, 1806) Myllaena elongata (Matthews, 1838) Myllaena gracilis (Matthews, A.H., 1838) Myllaena infuscata Kraatz, 1853 Myllaena intermedia Erichson, 1837 Myllaena kraatzi Sharp, 1871 Myllaena minuta (Gravenhorst, 1806) Oxypoda alternans (Gravenhorst, 1802) Oxypoda annularis Mannerheim, 1830 Oxypoda brachyptera (Stephens, 1832) Oxypoda elongatula Aubé, 1850 Oxypoda exoleta Erichson, 1839 Oxypoda ferruginea Erichson, 1839 Oxypoda flavicornis Kraatz, 1856 = amoena Fairmaire & Laboulbène, 1856 Oxypoda formiceticola Märkel, 1841 Oxypoda haemorrhoa (Mannerheim, 1830) Oxypoda induta Mulsant & Rey, 1861 Oxypoda lentula Erichson, 1837 Oxypoda lividipennis Mannerheim, 1830 SUB VCH 1907- Notes 2005 (N) [Woodlands in litter]. ++ Local. lists ++ ++ ++ ++ ++ + ++ Widespread in rotting fungi. Widespread in rotting fungi. Widespread in rotting fungi. Widespread in rotting fungi. Local, in decaying vegeatation, manure heaps, rotting fungi etc. Local in a wide variety of habitats, including woods, and grassland ? [In litter] Rare? In leaf litter in woods etc. + (Nb) ? Woodlands.. + Widespread in leaf litter and moss in woodlands. + Rare in Sphagnum and other wetland mosses beside bog pools and weedy ponds. Local in litter beside streams ditche and ponds. Tolerant of heavy shading Local in wetlands Local in litter in wetlands. Kennet (JSD) (N) Local on exposed sediments beside streams and rivers. Shade tolerant. Holybrook 2003 (JSD) Rare in wetlands. Local in litter in wetlands Local in litter in wetlands (N) Rare in Sphagnum mires. Local in wetlands in marsh litter and moss Widespread on rotting fungi in wooded areas. ? [In decaying vegetation] Rare? in garden Widespread in litter in marshes and wet woodland areas. (N) Rare, eurytopic. ? lists. , (N) Rare in fungoid leaf litter in old woods. + ++ Local in wood ant Formica rufa nests Rare in nests of wood ants Formica rufa + ++ ++ Rare. Rare in wet moss. Local in carrion and decaying litter + + + ++ ++ + + ++ + + ++ ++ ++ + 25 Oxypoda lurida Wollaston, 1857 Oxypoda nigrocincta Mulsant & Rey, 1875 Oxypoda opaca (Gravenhorst, 1802) Oxypoda recondita Kraatz, 1856 Oxypoda sericea Heer, 1839 Oxypoda spectabilis Märkel,, 1844 Oxypoda tarda Sharp, 1871 Oxypoda umbrata (Gyllenhal, 1810) Oxypoda vittata Märkel, 1842 Ocyusa maura (Erichson, 1837) Ocyusa (=Deubelia) picina (Aubé, 1850) Calodera aethiops (Gravenhorst, 1802) Calodera nigrita Mannerheim, 1830 Calodera riparia Erichson, 1837 Parocyusa (=Chiloporata) longitarsis (Erichson, 1837) Ischnoglossa prolixa (Gravenhorst, 1802) Ischnoglossa turcica Wunderle, 1992 Dexiogyia corticina (Erichson, 1837) Homoeusa acuminata (Märkel, 1842) Thiasophila angulata (Erichson, 1837) Thiasophila inquilina (Märkel,, 1842) Crataraea suturalis (Mannerheim, 1830) Haploglossa gentilis (Märkel, 1844) Haploglossa marginalis (Gravenhorst, 1806) Haploglossa nidicola (Fairmaire, 1852) Haploglossa pulla (Gyllenhal, 1827) Mniusa incrassata (Mulsant & Rey, 1852) Ocalea badia Erichson, 1837 Ocalea picata (Stephens, 1832) Ocalea rivularis Miller, L., 1851 Ilyobates bennetti Donisthorpe, 1914 Ilyobates nigricollis (Paykull, 1800) Ilyobates propinquus (Aubé, 1850) Amarochara forticornis (Boisduval & Lacordaire, 1835 Amarochara umbrosa (Erichson, 1837) Phloeopora bernhaueri Lohse 1984 = teres auct. Brit.partim? not (Gravenhorst, 1802) Phloeopora testacea (Mannerheim, 1830) Dinarda dentata (Gravenhorst, 1806) Dinarda maerkeli Kiesenwetter, 1843 Meotica apicalis Benick, 1954 Meotica exilis (Gravenhorst, 1806) * + + + + + + + ++ ++ + ++ + + + * + ++ + ? (N) ? as perplexa Mulsant (RDBIn) Windsor 1937 (HSD) Local in litter and tussocks in woods etc. (N) Rare, in hollow trees esp. with ants. ? In litter, grass heaps etc.? (N) ? [Mole nests] ? (P.Whitehead) Local in litter etc. in woods. ? Downland Rare with the ant Lasius fuliginosus Local in wetlands, adults often winter in Typha stems Local in wetlands Rare in fens and wetlands. (N) [In wetlands] (N) [In wetlands] last recorded in 1909 Local in wetlands Hutchins Copse (JSD) Rare, under bark. Rare, in rotting wood and under bark. (N) Rare under bark in old woodlands. (RDBK) ? With ants on downland. Rare with wood ant Formica rufa and Lasius fuliginosus etc. (N) ? With Lasius fuliginosus, last recorded at in 1914 Local in decaying vegetation esp. grass heaps. Rare? In bird’s nests and with Lasius fuliginosus. (N) Rare in hollow trees in old nests. lists + + + + + ++ * Rare? In ant and bird’s nests. ? Open sandy ground? ? Open sandy grassland. Local in moss and litter in damp areas. ? woodland. Rare. Sandy river-banks. Widespread, in grasslands, and woods. (RDBK) Rare in litter, mammal runs in woods. (N) Rare. Sandy ground near water [by rivers and in sandpits] (RDBK) ? [Subterranean] + ++ (N) [Subterranean in mammal runs] 1908 Rare? Under bark Hitchcopse Pit 1994 (DC) ++ + + + + Local under bark of hardwoods. (N) V.local, but present in most nests of the Slave-Making Ant Formica sanguinea on heaths and commons. Rare. In nests of Wood Ants Formica rufa. ? , ? [Eurytopic in litter and moss] + 26 Meotica exillima Sharp, 1915 Ischnopoda (=Tachyusa) atra (Gravenhorst, 1806) Ischnopoda (=Tachyusa) coarctata Erichson, 1837 Ischnopoda (=Tachyusa) constricta Erichson, 1837 Ischnopoda (=Tachyusa) leucopus (Marsham, 1802) Ischnopoda (=Tachyusa) umbratica Erichson, 1837 Gnypeta carbonaria (Mannerheim, 1830) Gnypeta ripicola (Kiesenwetter, 1844) Gnypeta rubrior Tottenham, 1939 Dasygnypeta (=Gnypeta) velata (Erichson, 1837) Brachyusa concolor (Erichson, 1839) Dacrila fallax (Kraatz, 1855) Callicerus rigidicornis (Erichson, 1839) Callicerus obscurus Gravenhorst, 1802 Schistoglossa gemina (Erichson, 1837) Boreophilia eremita (Rye, 1866) Aloconota gregaria (Erichson, 1839) Aloconota insecta (Thomson, 1856) Aloconota languida (Erichson, 1837) Aloconota longicollis (Mulsant & Rey, 1852) Aloconota sulcifrons (Stephens, 1832) Pycnota paradoxa (Mulsant & Rey, 1861) Amischa analis (Gravenhorst, 1802) Amischa decipiens (Sharp, 1869) Amischa forcipata Mulsant & Rey, 1873 Amischa nigrofusca (Stephens, 1829) =soror (Kraatz, 1856) Nehemitropia lividipennis (Mannerheim, 1830) = sordida (Marsham, 1802) Notothecta confusa (Markel, 1844) Notothecta flavipes (Gravenhorst, 1806) Lyprocorrhe anceps (Erichson, 1837) Alaobia (Atheta) gagatina (Baudi, 1848) Alaobia (Atheta) pallidicornis (Thomson, 1856) Alaobia (Atheta) scapularis (Sahlberg, 1831) Alaobia (Atheta) sodalis (Erichson, 1837) Alaobia (Atheta) taxiceroides Munster, 1932 Alaobia (Atheta) trinotata (Kraatz, 1856) Dochmonota clancula (Erichson, 1837) Ousipalia caesula (Erichson, 1839) + ++ Very local in Sphagnum in mires and mossy carr. ? Bare margins of water Green park 2002 (JSD) ++ (N) Local on bare ground near water. R.Kennet, Burghfield Bridge 2003 (JSD) ++ Local on bare sand and silt at margins of rivers and streams. R.Kennet, Burghfield Bridge 2003 (JSD) ++ Local on bare sand and silt at margins of rivers. R.Kennet, Burghfield Bridge 2003 (JSD) + Local on bare sand and silt at margins of rivers. , + + + ++ Rare on bare ground at margins of water (N) V.local on exposed sedimenst by flowing water. etc. V.local, on exposed sedimenst by flowing water. etc. (N) Recent immigrant, now locally common at margins of ponds etc. + + ++ ++ + + (N) Local at margins of rivers and ponds. ? [In wetlands] Rare in damp woods. Rare? In damp woods often near streams. (N) Rare wetland species. Rare. [Mainly an Widespread, eurytopic in litter and moss Rare? On exposed sediments beside rivers. (N) [Wetlands] (N) Rare in litter in marshy areas. ? [Riparian] In (N)? [Mole nests] 1907 Widespread and often abundant in litter and moss including compost heaps Widespread in litter and moss including compost heaps Rare? in damp litter. Local in litter + Local in grass heaps + + + + + + > ++ ++ (N) ? Hollow trees and with the ant Lasius fuliginosus. Local in nests of wood ants Formica rufa. Tubney 1908 Local in nests of wood ants Formica rufa. Tubney 1908 V.local in moss, litter and under bark. ? Woodland? (N) [Moss on downland] Local in litter in woods etc. Rare in nests of birds & mammals in hollow trees. . ? Woodland? Wytham Wood 1990 (V.Gusarov) Sheffield Bottom 2003 (JSD) (N) Rare in wetlands. Dinton Pastures 2003 (JSD) ? lists. + ++ + > ++ + + 27 Geostiba circellaris (Gravenhorst, 1806) Dinaraea aequata (Erichson, 1837) Dinaraea angustula (Gyllenhal, 1810) Dinaraea linearis (Gravenhorst, 1802) Paranopleta inhabilis (Kraatz, 1856) Dadobia immersa (Erichson, 1837) Plataraea brunnea (Fabricius, 1798) Liogluta longiuscula (Gravenhorst, 1802) Liogluta nitidula (Kraatz, 1856) Liogluta oblongiuscula (Sharp, 1869) Liogluta pagana (Erichson, 1839) Philhygra (Atheta) debilis (Erichson, 1837) Philhygra (Atheta) deformis (Kraatz, 1856) Philhygra (Atheta) elongatula (Gravenhorst, 1802) Philhygra (Atheta) fallaciosa (Sharp, 1869) Philhygra (Atheta) gyllenhali (Thomson, 1856) Philhygra (Atheta) hygrobia (Thomson, 1856) Philhygra (Atheta) hygrotopora (Kraatz, 1856) Philhygra (Atheta) luridipennis (Mannerheim, 1830) Philhygra (Atheta) malleus Joy, 1913 Philhygra (Atheta) nannion Joy, 1931 Philhygra (Atheta) obtusangula Joy, 1913 Philhygra (Atheta) palustris (Kiesenwetter, 1844) Philhygra (Atheta) volans (Scriba, 1859) Dilacra (Atheta) luteipes (Erichson, 1837) Dilacra (Atheta) vilis (Erichson, 1837) Parameotica (Atheta) difficilis (Brisout, 1860) Enaladroma hepatica (Erichson, 1839) Bessobia (Atheta) excellens (Kraatz, 1856) Bessobia (Atheta) fungivora (Thomson, 1867) Bessobia (Atheta) monticola (Thomson, 1852) Bessobia (Atheta) occulta (Erichson, 1837) Anopleta (Atheta) corvina (Thomson, 1856) Anopleta (Atheta) puberula (Sharp, 1869) Anopleta (Atheta) soedermani Bernhauer, 1931 Microdota (Atheta) aegra (Heer, 1841) Microdota (Atheta) amicula (Stephens, 1832) Microdota (Atheta) atomaria (Kraatz, 1856) Microdota (Atheta) atricolor (Sharp, 1869) Microdota (Atheta) benickiella (Brundin, 1948) Microdota (Atheta) boreella (Brundin, 1948) Microdota (Atheta) glabricula (Thomson, 1867) ++ ++ ++ + * + ++ + + ++ ++ ++ Widespread in litter and debris in a variety of habitats including gardens. Widespread on dead wood and under bark esp. on moribund attached boughs of hardwoods esp. oak Rare under bark, and at tree roots Local under bark and in decaying hardwoods. (RDBK) Rare. Local under pine bark. Widespread in litter esp. in woodlands ? Wytham Rare in nests of the ant Lasius fuliginosus [and L.brunneus] Rare in woodlands? (N) ? [In woods, parks and wetlands, probably associated with small mammal runs] Wytham Wood 1990 (V.Gusarov) Rare? In litter in wetlands. lists (N) [In moss in wetlands] Local in wetlands in litter and moss. + + + + Rare? [Wetlands] Rare in wetlands. (N) Rare [In wetlands esp. carr] Rare in litter and moss in wetland areas. Rare [exposed sediments near water] ++ * + + > + + + + + + + + * ? + ++ + + + + Local in litter in marshes, and river-banks. R.Kennet (JSD) Rare in wet areas. Rare in wetlands ? In wetlands ? [Riparian] ? Rare in marshes and carr (N) Rare? in wetlands Widespread in leaf litter etc. in deciduous woodland. Rare in woodlands? ? ? In interception trap in woodland at Local in litter and decaying vegetation in woods and gardens Local in fungi in woodlands. (RDBK) ? [Subterranean?] (N) [In dung, carrion and litter in a variety of open habitats] lists (N) Rare. Leaf litter on downland, possibly subterranean. ? In woodlands [in birch brackets] Wytham Wood 1990 (V.Gusarov) ? [In litter] lists Rare? [Moss and haystack refuse] (N) Local? Recorded from 3 sites by ? (N) ? [Woodlands] 28 Microdota (Atheta) indubia (Sharp, 1869) Microdota (Atheta) inquinula (Gravenhorst, 1802) Microdota (Atheta) liliputana (Brisout, 1860) Microdota (Atheta) minuscula (Brisout, 1860) Microdota (Atheta) palleola (Erichson,1837) Microdota (Atheta) subtilis (Scriba, 1866) Pachyatheta cribrata (Kraatz, 1856) Pachyatheta mortuorum Thomson, 1867 Ceritaxa (Atheta) pervagata (Benick, 1974) =dilaticornis auct. not (Kraatz, 1856) Ceritaxa (Atheta) testaceipes (Heer, 1839) Cadaverota (Atheta) cadaverina (Brisout, 1860) Mocyta (Atheta) amplicollis (Mulsant & Rey, 1873) Mocyta (Atheta) clientula (Erichson, 1839) Mocyta (Atheta) fungi (Gravenhorst, 1806) Mocyta (Atheta) gilvicollis Scheerpeltz Mocyta (Atheta) negligens Mulsant & Rey Mocyta (Atheta) orbata (Erichson, 1837) Mocyta (Atheta) orphana (Erichson, 1837) Rhagocneme subsinuata (Erichson, 1839) Datomicra canescens (Sharp, 1869) Datomicra celata (Erichson, 1837) Datomicra dadopora Thomson, 1867 Datomicra nigra (Kraatz, 1856) Datomicra sordidula (Erichson, 1837) Datomicra zosterae (Thomson, 1856) Atheta aquatica (Thomson, 1852) Atheta castanoptera (Mannerheim, 1830) Atheta graminicola (Gravenhorst, 1806) Atheta hypnorum (Kiesenwetter, 1850 Atheta triangulum (Kraatz, 1856) Atheta xanthopus (Thomson, 1856) Atheta basicornis (Mulsant & Rey, 1852) Atheta britanniae (Bernhauer & Scheerpeltz, 1926) Atheta coriaria (Kraatz, 1856) Atheta crassicornis (Fabricius, 1792) Atheta (type III) divisa (Märkel, 1844) Atheta euryptera (Stephens, 1832) Atheta fungicola (Thomson, 1852) Atheta harwoodi Williams, 1930 Atheta intermedia (Thomson, 1852) Atheta liturata (Stephens, 1832) ? ++ ++ ? Woods and ? Wytham Wood 1990 (V.Gusarov) (N) ? [Grassheaps, dung etc. in woods and gardens] Silwood Park 1998-9 (RGB) in FIT + ++ * + * Rare? ? Wytham Wood 1990 (V.Gusarov) (RDBIn) [Woodland in decaying fungi] ? in (AMM) coll. (N) ? [Woodlands in litter] (RDBK) and Tubney 1908 (JJW) (RDBK) Woodland in fungi. + + + (RDBK) [In fungi, moss etc on dry soils] Tubney Local, eurytopic in litter etc. . lists Widespread. . lists + + + + + * + + + + + + + ++ + ++ + + > ++ + ? In litter and moss? Very common in most habitats. ? . Taxonomically problematic species. ? 1995-7 Taxonomically problematic species. ? [In litter] lists (N) ? Local in litter and grass heaps. Local, in dung etc. ? ? Widespread in litter, grassheaps etc. ?. (N) [Wetlands in moss and litter] Local in leaf litter. Local in fungi. Widespread in marshland in litter and tussocks. Local in litter esp. in woods. ? [In damp litter] ? In litter. lists (N) Rare [in wet woodland]. ? Woodland. . + ++ + ? Woodland. . Local in litter ? Woods and downland? ? [In haystacks, moss etc.] ? lists Downland? ? (Fowler, 1888) Local on fungi on wood. + ++ 29 Atheta nidicola (Johansen, 1914) Atheta nigricornis (Thomson, 1852) Atheta nigritula (Gravenhorst, 1802) Atheta oblita (Erichson, 1839) Atheta paracrassicornis Brundin Atheta pertyi (Heer, 1839) ++ + > ?? + 1977 . Kennet 2003 (JSD) Local, in old bird nests and squirrel dreys. Usually present when looked for!! (N) [Woods and heaths in fungi, and other decaying debris] ? in grassheaps. Wytham Wood 1990 (V.Gusarov) Rare in rotting fungi. Atheta pilicornis (Thomson, 1852) Atheta ravilla (Erichson, 1839) Mycetota (Atheta) laticollis (Stephens, 1832) Dimetrota atramentaria (Gyllenhal, 1810) Dimetrota cauta (Erichson, 1837) Dimetrota cinnamoptera (Thomson, 1856) Dimetrota ischnocera Thomson, 1870 Dimetrota laevana (Mulsant & Rey, 1852) Dimetrota marcida (Erichson, 1837) Dimetrota nigripes (Thomson, 1856) Dimetrota setigera (Sharp, 1869) Badura (=Dimetrota) macrocera (Thomson, 1856) Chaetida longicornis (Gravenhorst, 1802) Coprothassa melanaria (Mannerheim, 1830) Acrotona aterrima (Gravenhorst, 1802) Acrotona benicki Allen, 1940 Acrotona consanguinea (Eppelsheim, 1875) Acrotona muscorum (Brisout, 1860) Acrotona parens (Mulsant & Rey, 1852) Acrotona parvula (Mannerheim, 1830) Acrotona pygmaea (Gravenhorst, 1802) Alevonota aurantiaca Fauvel, 1895 Alevonota egraria (Rye, 1876) Alevonota gracilenta (Erichson, 1839) Alevonota rufotestacea (Kraatz, 1856 Alianta incana (Erichson, 1837) Pachnida nigella (Erichson, 1837) Trichiusa immigrata Lohse, 1984 Thamiaraea cinnamomea (Gravenhorst, 1802) Thamiaraea hospita (Märkel,, 1844) Aleochara bilineata Gyllenhal, 1810 Aleochara bipustulata (L.innaeus,, 1761) Aleochara brevipennis Gravenhorst, 1806 Aleochara cuniculorum Kraatz, 1858 Aleochara curtula (Goeze, 1777) Aleochara discipennis Mulsant & Rey, 1853 + + ++ ++ + + + + + ++ ++ (N) Rare in rotting wood and under bark. ? Downland? ? Downland? [Wetlands in litter] ? Woodland. Wytham Wood 1990 (V.Gusarov) ? Woodland. ? Downland. ? Woodland. ? Woodland. ? Woodland. ? Wytham Wood 1990 (V.Gusarov) ? Wytham Wood 1990 (V.Gusarov) + + + + + ? Downland. . ? ? Downland. (RDBK) (RDBK) [Woodland in hollow trees and with the ant Lasius umbratus] lists (N) ? Silwood Park 1998 (RGB) in FIT ? Downland ? lists (RDBIn) Rare downland, including a garden on the . (RDBIn) [Woods, probably subterranean in mammal nests] (RDBK) ? Downland (N) Woods, hedgebanks probably subterranean. Silwood Park 1998-9 (RGB) in FIT Widespread in wetlands and pond margins in litter, winters in Typha stems. Rare in wetland in marsh litter. Recent immigrant increasing, Rare at goat moth Cossus trees. (N) Rare in woodlands [At goat moth Cossus trees] Rare. [In dung and vegetable debris] Hitchcopse Pit 1996 (TDH) Local in dung, carrion and rotting vegetable matter (N) Local, in wetland areas. Taken in interception trap at Rare in vicinity of rabbit warrens [& badger setts]. Taken in interception trap at Widespread in carrion and in rotting fungi. (N) Rare, in dung esp. horse and in rotten wood.. ++ + + + ++ + + + + + ++ ++ ++ + ++ * 30 Aleochara fumata (Gravenhorst, 1802) Aleochara funebris Wollaston, 1864 Aleochara intricata Mannerheim, 1830 Aleochara kamila Livossky, 1965 Aleochara lanuginosa Gravenhorst, 1802 Aleochara lata Gravenhorst, 1802 Aleochara lygaea Kraatz, 1862 Aleochara maculata Brisout, 1863 Aleochara moerens Gyllenhal, 1827 Aleochara moesta Gravenhorst, 1802 Aleochara obscurella Gravenhorst Aleochara ruficornis Gravenhorst, 1802 Aleochara sanguinea (Linnaeus, 1758) Aleochara spadicea (Erichson, 1837) Aleochara sparsa Heer, 1839 Aleochara stichai Likovsky, 1965 Aleochara tristis Gravenhorst, 1806 Aleochara verna Say, 1839 Aleochara villosa Mannerheim, 1830 Tinotus morion (Gravenhorst, 1802) Drusilla canaliculata (Fabricius, 1787) Zyras cognatus (Märkel, 1842) Zyras collaris (Paykull, 1789) Zyras funestus (Gravenhorst, 1806) Zyras haworthi Stephens, 1835 Zyras humeralis (Gravenhorst, 1802) Zyras laticollis (Märkel, 1842) Zyras limbatus (Paykull, 1789) Zyras lugens (Gravenhorst, 1802) Lomechusoides strumosus (Fabricius, 1792) Lomechusa emarginata (Paykull, 1789) Cordalia obscura (Gravenhorst, 1802) Myrmecocephalus concinnus Erichson, 1839 Falagrioma (Falagria) thoracica Stephens, 1832 Falagria caesa Erichson, 1837 Falagria sulcatula (Gravenhorst, 1806) Autalia impressa (Olivier, 1795) Autalia longicornis Scheerpeltz, 1947 Autalia rivularis (Gravenhorst, 1802) Encephalus complicans Stephens, 1832 Gyrophaena affinis Mannerheim, 1830 Gyrophaena angustata (Stephens, 1832) Gyrophaena bihamata Thomson, 1867 Gyrophaena congrua Erichson, 1837 Gyrophaena fasciata (Marsham, 1802) + + ++ + + ++ * * * * + + + > ++ + ++ ++ + + + + + + ++ + > + + + ++ + ++ + ++ + ++ ++ ++ + (RDBK) Wytham Wood 1909 Rare? Woods, parkland, downland? . Rare? In herbivore dung esp. horse. Wytham Wood 1990 (V.Gusarov) (N) a diversa Rare in wood/parkland on bracket fungi and in hollow trees etc. Widespread in herbivore dung esp. cow, and carrion and on fungi. ? Rare? [In carrion etc.] (RDBIn) [In carrion, dung etc.] (RDBIn) [Woodlands in litter and moss] (N) Extinct? (RDBK) Windsor 1927-34 ? (N) ? [Elusive, and can be abundant in pitfall traps] (N) [Woodlands in litter, dung, fungi and birds nests] Rare (under-recorded?) subterranean in mole nests Local? Woods, and . (N) ? [In dung and carrion] Rare in dung and carrion. (RDBK) Local. Recently separated species. Hutchins Copse 1998 (JSD) (RDBK) Local in dung and carrion Common in open habitats usually near to nests of Myrmica (RDBK) Rare with the ant Lasius fuliginosus in runs and nests. Rare [With Myrmica ants] 1907 Rare with the ant Lasius fuliginosus in runs and nests. (Nb) Rare in woodlands sometimes with Lasius fuliginosus. Aldermaston 1988 (S.Grove) Local, over-looked as it can be frequent in pitfall samples in a wide variety of open habitats. Rare with the ant Lasius fuliginosus in runs and nests. Rare with the ant Lasius fuliginosus in runs and nests. (N) Rare with the ant Lasius fuliginosus in runs and nests. (RDBIn) Rare, in nests of Formica sanguinea. Bagshott (JP) (N) Rare, with ants Myrmica etc. on heaths and downland Rare [in litter in marshy areas] Local in fungi on dead wood, and in compost heaps etc. Rare in damp litter in marshy areas Local in decaying grass etc. ? [Marshland] lists, Walton Widespread and frequnet on rotting fungi. Local on rotting fungi Local in decaying grass etc. Rare in damp areas [Flushes] Local on bracket fungi on trees. Sulhamstead 2003 (JSD) (N) Rare on bracket fungi esp. Polyporus squamosus. Sulhamstead 2003 (JSD) Local on bracket fungi esp. Polyporus squamosus. Sulhamstead 2003 (JSD) (N) Rare on bracket fungi. , , (Fowler, 1888) Local on bracket fungi 31 Gyrophaena gentilis Erichson, 1839 Gyrophaena hanseni Strand, 1946 Gyrophaena joyi Wendeler, 1924 Gyrophaena joyioides Wuesthoff, 1937 Gyrophaena minima Erichson, 1837 Gyrophaena nana (Paykull, 1800) Gyrophaena poweri Crotch, 1866 Gyrophaena pulchella Heer, 1839 Gyrophaena strictula Erichson, 1839 Gyrophaena williamsi Strand, 1935 Agaricochara (=Gyrophaena) latissima (Stephens, 1832) Homalota plana (Gyllenhal, 1810) Anomognathus cuspidatus (Erichson, 1839) Silusa rubiginosa Erichson, 1837 Leptusa fumida Kraatz, 1839 Leptusa norvegica Strand, 1941 Leptusa pulchella (Mannerheim, 1830) Leptusa ruficollis (Erichson, 1839) Euryusa optabilis Heer, 1839 Euryusa sinuata Erichson, 1837 * Tachyusida gracilis (Erichson, 1837) Bolitochara bella Märkel, 1844 Bolitochara lucida (Gravenhorst, 1802) Bolitochara obliqua Erichson, 1837 Placusa complanata Erichson, 1846 Placusa depressa Maeklin, 1845 Placusa pumilio (Gravenhorst, 1802) Placusa tachyporoides (Waltl, 1838) Hygronoma dimidiata (Gravenhorst, 1806) Cypha discoidea (Erichson, 1839) Cypha hanseni (Palm, 1949) Cypha longicornis (Paykull, 1800) Cypha ovulum (Heer, 1839) Cypha pulicaria (Erichson, 1839) Cypha punctum (Motschulsky, 1857) Cypah seminulum ?? Olobus (=Oligota) apicata Erichson, 1837 Olobus (=Oligota) flavicornis (Lacordaire, 1835) Oligota granaria Erichson, 1837 Oligota inflata (Mannerheim, 1830) Oligota parva Kraatz, 1862 Oligota picipes (Stephens, 1832) Oligota pumilio Kiesenwetter, 1858 ++ + + ++ ++ * + + + ++ ++ ++ + ++ ++ ++ ++ ++ + ++ ++ + + + + + ++ ++ + ++ * + + 195 1 + + + + + Local in bracket fungi. 1987 Local on fungi esp. sulphur tuft Hypholoma fasciculare and other yellow gill fungi. (N) [On fungi in wet woods] 1963 (N) Rare on fungi in woods including Polyporus squamosus. Sulhamstead 2003 (JSD) Widespread in yellow gill fungi esp. sulphur tuft in woodlands ? [In fungi], , , (RDBK) [In wood rotting fungi] (RDBK) Rare in fungi 1914 (N) Rare in maze-gill Daedalea quercina on oak & sweet chestnut. 1967 Rare. [In woodland and carr on fungi] Local, but widespread in fungi. Local, under bark of beech. Local, under fungoid bark (N) Rare at sap and under bark. Widespread on dead and moribund hardwoods esp. oak boughs. (N) [Under bark] Rare under bark of trees. Widespread on dead and moribund hardwoods (RDBIn) [Old woodlands in decaying trees often with ants] (RDBIn) Rare, in frass in nests of Lasius brunneus. Windsor park Silwood Park Windsor Widespread in fungi in woodlands Local on fleshy fungi on dead wood. Local under bark and in soft bracket fungi Recent colonist. Rare?. (N) In burrows of Tomicus piniperda on pine. Local under oak bark. (N) Rare at fresh sap of hardwood Local amongst tall emergent vegetation in wetlands, (N) Rare in dry litter ‘suspended’ off the ground in reed beds. Virginia Water 1998, Searles lane 2003 (JSD) Local in woodlands. Widespread in litter, grass heaps etc. in wild and man-made habitats. (RDBK) [In moss] (N) [Wetlands] 1961 ? 1999 (N) Local on soft bracket fungi. (N) ? Woods and parks. , ? 1910 Widespread in grass and compost heaps. Local in compost heaps etc. In SUB as O. atomaria (Gravenhorst, 1806) Local in compost heaps etc. 32 Oligota punctulata Heer, 1839 Oligota pusillima (Gravenhorst, 1806) Scaphidiinae (shining fungus beetles) Scaphidium quadrimaculatum Olivier, 1790 Scaphisoma agaricinum (Linnaeus, 1758) Scaphisoma boleti (Panzer, 1793) Piestinae Siagonum quadricorne Kirby, 1815 Oxytelinae Deleaster dichrous (Gravenhorst, 1802) Syntomium aeneum (Müller, 1821) Coprophilus striatulus (Fabricius, 1792) Manda mandibularis (Gyllenhal, 1827) Planeustomus palpalis (Erichson, 1839) Aploderus caelatus (Gravenhorst, 1802) Bledius dissimilis Erichson, 1840 Bledius femoralis (Gyllenhal, 1827) Bledius fergussoni Joy, 1912 Bledius gallicus (Gravenhorst, 1806) Bledius limicola Tottenham, 1940 =germanicus Wagner, 1935 Bledius longulus Erichson, 1839 Bledius opacus (Block, 1799) Bledius pallipes (Gravenhorst, 1806) Bledius subterraneus Erichson, 1839 Bledius terebrans (Schioedte, 1866) Bledius unicornis (Germar, 1825) Ochthephilus angustior (Bernhauer, 1943) = venustulus auct Brit. not (Rosenhaeur, 1856) Ochthephilus aureus (Fauvel, 1871) Carpelimus bilineatus Stephens, 1834 Carpelimus corticinus (Gravenhorst, 1806) Carpelimus elongatulus (Erichson, 1839) Carpelimus fuliginosus (Gravenhorst, 1802) Carpelimus gracilis (Mannerheim, 1830) Carpelimus impressus (Boisduval & Lacordaire, 1835) Carpelimus lindrothi Palm, 1942 Carpelimus obesus (Kiesenwetter, 1844) Carpelimus pusillus (Gravenhorst, 1802) Carpelimus rivularis (Motschulsky, 1860) Carpelimus subtilicornis (Roubal, 1946) Carpelimus subtilis (Erichson, 1839) ++ + SUB Rom VCH 1907 2005 + + + + + + + + LRom ++ * ++ + + ++ + + ESax Widespread, 3 sites ? 1907 Notes Local in dead wood of many kinds, avery distinctive species, >40 post 1980 records Local in dead wood of many kinds. Often found in between stacked slices of sawn up trunks (Nb) Rare, in dead wood of many kinds. Local under bark. Especially on elm attacked by Scolytus. (Nb) V.local on banks of streams and rivers. Rare in moss in woods. Local in rotting vegetation including compost heaps. (RDB1) Rare, pools in woodlands. (RDBK) Wet woodlands ? [In dung etc] (RDBIn) Sheffield Bottom 2003 (JSD) (Na) Rare on bare ground at margins of water. Rare in flooded sandpit. Local on sandy margins of , pond margins as at 1997 Rare on bare ground near water. s ? [Bare ground near water] ? [Bare ground near water] and at MV light Theale Lake 2003 (JSD) Local on sandy margins of rivers. Local on sandy margins of rivers esp. (RDBK) 1898 & 1927 ? in Fowler (1888) (N) ? + ++ ++ ++ + + + Rare on stoney margins of streams, shingle bars etc. Widespread on ground beside standing and flowing water Local on ground beside standing and flowing water Local in litter in marshy margins of water. (N) [Wetlands in tussocks etc.] ? Muddy pond margins. ? Wet areas, pond margins. + + + + > > (N) Rare in wetlands and pond margins. (N) Rare on muddy drawdown zone of ponds Local esp. near water, but also taken in gardens. Local on silt and sand, at margins of running water. Local on banks of rivers. (N) Damp open sandy ground. 33 Carpelimus zealandicus (Sharp, 1900) Thinodromus arcuatus (Stephens, 1834) Platystethus alutaceus Thomson, 1861 Platystethus arenarius (Fourcroy, 1785) Platystethus capito Heer, 1839 Platystethus cornutus (Gravenhorst, 1802) Platystethus degener Mulsant & Rey, 1878 Platystethus nitens (Sahlberg, 1832) Platystethus nodifrons Mannerheim, 1830 Anotylus clypeonitens (Pandelle, 1867) Anotylus complanatus (Erichson, 1839) Anotylus hamatus (Fairmaire & Laboulbène, 1856) Anotylus insecatus (Gravenhorst, 1806) Anotylus inustus (Gravenhorst, 1806) Anotylus mutator (Lohse, 1963) Anotylus nitidulus (Gravenhorst, 1802) Anotylus rugosus (Fabricius, 1775) Anotylus saulcyi (Pandelle, 1867) Anotylus sculpturatus (Gravenhorst, 1806) Anotylus tetracarinatus (Block, 1799) Oxytelus laqueatus (Marsham, 1802) Oxytelus migrator Fauvel Oxytelus sculptus Gravenhorst, 1806 Oxyporinae Oxyporus rufus (Linnaeus, 1758) Steninae (Camphor beetles) Stenus aceris Stephens, 1833 Stenus argus Gravenhorst, 1806 Stenus ater Mannerheim, 1830 Stenus atratulus Erichson, 1839 Stenus bifoveolatus Gyllenhal, 1827 Stenus biguttatus (Linnaeus, 1758) Stenus bimaculatus Gyllenhal, 1810 Stenus binotatus Ljungh, 1804 Stenus boops Ljungh, 1810 Stenus brevipennis Thomson, 1851 Stenus brunnipes Stephens, 1833 Stenus butrintensis Smetana, 1959 Stenus canaliculatus Gyllenhal, 1827 Stenus canescens Rosenhauer, 1856 Stenus carbonarius Gyllenhal, 1827 Stenus cicindeloides (Schaller, 1783) Stenus circularis Gravenhorst, 1802 Stenus clavicornis (Scopoli, 1763) R/Sax Rom R/Sax LRom LRom BrAge Rom/S ax Ro/sa x LRom + + + ++ + ++ + + > > + + Local in open situations. Local on exposed sediments by streams, shingle bars etc. Local in damp silt and sand at margins of still and flowing water. Widespread in dung and rotting vegetation Rare, on sparsely vegetated open chalk grassland. Widespread. Burrowing in sediment near water, esp. in sand and clay pits. Local in organically enriched mud by water Local on damp sediment or litter on margins of ponds and rivers. (N) Rare in marshy localities. ? In compost, and litter. Local in rotting vegetation (N) Rare [In dung in grasslands and woods] + ++ ++ (N) ? [In litter, dung and other decaying vegtable matter] Widespread in herbivore dung and rotting plant debris. (N) ? ? Rare? Extinct, much declined in carrion dung etc. Common on ground in most habitats, and a strong flyer + ++ (N) Rare? in subterranean mammal runs (mole, badger, rabbit). Widespread in dung and rotting vegetation ++ ++ + + + Common on dung and rotting vegetation. Local in herbivore dung Naturalised. Increasing, ? In woods [Also near water] Rare and declining in rotting fungi. + Local in litter in grasslands, not a wetland species. (Nb) [Marshes, fens and other rich wetlands] (Nb) [A dryland species of open habitats] (Nb) [Wetland species] (J. Linnell) Local in wetlands in moss and marsh litter (Should be Na) [Exposed sediments beside rivers and streams] Local in shaded marshy ground amongst emergent vegetation Local in wetlands amongst tall emergent vegetation Local on exposed sediments by ponds and flowing water. Rare, in mires and bogs. , . Common in dry grasslands areas (N) Local in tall emergent waterside vegetation. Winters in Typha stems. Local. Margins of still and running water, often on exposed sediments. (Nb) Rare, at marshy margins of ponds and streams. [Fens, and mesotrophic wetlands] Common in richly vegetated wetlands, river margins etc. A strong flier. (Nb) Rare, most often found in flood refuse. Kennet valley, Southcote 2003 (JSD) Widespread in grassy habitats. A dry land species, and a strong flier ++ ++ + ++ > ++ ++ ++ + ++ ++ ++ 34 Stenus comma LeConte, 1863 Stenus contumax Assing, 1994 Stenus crassus Stephens, 1833 Stenus exiguus Erichson, 1840 Stenus europaeus Puthz, 1966 Stenus flavipes Stephens, 1833 Stenus formicetorum Mannerheim, 1843 Stenus fornicatus Stephens, 1833 Stenus fulvicornis Stephens, 1833 Stenus fuscicornis Erichson, 1840 Stenus fuscipes Gravenhorst, 1802 Stenus geniculatus Gravenhorst, 1806 Stenus guttula Müller, 1821 Stenus guynemeri du Val, 1850 Stenus impressus Germar, 1824 Stenus incrassatus Erichson, 1839 Stenus juno (Paykull, 1789) Stenus kiesenwetteri Rosenhauer, 1856 Stenus latifrons Erichson, 1839 Stenus longitarsis Thomson, 1857 Stenus lustrator Erichson, 1839 Stenus melanarius Stephens, 1833 Stenus melanopus (Marsham, 1802) Stenus nanus Stephens, 1833 Stenus nigritulus Gyllenhal, 1827 Stenus nitens Stephens, 1833 Stenus nitidiusculus Stephens, 1833 Stenus niveus Fauvel, 1865 Stenus ochropus Kiesenwetter, 1858 Stenus opticus Gravenhorst, 1806 Stenus ossium Stephens, 1833 Stenus pallipes Gravenhorst, 1802 Stenus pallitarsis Stephens, 1833 Stenus palustris Erichson, 1839 Stenus picipennis Erichson, 1840 Stenus picipes Stephens, 1830 Stenus providus Erichson, 1839 Stenus pubescens Stephens, 1833 Stenus pusillus Stephens, 1833 Stenus similis (Herbst, 1784) Stenus solutus Erichson, 1840 Stenus subaeneus Erichson, 1840 Stenus subdepressus Mulsant & Rey, 1861 Stenus tarsalis Ljungh, 1810 Dianous coerulescens (Gyllenhal, 1810) ++ > + ++ + ++ + + + + + LBrA ? ++ + ++ * ++ + + ++ ++ + * ++ ++ + + ++ ++ ++ ++ ++ ++ + * + + Local on exposed sediments by water. Rapid coloniser of clay/sand pits. (N) Rare? In open dry situations. Rare [Wetlands and dry areas?] (Nb) Rare on ground in damp open areas? . Probably merely a form of S.pusillus (Nb) ? [Fens and marshes] Widespread but local in marshy wetlands including bogs Rare in moss and litter in damp areas. (Nb) Local, amongst richly vegetated marshy areas beside old ponds etc. Local in wetlands in marsh litter (Nb) Rare in dry sparsely vegetated places including gardens. [Chalky screes in old pits] Rare. 1 Rare. In heather litter on open heaths. Local on exposed sediments by rivers and streams also in sand and clay pits. Burghfield (Robinson, 1992) Rare [by waterfalls and swift flowing stoney streams in moss] Widespread in woodlands in leaf litter Local on margins of ponds etc. Widespread in wetlands on emergent vegetation (RDB2) Rare. In quaking bogs especially with Carex rostrata. Greenham Local in marshy wetlands and pond-margins (RDBIn) ? Wetlands including mires. Rare ? [Wetlands] Rare. In Sphagnum in bogs and mires. Local [In damp places such as manure heaps] Local in moss and litter in open grassland, a dry land species. (Nb) [Fens and mesotrophic wetlands] [In bogs and mires] Local in wetlands esp. in moss and marsh litter. (Nb) Rare in marshy wetland and bogs, Rare in moss on open grassland on the chalk. (Na) ? [In bogs and mires] (Fowler, 1888) Widespread in moss and litter in grasslands Rare in wetlands, bogs. Local on tall emergent waterside vegetation. Winters in Typha. (Nb)? [Fens] Local, in springfed wetlands, watercress beds etc. Widespread in dry areas. Local in wetlands Local in wetlands Local in moss and litter mosty in wet areas. Widespread in rough grassland etc. but not a wetland species. Widespread in wetlands Local, usually in man-made habitats, such as wet dung heaps. (RDBIn) Rare in litter under heather. Local in wetlands, frequent in flood debris Rare mossy stream margins 35 Euasthetinae Euaesthetus bipunctatus (Ljungh, 1804) Euaesthetus laeviusculus Mannerheim, 1844 Euaesthetus ruficapillus Lacordaire, 1835 Paederinae Paederus littoralis Gravenhorst, 1802 Paederus riparius (Linnaeus, 1758) Astenus immaculatus Stephens, 1833 Astenus lyonessius (Joy, 1908) Astenus pulchellus (Heer, 1839) Rugilus erichsoni (Fauvel, 1867) Rugilus fragilis (Gravenhorst, 1806) Rugilus orbiculatus (Paykull, 1789) Rugilus rufipes Germar, 1835 Rugilus similis (Erichson, 1839) Medon apicalis (Kraatz, 1857) Medon brunneus (Erichson, 1839) Medon castaneus (Gravenhorst, 1802) Medon dilutus (Erichson, 1839) Medon ripicola (Kraatz, 1854) Sunius bicolor (Olivier, 1795) Sunius melanocephalus (Fabricius, 1792) Sunius propinquus (Brisout, 1867) Pseudomedon (=Lithocharis) obscurella Erichson, 1840 Pseudomedon (=Lithocharis) obsoleta (von Nordmann, 1837) Lithocharis nigriceps Kraatz, 1859 Lithocharis ochracea (Gravenhorst, 1802) Hypomedon debilicornis (Wollaston) Scopaeus laevigatus (Gyllenhal, 1827) Scopaeus sulcicollis (Stephens, 1833) Lathrobium brunnipes (Fabricius, 1792) Lathrobium elongatum (Linnaeus, 1767) Lathrobium fovulum Stephens, 1833 Lathrobium fulvipenne (Gravenhorst, 1806) Lathrobium impressum Heer, 1841 Lathrobium longulum Gravenhorst, 1802 Lathrobium multipunctum Gravenhorst, 1802 Lathrobium pallidipenne Hochhuth, 1851 = ripicola Czwalina, 1888 Lathrobium pallidum von Nordmann, 1837 Lathrobium quadratum (Paykull, 1789) + LRom ++ ++ ++ ++ + ++ ++ ++ + LRom ? + * + + ++ + ? [Fens] formerly at , , , , ? [Mires] (Fowler, 1888) Local in moss esp. Sphagnum in marshes and bogs. Shade tolerant. Widespread in wetlands, also in open grassland esp. on chalk . Local in marshy wetlands including bogs. (N) [Marshy areas] Local in rough grassy areas esp amongst ruderal vegetation Local in decaying vegetation. Local? In litter and decaying grass in damp grassland. (N) Rare. Virginia Water 2004 (PMH) Widespread in open areas esp.with ruderal vegetation Local in many habitats at ground level (N) Rare [Eurytopic] (N) ? [Litter in woods etc.] Rare in moss and litter in (RDBIn) Rare but under-recorded in mole nests esp in sandy areas. Rare at roots of hardwoods in old parkland (N) ? [Exposed sediments by rivers] (RDBK) ? (N) [Litter in open habitats] Local in grassland (N) ? [Eurytopic] (RDBIn) ? ++ + ++ + Local, synanthropic in grass heaps esp. in gardens. An introduced Asian spp. First added to British list in 1962 by AAA Widespread in compost and grass heaps Recent immigrant, increasing in dung and compost heaps. (RDBIn) ? ? , Local on ground in damp litter and debris in woods and marshy areas. Rare in wetlands. Rare in wetlands. Rare in woods and grassland esp. near water. [in marshes and margins of fresh water] Rare in grassland, marshes and margins of freshwater. Burghfield (Robinson, 1992) Local in moss by flowing water also in manure heaps etc (N) ++ (RDBK) Rare only records relate to two found in flood debris from 1928 Rare in wetlands. Calcott 2003 (JSD) ++ ++ + + ++ + > + LBrA [ 36 Lathrobium terminatum Gravenhorst, 1802 Lathrobium volgense Hochhuth, 1851= geminum Kraatz, 1857 Achenium depressum (Gravenhorst, 1802) Achenium humile (Nicolai, 1822) Ochthephilum fracticorne (Paykull, 1800) Neobisnius lathrobioides (Baudi, 1848) Neobisnius procerulus (Gravenhorst, 1806) Neobisnius prolixus (Erichson, 1840) Neobisnius villosulus (Stephens, 1833) Erichsonius cinerascens (Gravenhorst, 1802) Erichsonius signaticornis (Mulsant & Rey, 185 Philonthus addendus Sharp, 1867 Philonthus albipes (Gravenhorst, 1802) Philonthus cognatus Stephens, 1832 Philonthus concinnus (Gravenhorst, 1802) Philonthus corruscus (Gravenhorst, 1802) Philonthus cruentatus (Gmelin, 1790) Philonthus debilis (Gravenhorst, 1802) Philonthus decorus (Gravenhorst, 1802) Philonthus discoideus (Gravenhorst, 1802) Philonthus ebeninus (Gravenhorst, 1802) Philonthus fumarius (Gravenhorst, 1806) Philonthus intermedius (Lacordaire, 1835) Philonthus jurgans Tottenham, 1937 Philonthus laminatus (Creutzer, 1799) Philonthus lepidus (Gravenhorst, 1802) Philonthus longicornis Stephens, 1832 Philonthus mannerheimi Fauvel, 1869 Philonthus marginatus (Stroem, 1768) Philonthus micans (Gravenhorst, 1802) Philonthus micantoides Benick & Lohse, 1956 Philonthus nigrita (Gravenhorst, 1806) Philonthus nigriventris Thomson, 1867 Philonthus nitidicollis (Boisduval & Lacordaire, 1835) Philonthus parvicornis (Gravenhorst, 1802) = agilis (Gravenhorst, 1806) Philonthus politus (Linnaeus, 1758) Philonthus quisquiliarius (Gyllenhal, 1810) Philonthus rectangulus Sharp, 1874 Philonthus rubripennis Stephens, 1832 Philonthus rufipes (Stephens, 1832) =immundus auct not (Gyllenhal, 1806) Philonthus sanguinolentus (Gravenhorst, 1802) LRom + + Local in moss and litter in bogs and marshy areas. Local in marshes and margins of fresh water + > + > > Local in litter in marshes and pond margins esp. on floodplains (Nb) ? Local in wet moss esp. Sphagnum in bogs and marshes. Rare ? [On exposed sediments by water] (RDBK) [On exposed sediments by water] ? (RDBK) Rare [at margins of water] also on downland at Local in marsh litter and moss in wetlands including Sphagnum bogs (Nb) [On exposed sediments by flowing water] Local in woodlands Local in dung compost and grass heaps as at Frequent in a many habitats esp. grassland and woods ? [In grass heaps] (RDBIn) ? [Carrion and decaying vegetation] Local in moss and litter in woods? etc ? most recent records I have are from Widespread in moss and litter esp. in woodland including conifer plantations Local in grass and compost heaps. (N) ? [Wetlands] (Nb) [Wetlands] Local in dung and rotting vegetation ? Widespread in dung and rotting vegetation (RDBK) Frilford 2001 JC [dunes and sandy grassland]. Rare ? (Nb) ? last recorded at Widespread in dung and carrion [Marshes] Rare in wetlands. Local in Sphagnum in bogs and wet heath areas. (PMH) (Nb) Rare. ? + Local in dung ++ ++ + Widespread in rotting vegetation including compost heaps Local in wetlands and pond margins in marsh litter. Naturalised, widespread in manure and compost heaps. + Local in dung and rotting grass cuttings + * * + ++ + ++ ++ + * + > ++ + + LRom ++ ++ + ++ 37 Philonthus spinipes Sharp, 1874 Philonthus splendens (Fabricius, 1792) Philonthus succicola Thomson, 1860 Philonthus tenuicornis Mulsant & Rey, 1853 Philonthus umbratilis (Gravenhorst, 1802) + ++ + ++ ++ Naturalised, recent immigrant. In dung heaps. Widespread in herbivore dung Local ? in carrion etc. Local ? ? [Marshes] lists Philonthus varians (Paykull, 1789) Philonthus varius (Gyllenhal, 1810) Philonthus ventralis (Gravenhorst, 1802) Gabronthus thermarum (Aubé, 1850) Bisnius (=Philonthus) cephalotes (Gravenhorst, 1802) Bisnius (Philonthus) fimetarius (Gravenhorst, 1802) Bisnius (=Philonthus) nigriventris Thomson, 1867 Bisnius (=Philonthus) pseudoparcus Brunne, 1976 Bisnius (=Philonthus) puella von Nordmann, 1837 Bisnius (=Philonthus) sordidus (Gravenhorst, 1802) Bisnius (=Philonthus) subuliformis (Gravenhorst, 1802) Gabrius exiguus (von Nordmann, 1837) Gabrius nigritulus (Gravenhorst, 1802) Gabrius osseticus (Kolenati, 1846) Gabrius pennatus Sharp, 1910 Gabrius piliger Mulsant & Rey, 1876 Gabrius splendidulus (Gravenhorst, 1802) Gabrius subnigritulus (Reitter, 1909) Gabrius trossulus (von Nordmann, 1837) Gabrius velox Sharp, 1910 Dinothenarus (=Platydracus) pubescens (Degeer, 1774) Ontholestes murinus (Linnaeus, 1758) Ontholestes tessellatus (Fourcroy, 1758) Platydracus fulvipes (Scopoli, 1763) Platydracus latebricola (Gravenhorst, 1806) Platydracus stercorarius (Olivier, 1795) Staphylinus dimidiaticornis Gemminger, 1851 Staphylinus erythropterus Linnaeus, 1758 Ocypus aeneocephalus (Degeer, 1774) Ocypus brunnipes (Fabricius, 1781) Ocypus fortunatarum Wollaston, 1871 Ocypus fuscatus (Gravenhorst, 1802) ++ ++ ++ ++ Widespread and frequent in dung etc. strong flyer esp. on warm summer evenings Widespread and frequent ? [In dung and decaying vegetable matter] Widespread in compost and dung heaps Widespread in compost heaps, and other rotting plant debris. ++ Widespread in moss and litter esp. in woods > (PMH) + (N) Only county record to date ++ In dung and rotting vegetation + Local in rotting vegetation, including compost heaps. + Rare ? ++ ++ + ++ + ++ + Rom ++ + > + ++ ++ ++ ++ ++ + ++ Rare. Rare in open sandy areas near water. (Nb) Wetlands in litter. Local in litter and moss near water ? [In litter and carrion] esp. on ? Widespread in dead wood and under bark in woodland countywide Local in litter in marshes and weedy pond margins Rare, in flood litter in marshy areas. (Nb) Rare at margins of a lake. (Nb) Rare. Uder dung, Widespread around mammal dung esp. cow and horse. Local on dung and carrion (Nb) Rare. (Nb) Rare. Open ground with Myrmica ants esp. on heaths, and downland as at Widespread and frequent in most habitats incl. gardens V.local in grasslands, & heaths. Local in grasslands. Local in open areas esp. on sandy soils, possibly associated with rabbit burrows. Primarily a winter species. Local on ground in a variety of open grassy habitats (Nb) Isi [dry sandy grassland and open heaths] (Nb) ? [Terrestrial, eurytopic] 38 Ocypus olens (Müller, 1764) Devil’s coach horse. Ocypus ophthalmicus (Scopoli, 1763) Tasgius (=Ocypus) ater (Gravenhorst, 1802) Tasgius (=Ocypus) globulifer (Fourcroy, 1785) Tasgius (=Ocypus) melanarius (Heer, 1839) Tasgius (=Ocypus) morsitans (Rossi, 1790) =compressus (Marsham, 1802) Tasgius (=Ocypus) winkleri (Bernhauer, 1906) Creophilus maxillosus (Linnaeus, 1758) Heterothops dissimilis (Gravenhorst, 1802) Heterothops minutus Wollaston Heterothops niger Kraatz, 1868 Heterothops praevius Erichson, 1839 Velleius dilatatus (Fabricius, 1787) Quedius aetolicus Kraatz, 1858 Quedius boops (Gravenhorst, 1802) Quedius brevicornis (Thomson, 1860) Quedius brevis Erichson, 1840 Quedius cinctus (Paykull, 1790) Quedius cruentus (Olivier, 1795) Quedius curtipennis Bernhauer, 1908 Quedius fulgidus (Fabricius, 1792) Quedius fuliginosus (Gravenhorst, 1802) Quedius fumatus (Stephens, 1833) Quedius humeralis Stephens, 1832 Quedius invreae Gridelli, 1924 Quedius lateralis (Gravenhorst, 1802) Quedius levicollis (Brullé, 1832) = tristis (Gravenhorst. 1802) Quedius longicornis Kraatz, 1857 Quedius maurorufus (Gravenhorst, 1806) Quedius maurus (Sahlberg, 1830) Quedius mesomelinus (Marsham, 1802) Quedius microps Gravenhorst, 1847 Quedius molochinus (Gravenhorst, 1806) Quedius nemoralis Baudi, 1848 Quedius nigriceps Kraatz, 1857 Quedius nigrocaeruleus Fauvel, 1874 Quedius nitipennis (Stephens, 1833) Quedius persimilis Muls & Rey, 1876 = aridulus Jansson, 1939 Quedius picipes (Mannerheim, 1830) Quedius puncticollis (Thomson, 1867) Rom ++ Widespread, under logs and stones etc. in most habitats including gardens ++ + (Na) ? [Open sandy ground, breckland] Frilford (JMC /JSD)+ Local in disturbed habitats including gardens. Appears to have become much more frequent inland over past century or so. Local ground dwelling in a variety of habitats. Local in woods, heaths etc. 1 Local on ground in a variety of habitats esp. on sandy heaths, but also in ruderal disturbed areas in suburbia. ++ ++ ++ ++ ++ > ++ ++ ++ ++ ++ + + ++ ++ ++ + ++ + ++ ++ + ++ Local on ground in a variety of habitats including ruderal disturbed areas in suburbia. Local in carrion. (RDBIn) ? referebale to minutus Local? in compost heaps Local, though under-recorded?, The most frequent beetle in mole breeding fortress nests. ? (RDB Windsor (Na) Rare, in old nests and squirrel dreys in hollow trees etc. Silwood Park 1998-9 (RGB) in FIT Rare? (Nb) Rare in hollow trees etc in birds nests. Local in wood ant Formica rufa nests. , 1908 Tubney Widespread in grass heaps, leaf litter etc. Widespread in grass heaps, also under bark. Local in wet areas (Nb) Synanthropic in rotten straw in old stables in Local in damp litter Local in woods and carr areas Rare in decaying vegetation. Wytham Wood 1990 (V.Gusarov) (Nb) Rare. In subterranean nests of mammals. Moulsford Downs 1997 (PJH) Widespread in rotting fungi and carrion in woods. Widespread on ground in open habitats including farmland + ++ + ++ + ++ + + + ++ + (Nb) Rare in mole nests. Local. On flooded moss and litter esp. Sphagnum in bogs and fens, and marshy pond margins. Local, saproxylic in hollow trees etc. also taken at roots of old trees at Widespread in carrion and compost and grass heaps (Nb) Rare in dead hardwoods Local in damp grassland Rare under bark and in litter in woods Rare in litter. (Nb) Rare, in mole nests at Rare in litter and moss near water. Wytham Wood 1990 (V.Gusarov) Local on ground in open areas esp. on sandy soils. ++ Widespread on ground in a variety of habitats (Nb) [Mole nests, more appropriately in the breeding ‘fortresses’!] 39 Quedius schatzmayri Gridelli, 1922 Quedius scintillans (Gravenhorst, 1806) Quedius scitus (Gravenhorst, 1806) Quedius semiaeneus (Stephens, 1833) Quedius semiobscurus (Marsham, 1802) Quedius truncicola Fairmaire & Laboul, 1856 = ventralis (Aragona, 1830) Othius angustus Stephens, 1833 Othius laeviusculus Stephens, 1833 Othius punctulatus (Goeze, 1777) Othius subuliformis Stephens, 1833 =myrmecophilus Kiesenwetter,1843 Atrecus affinis (Paykull, 1789) Leptacinus batychrus (Gyllenhal, 1827) Leptacinus formicetorum Märkel, 1841 Leptacinus intermedius Donisthorpe, 1936 Leptacinus pusillus (Stephens, 1833) Phacophallus parumpunctatus (Gyllenhal, 1827) Phacophallus tricolor (Kraatz, 1859) =pallidipennis (Motschulsky, 1858 Gauropterus fulgidus (Fabricius, 1787) Nudobius lentus (Gravenhorst, 1806) Gyrohypnus angustatus Stephens, 1833 Gyrohypnus atratus (Heer, 1839) Gyrohypnus fracticornis (Müller, 1776) Gyrohypnus liebei Scheerpeltz,1926 =punctulatus (Paykull, 1789) Megalinus (=Xantholinus) glabratus (Gravenhorst, 1802) Hypogyra (=Xantholinus) angularis Ganglbauer, 1895 Xantholinus elegans (Olivier, 1795) = jarrigei Coiffait, 1956 Xantholinus gallicus Coiffait, 1956 Xantholinus linearis (Olivier, 1795) LRom Rom Ro/S ax Rom Xantholinus longiventris Heer, 1839 Ro/S ax Rom Lucanidae (Stag beetles) 3 species SUB Lucanus cervus (Linnaeus, 1758) Stag beetle ++ ++ + + ++ ++ Local on ground in a variety of habitats Local (Nb) Rare, saproxylic in hollow trees etc. Rare? Local on ground in open habitats esp. on sandy soils? (Nb) Rare, saproxylic in hollow trees in wood mould etc.. + + + ? Old records from lists. Local in many habitats at ground level. Widespread in litter and moss in woodlands Widespread in many habitats at ground level, frequent in leaf litter. + + + Widespread in dead wood and under bark in woodlands Local in grass and compost heaps Local, but often abundant in the nests of the Wood Ant Formica rufa. > + + ? In compost heaps etc. Local in grass and compost heaps Local in grass and compost heaps. ? [In grass and compost heaps] ++ + + + ++ Local. In dung and compost heaps. Local. Under bark of pines and firs. Local in many habitats at ground level (Nb) Local in nests of the Wood Ant Formica rufa Local in dung and rotting vegetation > ? + Widespread in dung heaps ++ (Na) Rare. Saproxylic in hollow trees, rotholes etc. Silwood Park 1998 (RGB) in FIT + Local in open habitats including ruderal areas. + + Local in woods and open heathland on sandy soils Common in many habitats at ground level + Common in many habitats at ground level VCH 1907 2005 + Notes (Nb) Widespread especially in suburban gardens 40 Dorcus parallelipipedus (Linnaeus, 1758) Lesser stag beetle Platycerus caraboides (L., 1758) Sinodendron cylindricum (Linnaeus,, 1758) Lesser rhino beetle Rom Trogidae 2 species recorded SUB Trox sabulosus (Linnaeus, 1758) Trox scaber (Linnaeus, 1767) Rom Rom Bolboceratidae 1 species recorded SUB Odonteus armiger (Scopoli, 1772) Geotrupidae (Dor Beetles) 7 species Typhaeus typhoeus (Linnaeus, 1758) Minotaur beetle Anoplotrupes (=Geotrupes) stercorosus (Scriba, 1791) Geotrupes mutator (Marsham, 1802) Geotrupes spiniger (Marsham, 1802) Geotrupes stercorarius (L.innaeus, 1758) Dumble-dor, Dor beetle, Lousy watchman Trypocopris (=Geotrupes) vernalis (Linnaeus, 1758) Scarabaeidae (Dung Beetles & Chafers) 60 species + 2 sub-fossil Aphodius ater (Degeer, 1774) Aphodius borealis Gyllenhal, 1827 Aphodius coenosus (Panzer, 1798) Aphodius conspurcatus (Linnaeus, 1758) Aphodius constans Duftschmid, 1805 Aphodius contaminatus (Herbst, 1783) Aphodius distinctus (Müller, 1776) Aphodius equestris (Panzer, 1798) Aphodius erraticus (Linnaeus, 1758) Aphodius fimetarius (Linnaeus, 1758) SUB + Widespread in hardwoods and occasionally in chipboard! * + Extinct Widespread in hardwoods especially ash. VCH 1907 2005 + Notes (Na) Extinct ? [Under dry carrion] Local in hollow trees in old bird nests, and also in compost heaps. VCH 1907 2005 > Notes VCH 1907 2005 + Notes (Na) Rare, subterranean, comes to light. Wellington College (Elton) Locally frequent on sandy heaths gathering dung esp.rabbit. Rare, extinct? + + (Nb) Extinct? [In dung] ESax SUB VCH LRom Rom LRom Sax Rom? (Nb) [In dung esp. cow]. Widespread but local in pastures and on heaths on a variety of dung Rare? 1907 2005 ++ + + ++ ++ ++ ++ ++ Notes Widespread in dung Local in dung esp. deer in parks. etc. (Nb) Local on sandy soils. (Nb) Rare in dung. Rare in dung. Widespread in dung esp. cow. (Nb) Rare? Extinct? on dung in sandy areas Local in dung esp. horse in shady places esp. on sandy heaths Widespread in dung esp. cow. Widespread in dung, but see pedellus below! 41 Aphodius fasciatus (Olivier, 1789) Aphodius foetens (Fabricius, 1787) Aphodius foetidus (Herbst, 1783) Aphodius fossor (Linnaeus, 1758) Aphodius granarius (Linnaeus, 1767) Aphodius haemorrhoidalis (Linnaeus, 1758) Aphodius ictericus (Laicharting, 1781) Aphodius luridus (Fabricius, 1775) Aphodius merdarius (Fabricius, 1775) Aphodius obliteratus Panzer, 1823 Aphodius paykulli Bedel, 1908 Aphodius pedellus (DeGeer, 1774) Aphodius porcus (Fabricius, 1792) Aphodius prodromus (Brahm, 1790) Aphodius pusillus (Herbst, 1789) Aphodius putridus (Fourcroy, 1785) Aphodius quadrimaculatus (Linnaeus, 1761) Aphodius rufipes (Linnaeus, 1758) Aphodius rufus (Moll, 1782) Aphodius sphacelatus (Panzer, 1798) Aphodius varians Duftschmid, 1805 Aphodius zenkeri Germar, 1813 Euheptaulacus villosus Gyllenhal, 1806 Oxyomus sylvestris (Scopoli, 1763) Saprosites natalensis Peringuy, 1901 Copris lunaris (Linnaeus, 1758) Rhino beetle Onthophagus coenobita (Herbst, 1783) Onthophagus joannae Goljan, 1953 Onthophagus similis (Scriba, 1790) Onthophagus vacca (Linnaeus, 1767) Serica brunnea (Linnaeus, 1758) Omaloplia ruricola (Fabricius, 1775) Downland chafer Amphimallon ochraceus (Knock, 1801) Amphimallon solstitialis (Linnaeus, 1758) Summer or Fern Chafer Melolontha melolontha (Linnaeus, 1758) Cock-chafer or Maybug Hoplia philanthus (Fuessly, 1775) Phyllopertha horticola (Linnaeus, 1758) Garden chafer or Bracken-clock Ro/S ax BrAg e Sax Rom LRom Ro/S ax LRom Rom BrAg e LRom Rom Rom BrAg e Rom ++ ++ (Nb) Sub-fossil only. Local on dung Local in dung esp. cow on sandy soils ++ ++ Widespread in dung esp. cow. Widespread in dung ++ Widespread in dung esp. cow. + > ++ + + + Widespread in dung. Rare much declined in 20th Century, with no record since 1951. Local on dung esp. horse in the autumn. (Nb) Rare. Abundant in LRoman times at Barton Court (Rob Local? see Wilson (2001) this recent ‘split’ comes too early to assess differences with fimetarius (Nb) Rare in dung on open grassland (parasitic on Geotrupes). F ++ ++ * * ++ ++ ++ Widespread in dung Widespread in dung (Nb) Extinct? Newbury (Harwood) (RDB1) Extinct? [Dung in open habitats] Widespread in dung, comes readily to light, esp. MV light traps. Widespread in dung Common in dung and frequently taken on the wing and drowned in troughs puddles etc. Ru (Nb) Rare? On deer dung locally abundant in deer parks but at very low density in woodland generally (Na) Rare, Newbury Streatley Local in compost heaps Local in old parkland introduced in 20th C (RDB1) Extinct? Tubney 1913 (JJW) ++ * ++ + + ++ ++ ++ ++ ++ Local in dung Local in sheep and rabbit dung on the , seems to avoid sandy soils in Surrey! Local on dung esp. on sandy soils ? Last recorded at in 1940s [in dung] Widespread esp. in sandy areas. Frequently comes to light (Nb) Rare on open downland, but can be numerous when found. * ++ (Na) Local swarms around trees in high summer. Also comes to light ++ Local, declining? Comes to light. Larvae develope at roots of grass etc. ++ Local esp. on heathland and heathy commons, also on downland. ++ Local in many habitats including gardens and rough grassland 42 Anomala dubia (Scopoli, 1763) Sand chafer Cetonia aurata (Linnaeus, 1758) Rose Chafer Gnorimus nobilis (Linnaeus, 1758) Noble chafer Gnorimus variabilis (Linnaeus, 1758) Dascillidae 1 species SUB Dascillus cervinus (Linnaeus, 1758) +V Clambidae (Minute beetles) SUB 7 species Calyptomerus dubius (Marsham, 1802) Clambus armadillo (Degeer, 1774) Clambus gibbulus (LeConte, 1850) Clambus pallidulus Reitter, 1911 Clambus pubescens Redtenbacher, 1849 Clambus punctulus (Beck, 1817) Clambus simsoni Blackburn, 1902 LRom Scirtidae (Marsh Beetles) SUB 16 species Elodes elongata Tournier,1868 =koelleri Klausnitzer, 1970 Elodes marginata (Fabricius, 1798) Elodes minuta (Linnaeus, 1767) Elodes pseudominuta Klausnitzer, 1971 Elodes tricuspis Nyholm, 1985 =elongata sensu Klausnitzer Microcara testacea (Linnaeus, 1767) Cyphon coarctatus Paykull, 1799 Cyphon hilaris Nyholm, 1944 Cyphon laevipennis Tournier, 1868 Cyphon ochraceus Stephens, 1830 Cyphon padi (Linnaeus, 1758) Cyphon palustris Thomson, 1855 Cyphon pubescens (Fabricius, 1792) Cyphon variabilis (Thunberg, 1787) Prionocyphon serricornis (Müller, 1821) Scirtes hemisphaericus (Linnaeus, 1758) VCH VCH + ++ ++ V.local in open sandy areas. Very local not uncommon in Blackwater Valley (RDB2) [In rotting wood esp. fruit trees] Windsor ++ (RDB1) [In red rotten oak] Windsor 1907 2005 ++ Notes 1907 2005 ++ + Notes ++ ++ + ++ VCH 1907 2005 ++ Local in grassland esp.on the , on flowers esp. umbellifers. Local in compost, grass heaps, also in wetlands. Wytham (JJW) ? Wetland species. Rare [In compost heaps, and other decaying vegetation] (RDBK) Silwood Park 1998-9 (RGB) in FIT Local in manure, old haystacks, grass heaps. ? In litter etc. Recent introduction. Silwood Park 1998-9 (RGB) in FIT Notes Local, larvae in streams, avoiding very acidic waters [Also in fens]. Adults on foliage esp. close to water. + ++ ++ ++ Rare. Shaded streams Local. Acidic flowing water on heaths and Seepages in acid areas in woods. Hutchin’s Copse 1998 (JSD) (RDBIn) Rare. Windsor [Breeds in streams, adults on trees and bushes] ++ ++ ++ ++ ++ ++ ++ ++ ++ ++ ++ Widespread in wooded areas. Adults on foliage, larvae in leaf filled puddles and pools Widespread in wetlands and near streams and ponds of all kinds. Often abundant on foliage. Local. Frequent on wet heaths, also in fens on peaty soils. Local. In well vegetated wetlands esp. in reed beds. Adults winter in Typha stems. Local in poor fen areas, absent from very acid wetlands. Local. In wetlands especially in areas of acid water on peaty soils, with Sphagnum esp. in open carr woodland. Rare. Near streams Virginia Water (PJH), (Nb) Rare in richer wetlands, poor fen areas and marshes. Local. In wetlands especially in areas of acid water (Nb) Widespread. Common in flooded rot holes in trees, commonest in areas with abundant large beech trees. Local. Weedy pond margins, larvae in flooded moss and debris. 43 Eucinetidae 1 species SUB 1907 2005 + Notes Notes Simplocaria semistriata (Fabricius, 1794) Cytilus sericeus (Forster, 1771) Byrrhus fasciatus (Forster, 1771) Byrrhus pilula (Linnaeus, 1758) Byrrhus pustulatus (Forster, 1771) Porcinolus murinus (Fabricius, 1794) Curimopsis maritima (Marsham, 1802) Chaetophora spinosa (Rossi, 1794) VCH 1907 2005 LRom ++ ++ + + + + ++ + Psephenidae 1 species recorded SUB VCH 1907 2005 ++ Notes VCH 1907 2005 ++ ++ Notes VCH 1907 2005 Notes Eucinetus meridionalis (Laporte de Castelnau, 1836) Byrrhidae (Pill Beetles) 8 species SUB Eubria palustris Germar, 1818 Heteroceridae 2 species SUB Heterocerus fenestratus (Thunberg, 1784) Heterocerus marginatus (Fabricius, 1787) Limnichidae 1 species Limnichus pygmaeus (Sturm, 1807) Dryopidae 6 species Pomatinus substriatus (Müller, 1806) Dryops auriculatus (Fourcroy, 1785) Dryops ernesti des Gozis, 1886 Dryops luridus (Erichson, 1847) Dryops similaris Bollow, 1936 VCH SUB LBrA SUB Recent immigrant? Rare. Under pine bark etc in plantations on heathland. Local on ground in open sandy areas. Widespread in moss and often on umbellifers. Rare on heathland and sandy areas. Local at roots and in moss on open grassland and heaths Rare on heaths. Tubney (JJW) (Nb) Rare on heaths. ? [Bare places on sand and chalk] Frilford (JMC/ JSD) Rare on bare places on the chalk [and sand] (RDB3) Rare in fens and [seepages] larvae aquatice adults often on sallow blossoms. Cothill Local on bare ground beside water in sand and clay pits and banks of rivers ( etc), also frequent at light. Local on bare ground beside water in sand and clay pits and banks of rivers ( etc), also frequent at light. (Nb) Late Bronze Age at Burghfield (Robinson, 1992) VCH 1907 2005 + ++ Notes (Na) ? [On logs in rivers] hard to find in deep water and easily overlooked] (Nb) [Wetlands, pond-margins] Local in sandy areas on river banks, damp fields etc. Widespread at margins of water (ponds, seepages etc.) (RDB3) [Fens] 44 (RDB3) Rare at margins of pools on sandy heaths. Dryops striatellus (Fairmaire & Brisout, 1859 Elmidae (Riffle Beetles) 7 species + 4 sub-fossil Elmis aenea (Müller, 1806) Esolus parallelepipedus (Müller, 1806) Limnius volckmari (Panzer, 1793) Macronychus quadrituberculatus Muller, 1806 Normandia nitens (Müller, 1817) Riolus cupreus (Müller, 1806) Riolus subviolaceus (Müller, 1817) * Oulimnius tuberculatus (Müller, 1806) Oulimnius troglodytes (Gyllenhal, 1827) Oulimnius rivularis (Rosenhauer, 1856) Stenelmis canaliculata (Gyllenhal, 1808) Buprestidae (Jewel Beetles) 11 species Melanophila acuminata (Degeer, 1774) Agrilus angustulus (Illiger, 1803) Agrilus biguttatus (Fabricius, 1777) =pannonicus (Piller & Mitterpacher, 1783) Agrilus laticornis (Illiger, 1803) Agrilus sinuatus (Olivier, 1790) Hawthorn jewel beetle Agrilus sulcicollis Lacordaire, 1835 * Agrilus viridis (Linnaeus, 1758) Aphanisticus emarginatus (Olivier, 1790) Aphanisticus pusillus (Olivier, 1790) Trachys minuta (Linnaeus, 1758) Trachys scrobiculatus Kiesenwetter, 1857 Trachys troglodytes Gyllenhal, 1817 Elateridae (Click Beetles) 43 species + 1 sub-fossil Lacon querceus (Herbst, 1784) Agrypnus murinus (Linnaeus, 1758) SUB LBrA LBrA LBrA LBrA VCH 1907 2005 ++ ++ ++ ++ ++ SUB VCH 1907 2005 + ++ ++ ++ ++ ++ ++ Rom SUB Ro/Sa x ++ + ++ VCH 1907 2005 ++ ++ Ampedus balteatus (Linnaeus, 1758) ++ Ampedus cardinalis (Schiodte, 1865) Ampedus elongantulus (Fabricius, 1787) ++ ++ Notes Widespread on stones, logs and plant roots in streams Late Bronze Age Burghfield (Robinson, 1992) [In streams] Widespread on stones, logs and plant roots in streams (RDB3) Holybrook Reading in 2003 (JSD) (RDB2) Burghfield (Robinson, 1992) (Nb) In streams. (Nb) Rare in streams. Local in weedy margins of ponds canals and flowing water. (Nb) ? Extinct? [Canals, lakes, streams] (Na) [Debris at margins of stil land slow flowing water] 1870 (RDB2) Notes Rare on burnt trees (pine & birch) on heathland (Nb) Saproxylic. Local on oak, hazel etc. (Na) Saproxylic. Much increased after 1987! Local on large oaks, where the D shaped emergence holes are charcteristic. (Nb) Saproxylic. Local on oaks in woods and hedgerows. (Na!) Saproxylic. Widespread on hawthorn, in hedgerows, parks, etc. Also in Pear trees Pyrus communis ? Recent colonist on oak, (Na) Rare on sallow etc. (RDB1) Extinct? On Juncus in disturbed areas in woods and heaths. (Nb) Local on the chalk on open downland turf, also on acid grassland Bradfield (NHJ) (RDB2) ? [On sallows, hazel etc.] (Na) Local on ground ivy Glechoma hederacea on chalk and clinkery waste ground. >10 post 1980 records. Rare. [On devil’s-bit scabious Succisa pratensis ] Notes (RDB1) Windsor Forest Local on sandy soils Local on various rotten stumps esp. pine quite a frequent sight on the large heaths in the west of the county. (RDB2) Rare in red-rotten oak. (Na) Rare in heathy areas on pine, and oak and beech. Dinton Pastures 1997 (AJH) 45 Ampedus nigerrimus (Lacordaire, 1835) Ampedus quercicola (du Buysson, 1887) Ampedus rufipennis (Stephens, 1830) Ampedus sanguinolentus (Schrank, 1776) ++ ++ ++ ++ Brachygonus ruficeps (Mulsant & Guillebeau, 1855) Ischnodes sanguinicollis (Panzer, 1793) Procraerus tibialis (Boisduval & Lacordaire, 1835) Megapenthes lugens (Redtenbacher, 1842) Hypnoidus riparius (Fabricius, 1792) Oedostethus quadripustulatus (Fabricius, 1792) Panspoeus guttatus Sharp, 1877 Cardiophorus asellus Erichson, 1840 Cardiophorus gramineus (Scopoli, 1763) Melanotus castaneipes Cidnopus aeruginosus (Olivier, 1790) Kibunea minuta (Linnaeus, 1758) Stenagostus rhombeus (Olivier, 1790) Athous bicolor (Goeze, 1777) Athous campyloides Newman, 1833 Athous haemorrhoidalis (Fabricius, 1801) Athous hirtus (Herbst, 1784) Athous subfuscus (Müller, 1764) Athous vittatus (Fabricius, 1792) Actenicerus sjaelandicus (Müller, 1764) Ctenicera cuprea (Fabricius, 1775) Ctenicera pectinicornis (Linnaeus, 1758) Calambus bipustulatus (Linnaeus, 1767) Aplotarsus incanus (Gyllenhal, 1827) Selatosomus cruciatus (L., 1758) Selatosomus nigricornis (Panzer, 1799) Prosternon tessellatum (Linnaeus, 1758) Elater ferrugineus Linnaeus, 1758 Agriotes acuminatus (Stephens, 1830) Agriotes lineatus (Linnaeus, 1767) Agriotes obscurus (Linnaeus, 1758) + (RDB1) Windsor Forest (Nb) Rare in dead wood esp. oak, birch and beech. (RDB2) Rare in white rot in various hardwoods including ash, beech, and elm (Na) V.local on heathland amongst heather, and under birch and pine bark. Snelsmore Common 2003 (JSD) (RDB1) Windsor ++ ++ (Na) Rare on rotting hardwoods. Silwood Park 1997 (MVB) (RDB3) Rare in oak, beech, ash, and horse chestnut) ++ ++ + ++ + + ++ + ++ ++ ++ ++ + ++ + ++ ++ (RDB1) [In decaying hardwood] Windsor Forest 1993 (DRC) Rare at margins of water. , (Na) Rare in a variety of habitats including wooded gardens Naturalised Australasian species, spreading on heaths and wooded commons. (Nb) Local on sandy heaths RDB app. Widespread in dead wood, also comes to MV light traps. Rare on sandy soils. Local in a variety of habitats esp. near woods. Local in dead wood esp. beech.. Frequently comes to MV light. Local in a variety of habitats esp. grassland. (Nb) Local but on the increase. In disturbed areas including gardens Common in grassy places countywide Local in woodlands often under bark, but not saproxylic. (RDB3) Increasing recent colonist? Particularly frequent on high ground, Widespread on vegetation in woods, commons. V.local in wetlands primarily on heaths. + ++ ++ * + ++ ++ ++ ++ ++ (Nb) Accidental import? (Nb) Rare, saproxylic on a variety of hardwoods Local in wetlands, frequently swept from tussocks and other emergent vegetation. Extinct. Near Windsor early 19th C (RDB3) Rare in floodplain grassland. Local in open areas esp. heaths (RDB1) Rare. In hollows in old trees oaks, horse chestnut, esp. in old bird’s nests. Windsor Widespread esp. in woodlands Widespread on ground in a variety of habitats Widespread on ground in a variety of habitats ++ ++ ++ ++ ++ ++ ++ Widespread esp. in woodlands Widespread on ground in a variety of habitats Widespread and frequent in most habitats Local esp. on sandy soils Local in woodlands (RDB1) Widespread, larvae in dead wood Agriotes pallidulus (Illiger, 1807) Agriotes sputator (Linnaeus, 1758) Dalopius marginatus (Linnaeus, 1758) Sericus brunneus (Linnaeus, 1758) Adrastus pallens (Fabricius, 1792) Limoniscus violaceus (Müller, 1821) Denticollis linearis (Linnaeus, 1758) LRom Rom LRom Rom lBrA LRom LRom Ro/Sa x LRom Rom Rom 46 Throscidae 5 species SUB Aulonothroscus brevicollis (Bonvouloir, 1859) Trixagus carinifrons (Bonvouloir, 1859) Trixagus dermestoides (Linnaeus, 1767) Trixagus obtusus (Curtis, 1827) Eucnemidae (False click beetles) 5 species recorded + 1 sub-fossil VCH 1907 2005 ++ + ++ LRom + Notes SUB VCH 1907 2005 ++ ++ ++ ++ ++ Notes VCH 1907 2005 + Notes VCH 1907 2005 ++ ++ ++ + >? ++ ++ ++ ++ ++ ++ ++ ++ ++ ++ ++ ++ Notes Melasis buprestoides (Linnaeus, 1761) Epiphanis cornutus Eschscholtz, 1829 Microrhagus pygmaeus (Fabricius, 1792) Hylis olexai (Palm, 1955) Eucnemis capucinus Ahrens, 1812 Dromaeolus barnabita (Villa, 1835?) Drilidae 1 species recorded SUB Drilus flavescens (Fourcroy, 1785) Cantharidae (Soldier Beetles) recorded 34 species Podabrus alpinus (Paykull, 1798) Cantharis cryptica Ashe, 1947 Cantharis decipiens Baudi, 1871 Cantharis figurata Mannerheim, 1843 Cantharis fusca Linnaeus, 1758 Cantharis lateralis Linnaeus, 1758 Cantharis livida Linnaeus, 1758 Cantharis nigra (Degeer, 1774) Cantharis nigricans (Müller, 1776) Cantharis pallida Goeze, 1777 Cantharis pellucida Fabricius, 1792 Cantharis rufa Linnaeus, 1758 Cantharis rustica Fallén, 1807 Cantharis thoracica (Olivier, 1790) Rhagonycha fulva (Scopoli, 1763) Rhagonycha lignosa (Müller, 1764) Rhagonycha limbata Thomson, 1864 SUB Rom Rom (RDB3) Saproxylic on old oaks etc. Windsor Silwood Park 1998 (RGB) in FIT Local in litter, moss etc. Widespread in moss litter and in dead wood and hollow trees. Local in litter and debris. (Nb) Local. Saproxylic, on oak etc. often on cut ends of fallen/felled trees. Recent immigrant? In dead wood. Moulsford Downs 1997 (PJH) Drayton Copse 1998 (JSD) (RDB3) Local,, increasing ? Saproxylic in old oak. (RDB3) Local, increasing? Saproxylic in hardwoods esp. beech, Silwood Park 1998 (RGB) in FIT (RDB1) Saproxylic, in hollow beech Sub-fossil only. (Nb) Local. Chalk grassland along , males strong fliers, often seen in gardens on flowers etc. Local and primarily Widespread esp. on trees. Widespread and eurytopic Rare, in damp grassland. 1993 (RDB3) Local in grassland, woods and parkland Widespread on tree and herbs esp. in woodlands and wooded commons. Widespread in grassland and on trees and herbs. Widespread on trees and herbs in woods and commons Local on vegetation near water. Widespread in most habitats Widespread in woods and parkland. Widespread in woods, heaths, and chalk grassland Local in wetland areas, margins of lakes, ponds, canals and rivers Ubiquitous on umbels etc. in high summer. Widespread on trees and herbs. Widespread on trees and herbs. 47 Rhagonycha lutea (Müller, 1764) Rhagonycha testacea (Linnaeus, 1758) Rhagonycha translucida (Krynicki, 1832) Silis ruficollis (Fabricius, 1775) Malthinus balteatus Suffrian, 1851 Malthinus flaveolus (Herbst, 1786) Malthinus frontalis (Marsham, 1802) Malthinus seriepunctatus Kiesenwetter, 1852 Malthodes crassicornis (Maeklin, 1846) Malthodes dispar (Germar, 1824) Malthodes fibulatus Kiesenwetter, 1852 Malthodes fuscus (Waltl, 1838) Malthodes guttifer Kiesenwetter, 1852 Malthodes marginatus (Latreille, 1806) Malthodes maurus (Castelnau, 1840) Malthodes minimus (Linnaeus, 1758) Malthodes mysticus Kiesenwetter, 1852 Malthodes pumilus (Brebisson, 1835) Lampyridae (Glow-worms) 1 species ++ ++ ++ ++ ++ ++ ++ ++ + ++ ++ + + ++ + ++ +? ++ SUB Lampyris noctiluca (Linnaeus, 1758) Glow worm Lycidae (Net-winged Beetles) 1 species SUB Platycis minutus (Fabricius, 1787) Dermestidae (Larder beetles and allies) species (see Peacock, 1993) Dermestes ater Degeer, 1774 Dermestes haemorrhoidalis Kuester, 1852 Dermestes lardarius L.innaeus, 1758 Larder or Bacon beetle Dermestes maculatus Degeer, 1774 Hide beetle Dermestes murinus Linnaeus, 1758 Dermestes peruvianus Castelnau, 1840 Peruvian Larder or Hide beetle Attagenus pellio (Linnaeus, 1758) Two-spotted carpet-beetle Attagenus smirnovi Zhantiev, 1973 SUB ESax (Nb) Local on trees and hedgerows esp. in old woods and parklands Local primarily on heaths and heathy commons, (Nb) Rare in woodlands. Rare on tall vegetation by water. , (Nb) Rare, saproxylic in damp woodland? Local but widespread. Saproxylic, in woods and hedgerows (Nb) Rare, saproxylic, esp. on trees in open parkland. Local but widespread, saproxylic (RDB3) Rare, breeds in red rotten oak. Rare? [Wet woodlands]. (Nb) Rare in woods on calcareous soils Rare, saproxylic in woodland. (Nb) Rare, saproxylic. Local but widespread, saproxylic Local but widespread, saproxylic Rare, woodlands. Rare. Saproxylic in hardwoods. VCH 1907 2005 ++ Notes VCH 1907 2005 ++ Notes VCH 1907 2005 Notes + ++ * Local in many habitats from grassland to bogs, commonest in calcareous areas with abundant snails.. (Nb) Local, saproxylic, on rotten hardwoods, adult mostly in late summer/early autumn. ? Synanthropic Synanthropic in bird’s nests and carrion,Windsor Forest 1986 (H Mendel). Local in carrion bird and wasp nests. Synanthropic in carrion Windsor Forest 1936 ++ + Rare, on carrion. Local, increasing, synanthropic in accumulations of pigeon droppings and debris etc. ++ Local, synanthropic in houses, warehouses etc. also in hollow trees outdoors. + Synanthropic 48 Attagenus unicolor (Brahm, 1791) Black carpet beetle Trogoderma granarium Everts Khaphra beetle Megatoma undata (Linnaeus, 1758) Anthrenocerus australis (Hope, 1843) Australian carpet beetle Ctesias serra (Fabricius, 1792) Cob-web beetle Globicornis rufitarsis (Panzer) = nigripes (Fabricius, 1792) Anthrenus fuscus Olivier, 1789 Anthrenus museorum (Linnaeus, 1761) Museum beetle Anthrenus verbasci (Linnaeus, 1767) Varied carpet-beetle Trinodes hirtus (Fabricius, 1781) Derodontidae (Tooth-necked fungus beetle) 1 species only SUB Laricobius erichsoni Rosenhauer, 1846 Anobiidae (Woodworms) 27 species Ptinomorphus imperialis (Linnaeus, 1767) Grynobius planus (Fabricius, 1787) Dryophilus anobioides Chevrolat, 1832 Dryophilus pusillus (Gyllenhal, 1808) Ochina ptinoides (Marsham, 1802) Xestobium rufovillosum (Degeer, 1774) Death-watch beetle Ernobius angusticollis (Ratzeburg, 1837) Ernobius gigas (Mulsant & Rey, 1863) Ernobius mollis (Linnaeus, 1758) Ernobius nigrinus (Sturm, 1837) Ernobius pini (Sturm, 1837) Gastrallus immarginatus (Müller, 1821) Hemicoelus nitidus (Herbst, 1793) Hadrobregmus denticollis (Creutzer in Panzer, 1796) Hemicoelus fulvicornis (Sturm, 1837) Anobium inexspectatum Lohse, 1954 Anobium punctatum (Degeer, 1774) Furniture beetle or Woodworm Xyletinus longitarsis Jansson, 1942 Stegobium paniceum (Linnaeus, 1758) Bread beetle + Synanthropic breeding in shops and factories + ++ Slough 1951 (AMM) (Nb) V.local, Under bark, in old bird nests, spiders webs etc. ++ ++ (Nb) Local under bark of trees in spider’s webs etc. (RDB1) Slough (Woodroffe) Windsor 1986 (JAO) + ++ Widespread on flowers breeding in debris in nests and buildings. Widespread on flowers breeding in debris in nests and buildings. ++ Widespread on flowers breeding in nests and in debris in habitations ++ (RDB3) Rare under bark of trees in spider’s webs etc. VCH 1907 2005 ++ SUB LRom Ro/S ax Rom Notes Introduced and spreading, on conifers ++ ++ ++ ++ ++ ++ Notes (Nb) Local, breeds in dead stems of ivy, also on dead oak boughs Widespread in dead wood (RDB3) Rare on dead broom Herons Nest 2003 (JSD) Naturalised. Local in dead pine and larch Widespread on dead, thick stems of ivy. Local in old timber in buildings and in the wild esp. red-rotten oak in parklands and old woods. > + ++ + ++ ++ ++ ++ Vagrant? On pines Naturalised, found on burnt pine. Local on pine and firs Local on pine and firs, native in Scotland, but introduced Naturalised. Local on and firs. (RDB1) Rare on field maple Acer campestre. Windsor (RDBIn) [On dead wood of maple and poplar etc.] (Nb) Saproxylic [in dead heartwood of various hardwoods] ++ ++ ++ Widespread in small dead branches of hardwoods. (Nb) Local on old Ivy stems in woods and hedgerows Ubiquitous, in untreated timber in buildings, and frequent in the wild. ++ (RDB2) [In oak, broom etc] Synanthropic, cosmopolitan, occasionally outdoors. 49 Ptilinus pectinicornis (Linnaeus, 1758) Fan-bearing wood-borer Lasioderma serricorne (Fabricius, 1792) Dorcatoma ambjoerni Baranowski, 1985 Dorcatoma chrysomelina Sturm, 1837 Dorcatoma dresdensis Herbst, 1792 Dorcatoma flavicornis (Fabricius, 1792) Dorcatoma serra Panzer, 1795 Caenocara bovistae (Hoffmann, 1803) Anitys rubens (Hoffmann, 1803) Ptinidae (Spider beetles) 12 species Gibbium psylloides (Czenpinski, 1778) Tipnus unicolor (Piller & Mitterpacher, 1783) Niptus hololeucus (Faldermann, 1835) Golden spider beetle Ptinus clavipes Panzer, 1792 Ptinus fur (Linnaeus, 1758) White marked spider beetle Ptinus lichenum Marsham, 1802 Ptinus palliatus Perris, 1847 Ptinus pilosus Müller, 1821 Ptinus sexpunctatus Panzer, 1792 Ptinus subpilosus Sturm, 1837 Ptinus tectus Boieldieu, 1856 SUB ++ ++ ++ ++ ++ ++ Synanthropic. Windsor Rare in dead wood esp. red rotten oak (Na) Rare. In hard brackets on alder, beech etc. (Nb) Rare. In red rotten oak. (Na) Rare in bracket fungi esp. Inonotus. (RDB3) ? In the puff balls Lycoperdon bovista & Bovista plumbea. Tubney (JJW) (Nb) Rare. In red rotten oak. Windsor VCH 1907 2005 Rom > ++ Extinct? Widespread mainly synanthropic but also on birds nests (esp. feral pigeon) and in hollow trees. ++ + Phloiophilidae SUB VCH Notes [Cosmopolitan, synanthropic.] Cosmopolitan, synanthropic. [Cosmopolitan, synanthropic.] * ++ SUB Phloiophilus edwardsi Stephens, 1830 Widespread in a variety of hardwoods ++ Bostrichidae (False powder post beetles) 4 species Rhyzopertha dominica (Fabricius, 1792) Lesser grain-borer Dinoderus minutus Fabricius, 1775 Lyctus brunneus (Stephens, 1830) Lyctus linearis (Goeze, 1777) 1 species ++ (RDB3) [Woodland] (Na) Rare. On dead wood, especially oak, and also on old posts. Wellman’s lakes 2003 (JSD). Introduced? Extinct. (Nb) Rare, in old nests, in dead wood, and indoors. Comes to light traps. (Nb) [In old hollow trees] Naturalised. 1907- Notes 2005 Cosmopolitan, Synanthropic [in stored products] ++ LRom VCH 1907 2005 ++ Synanthropic. Saproxylic, breeds in imported timber and in the wild in hardwoods. Green park 2002 (JSD) (Nb) ? Rare, saproxylic in oak, beech etc. Notes (Nb) Rare, in the fungi Phlebia merismoides on dead hardwood branches. 50 Trogossitidae (Bark-gnawing beetles) 1 species SUB Tenebroides mauritanicus (Linnaeus, 1758) Cadelle Cleridae (Checkered-beetles) 11 species SUB Paratillus carus (Newman, 1840) Tillus elongatus (Linnaeus, 1758) Tilloidea unifasciatus (Fabricius, 1787) Opilo mollis (Linnaeus, 1758) Thanasimus formicarius (Linnaeus, 1758) Ant beetle Korynetes caeruleus (Degeer, 1775) Necrobia ruficollis (Fabricius, 1775) Necrobia rufipes (Degeer, 1775) Copra beetle or Red-legged ham beetle Necrobia violacea (Linnaeus, 1758) Melyridae (Soft-winged flower beetles) 16 species Aplocnemus pini (Redtenbacher, 1849) Dasytes aeratus Stephens, 1830 Dasytes niger (Linnaeus, 1761) Dasytes plumbeus (Müller, 1776) Dasytes puncticollis Reitter, 1888 Ebaeus pedicularis (Linnaeus, 1758) Axinotarsus marginalis (Castelnau, 1840) Axinotarsus ruficollis (Olivier, 1790) Malachius aeneus (Linnaeus, 1758) Malachius bipustulatus (Linnaeus, 1758) Malachius marginellus Olivier, 1790 Malachius viridis Fabricius, 1787 Anthocomus fasciatus (Linnaeus, 1758) Anthocomus rufus (Herbst, 1784) Sailor beetle VCH 1907 2005 > Notes VCH 19072005 + ++ * ++ ++ Notes + Synanthropic, cosmopolitan Naturalised Australasian spp. Predates Lyctus in timber. (Nb) Local in dead wood in woods and hedgerows. Attacking Scolytids. [Predates Lyctus in old stumps and posts] (Nb) Local. In hollow trees also in houses attaching Anobiids in timber. Local, on wide variety of trees attacking bark beetles, including scot’s pine on heaths. (Nb) Rare in wood attacking Anobiids. Also in Synanthropic sites. Cosmopolitan [in carrion, old bones etc.] ? [In stored seeds and beans] Cosmopolitan. Local in carrion, old bones SUB VCH 19072005 ++ ++ ++ ++ * ++ + * ++ Rom ++ ++ ++ Lymnexylidae (Ship-timber beetles) 1 species Lymexylon navale (Linnaeus, 1758) SUB Kateritidae 3 species. For list of published records see Kirk-Spriggs (1996) SUB Notes (Nb) Rare in ancient wood/parkland. Widespread and frequent esp. on blossoms (hawthorn etc.) (Na) Rare in grassland but breeding in dead wood. (Nb) Local in woods and hedgerows. (Nb) ? [Grasslands] Extinct Widespread, spreading rapidly in 20th C on grass and herbage, breeding in dead wood esp. oak. Very local develops in dead wood (RDB3) Rare, in old unimproved grasslands. Widespread on flowers and herbage. Saproxylic. (Nb) Widespread on flowers and herbage. Local, in woods, parks and rural gardens. Saproxylic. Rare, locally abundant in tall vegetation near water esp. reed Phragmites beds. VCH 19072005 ++ Notes VCH 19072005 Notes (RDB2) Saproxylic in heartwood of Oaks. 51 Kateretes pusillus (Thunberg, 1794 Kateretes pedicularius (Linnaeus, 1758) Kateretes rufilabris (Latreille, 1807) Nitidulidae (Pollen & Sap beetles) 77 species For list of published records see Kirk-Spriggs (1996) Brachypterus glaber (Stephens, 1835) Brachypterus urticae (Fabricius, 1792) Brachypterolus linariae (Stephens, 1830) Brachypterolus pulicarius (Linnaeus, 1758) Brachypterolus villiger (Reitter, 1885) Brachypterolus vestitus (Kiesenwetter, 1850) Carpophilus hemipterus (Linnaeus, 1758) Dried fruit beetle Carpophilus marginellus Motschulsky, 1858 Carpophilus sexpustulatus (Fabricius, 1792) Pria dulcamarae (Scopoli, 1763) Meligethes aeneus (Fabricius, 1775) Meligethes atramentarius Foerster, 1849 Meligethes atratus (Olivier, 1790) Meligethes bidens Brisout, 1863 Meligethes bidentatus Brisout, 1863 Meligethes brunnicornis Sturm, 1845 Meligethes carinulatus Förster 1849 Meligethes difficilis (Heer, 1841) Meligethes erichsoni Brisout, 1863 Meligethes flavimanus Stephens, 1830 Meligethes fulvipes Brisout, 1863 Meligethes gagathinus Erichson, 1845 Meligethes incanus Sturm, 1845 Meligethes kunzei Erichson, 1845 Meligethes lugubris Sturm, 1845 Meligethes morosus Erichson, 1845 Meligethes nigrescens Stephens, 1830 Meligethes obscurus Erichson, 1845 Meligethes ochropus Sturm, 1845 Meligethes ovatus Sturm, 1845 Meligethes pedicularius (Gyllenhal, 1808) Meligethes persicus Falderman, 1837 Meligethes planiusculus (Heer, 1841) Meligethes rotundicollis Brisout, 1863 Meligethes ruficornis (Marsham, 1802) Meligethes solidus (Kugelann, 1794) Meligethes umbrosus Sturm, 1845 LBrA SUB ESax ++ ++ ++ VCH 19072005 Local on sedges Carex in damp areas, marshes etc. Local on sedges Carex in bogs, marshes etc. Also on flowers esp. meadowsweet Filipendula ulmaria Local on sedges and rushes in damp areas, marshes etc. Notes ++ ++ ++ + > > Common on nettles Urtica. Spp. Common on nettles Urtica. Spp. Local on toadflax Linaria spp. in grassy places including road verges Rare on Linaria. (RDBK) Rare in parks and gardens on Antirrhinum 1 Rare in parks and gardens on Antirrhinum. Cosmopolitan, synanthropic. ++ ++ ++ ++ + ++ + Widespread, synanthropic, in houses and gardens, also at sap in the wild! Rare in sappy timber. Local on bittersweet Solanum dulcamarae. Ubiquitous on various yellow crucifers, also attracted to bright clothing, one of the most common of beetles! (N) Rare on yellow archangel Lamiastrum galeobdolon Local on Rosa especially dog-rose R.canina. Rare on wild basil Clinopodium vulgare on the (RDBK) [On dyer’s greenweed Genista tinctoria] Widespread on hedge woundwort Stachys sylvatica Widespread on bird’s-foot trefoil Lotus corniculatus Widespread on white dead-nettle Lamium album (N) Rare on horseshoe vetch Hippocrepis comosa on the . Widespread on roses esp. dog-rose. Rosa canina + + + * + * + + * + + + * + + > + + + + > (N) Rare on corn mint Mentha arvensis. (N) Rare, on cat-mint Nepeta cataria. Local on yellow archangel Lamiastrum galeobdolon (N) Rare/Extinct? On wild thyme Thymus Widespread on white dead-nettle Lamium album Widespread on white clover Trifolium repens Widespread on wood-sage Teucrium scorodonia (N) Rare on marsh woundwort Stachys palustris Local on ground-ivy Glechoma hederacea Local on common hemp-nettle Galeopsis tetrahit Local on betony Stachys officinalis Very local on viper’s bugloss Echium vulgare. (N) Rare on crucifers, charlock Sinapis arvensis & hedge mustard Sisymbrium officinale Local on black horehound Ballota nigra (N) Very local on rock-rose Helianthemum on the (N) Rare. on self-heal Prunella vulgaris. 52 Meligethes viridescens (Fabricius, 1787) Epuraea aestiva (Linnaeus, 1758) Epuraea angustula Sturm, 1844 Epuraea biguttata (Thunberg, 1784) Epuraea distincta (Grimmer, 1841)* Epuraea fuscicollis (Stephens, 1832) Epuraea guttata (Olivier, 1811) Epuraea limbata (Fabricius, 1787) Epuraea longula Erichson, 1845 Epuraea melanocephala (Marsham, 1802) Epuraea melina Erichson, 1843 Epuraea neglecta (Heer, 1841) Epuraea pallescens (Stephens, 1832) Epuraea pusilla (Illiger, 1798) Epuraea rufomarginata (Stephens, 1830) Epuraea silacea (Herbst, 1784) = deleta Sturm, 1844 Epuraea terminalis (Mannerheim, 1843) Epuraea thoracica Tournier, 1872 Epuraea unicolor (Olivier, 1790) Nitidula bipunctata (Linnaeus, 1758) Nitidula carnaria (Schaller, 1783) Nitidula rufipes (Linnaeus, 1767) Omosita colon (Linnaeus, 1758) Omosita depressa (Linnaeus, 1758) Omosita discoidea (Fabricius, 1775) Soronia grisea (Linnaeus, 1758) Soronia punctatissima (Illiger, 1794) Amphotis marginata (Fabricius, 1781) Pocadius ferrugineus (Fabricius, 1775) Pocadius adustus Reitter, 1888 =lanuginosus Franz, 1969 Thalycra fervida (Olivier, 1790) Cychramus luteus (Fabricius, 1787) Cryptarcha strigata (Fabricius, 1787) Cryptarcha undata (Olivier, 1790) Pityophagus ferrugineus (Linnaeus, 1761) Glischrochilus hortensis (Fourcroy, 1785) Picnic beetle Glischrochilus quadriguttatus (Fabricius, 1777) Cybocephalus fodori Endrödy-Younga Monotomidae (= Rhizophagidae) 21 species Rhizophagus bipustulatus (Fabricius, 1792) ++ + ++ + + Rom + + + + + ++ ++ ++ ++ ++ ++ + ++ + + (RDBIn) [Under sappy bark of oak, birch and pine] (N) Rare, under bark of conifers, and at sap. Local at sap on hardwoods, and in compost heaps. . Local in carrion ? [In carrion] Local in carrion Local in carrion Local in carrion Local in carrion Under bark of ash. Local at sap, also at trees attacked by goat mot (RDBK) Rare, in or near to nests of the ant Lasius fuliginosus. Wellman’s lake 2003 (JSD) Local in fungi esp. puffballs Lycoperdon spp. Local in rotting fungi * ++ ++ ++ ++ ++ (Nb) [At sap including Cossus trees] Bradfield (NHJ) Local on blossom and fungi (Nb) Local at sap. . Bradfield (NHJ) (Nb) Rare, at sap. Bradfield (NHJ) Rare on pines. Besselsleigh Oxo?? Local at sap, in fungi, etc. ++ Local at sap. New for Britain, at + + > LRom LRom SUB Widespread on crucifers Widespread on flowers of trees and shrubs, esp. rowan, hawthorn and Gorse. (Nb) [In borings of Xyloterus beetles in deadwood] Rare? [Under bark and in fungi] (N) on soft barcjet fungi on willows in carr /fen etc. Burghfield Bridge 2003 (JSD) (Nb) Local at sap. Bradfield (NHJ) (Nb) [At sap, including Cossus trees] Bradfield (NHJ) [In fungi] (Nb) Rare in old woodlands Widespread on blossoms Widespread on blossoms (RDBIn) [Rare at sap and under bark] Widespread? On flowers, at sap and in fungi Local on blossoms [and under bark] ? [In borings of ambrosia beetles and in fungi] Widespread, saproxylic at sap, in fungi and under bark. VCH 19072005 ++ Notes Frequent under bark of many kinds of tree. 53 Rhizophagus cribratus Gyllenhal, 1827 Rhizophagus depressus (Fabricius, 1792) Rhizophagus dispar (Paykull, 1800) Rhizophagus ferrugineus (Paykull, 1800) Rhizophagus nitidulus (Fabricius, 1798) Rhizophagus oblongicollis Blatch & Horner, 1892 Rhizophagus parallelocollis Gyllenhal, 1827 Graveyard beetle Rhizophagus perforatus Erichson, 1845 Rhizophagus picipes (Olivier, 1790) Monotoma angusticollis (Gyllenhal, 1827) Monotoma bicolor Villa, 1835 Monotoma brevicollis Aubé, 1837 Monotoma conicicollis Aubé, 1837 Monotoma longicollis (Gyllenhal, 1827) Monotoma picipes Herbst, 1793 Monotoma quadricollis Aubé, 1837 Monotoma quadrifoveolata Aubé, 1837 Monotoma spinicollis Aubé, 1837 Monotoma testacea Motschulsky, 1845 Sphindidae (Dry-fungus beetles) 2 species Rom 2 species LRom Cryptolestes duplicatus (Waltl, 1839) Cryptolestes ferrugineus (Stephens, 1831) Cryptolestes pusillus (Schoenherr, 1817) Cryptolestes turcicus (Grouvelle, 1876) Cryptolestes spartii (Curtis, 1834) Notolaemus unifasciatus (Latreille, 1804) Laemophloeus monilis (Fabricius, 1787) ++ Local under dead hardwood bark, in carrion and often subterranean. ++ Under dead hardwood bark, also in compost and in large numbers in subterranean traps in gardens and woodland (Na) Rare, saproxylic, under sapy bark of various trees (RDB3) V.local in wood ant Formica rufa nests. Wellington College ? Compost heaps. ? [In grass and compost heaps] V.local in wood ant nests. Wellington College (NHJ) Widespread in compost heaps, and other decaying vegetable matter. Widespread in compost heaps, and other decaying vegetable matter. Regularly comes to light. Local in decaying vegetable matter. ? Extinct? Rare? Widespread in compost & grass heaps etc. + ++ ++ ++ + ++ SUB SUB Pediacus depressus (Herbst, 1797) Pediacus dermestoides (Fabricius, 1792) Laemophloidae 7 species Local under bark of many kinds of tree Local under bark of many kinds of tree Local under bark of many kinds of tree Frequent under bark of many kinds of tree, (Nb) Rare under bark of many kinds of tree (RDB1) Rare, saproxylic, subterranean?. * * ++ Sphindus dubius (Gyllenhal, 1808) Aspidiphorus orbiculatus (Gyllenhal, 1808) Cucujidae (Flat bark beetles) ++ ++ ++ ++ ++ ++ SUB VCH 19072005 ++ ++ Notes VCH 1907 2005 * ++ Notes VCH 1907 2005 + ++ Notes ++ ++ * (Nb) Local, In fungoid wood, feeding on slime fungus. Widespread, saproxylic. Frequent in summer swarms and in FITs, rarely encountered by other means. (Na) Rare [Sappy timber and Goat Moth Cossus workings] Local, under hardwood bark, especially beech. Local under bark of trees. Local under bark of hard and softwoods Synanthropic? (Na) Rare in dead broom bark, (Na) Rare. Under hardwood bark esp.oak. RDB1) 54 Silvanidae 9 species SUB Uleiota planata (Linnaeus, 1761) Psammoecus bipunctatus (Fabricius, 1792) Nausibius clavicornis (Kugelann, 1794 Ahasverus advena (Waltl, 1834) Foreign grain beetle Oryzaephilus mercator (Fauvel, 1889) Merchant’s grain beetle Oryzaephilus surinamensis (Linnaeus, 1758) Saw-toothed grain beetle Silvanus bidentatus (Fabricius, 1792) Silvanus unidentatus (Olivier, 1790) Cryptophagidae (Silken fungus beetles) 71 species Telmatophilus caricis (Olivier, 1790) Telmatophilus schoenherri (Gyllenhal, 1808) * Telmatophilus sparganii (Ahrens, 1812) Telmatophilus typhae (Fallén, 1802) Paramecosoma melanocephalum (Herbst, 1793) Henoticus californicus (Mannerheim, 1843) Henoticus serratus (Gyllenhal, 1808) Cryptophagus acutangulus Gyllenhal, 1827 Cryptophagus angustus Ganglbauer, 1899 Cryptophagus badius Sturm, 1845 Cryptophagus cellaris (Scopoli, 1763) Cryptophagus confusus Bruce, 1934 Cryptophagus dentatus (Herbst, 1793) Cryptophagus distinguendus Sturm, 1845 Cryptophagus falcozi Roubal, 1927 Cryptophagus intermedius Bruce, 1934 Cryptophagus labilis Erichson, 1846 Cryptophagus laticollis Lucas, 1846 Cryptophagus lycoperdi (Scopoli, 1763) Cryptophagus micaceus Rey, 1889 Cryptophagus pallidus Sturm, 1845 Cryptophagus pilosus Gyllenhal, 1827 Cryptophagus populi Paykull, 1800 Cryptophagus postpositus Sahlberg, 1903 Cryptophagus pseudodentatus Bruce, 1934 Cryptophagus pubescens Sturm, 1845 VCH 1907 2005 ++ ++ ++ Notes (Na) Increasing under bark of hardwoods and occasionally on pine. formerly placed in Cucujidae. Local, in sedge and marsh litter in wetlands. Also in tussocks. ? Introduced, Synanthropic. Local in compost and manure heaps. Introduced in stored products. Synanthropic, cosmopolitan. ++ ++ SUB VCH 1907 2005 ++ + ++ + + + + ++ + + ++ + ++ + ++ + + + + + (Nb) Local under bark, of hardwoods and pine. Local. Under bark of hard and softwoods. Notes Local on Carex in wetlands (RDBK) Rare? On lesser reedmace Typha latifolia on seed heads, also wintering in stems. (RDBK) Rare? On Sparganium and Typha, Local but frequent on Typha on flower heads and hiding in leaf sheaths. Rare. In flood debris suspended in trees, bridgework wire fences etc. ? Synanthropic, cosmopolitan Rare, saproxylic. Under bark, at blossoms. Bradfield (NHJ) ? Synanthropic, in cellars, warehouses etc. in mouldy organic matter. (N) [In pine woodland under bark etc.] Last recorded in 1907? (RDBIn) [In woodland in squirrel dreys, and bird’s nests, in birds nests and granaries] ? [Synanthropic in granaries, warehouse etc.] (RDBK) Rare in crumbly dead beech. Windsor High Standing Hill 1987 (ABD) Silwood Park 1998-9 (RGB) in FIT Local on dead wood, fungi and in buildings Local in fungi and haystack and other mouldy vegetable matter. Windsor Silwood Park 1998-9 (RGB) in FIT (N) Rare, in rotten wood. Silwood Park 1998-9 (RGB) in FIT ? lists. Local in Scleroderma fungi (RDBK) Rare in trees in debris including old wasp nests. Silwood Park 1998-9 (RGB) in FIT Local? In decaying organic matter including dead wood and fungi Local usually synanthropic in haystack refuse, grass clippings, and hotbeds (N) [Woods, fens and river banks in litter and decaying organic matter] ? Local? In decaying organic matter. Bradfield (NHJ) 55 Cryptophagus rotundatus Coombs & Woodroffe, 1955 Cryptophagus ruficornis Stephens, 1830 Cryptophagus saginatus Sturm, 1845 Cryptophagus scanicus (Linnaeus, 1758) + Cryptophagus scutellatus Newman, 1834 Cryptophagus setulosus Sturm, 1845 Cryptophagus subdepressus Gyllenhal, 1827 Micrambe abietis (Paykull, 1798) Micrambe vini (Panzer, 1797) Antherophagus canescens Grouvelle, 1916 Antherophagus nigricornis (Fabricius, 1787) Antherophagus pallens (Linnaeus, 1758) Caenoscelis ferruginea (Sahlberg, 1820) Caenoscelis subdeplanata Brisout, 1882 Atomaria apicalis Erichson, 1846 Atomaria atra (Herbst, 1793) Atomaria atricapilla Stephens, 1830 Atomaria basalis Erichson, 1846 + + Atomaria berolinensis Kraatz, 1853 Atomaria fuscata (Schoenherr, 1808) Atomaria fuscipes (Gyllenhal, 1808) Atomaria gutta Stephens, 1835 Atomaria lewisi Reitter, 1877 Atomaria mesomela (Herbst, 1792) Atomaria munda Erichson, 1846 Atomaria nigripennis (Kugelann, 1794) Atomaria nitidula (Marsham, 1802) Atomaria peltata Kraatz, 1853 Atomaria pusilla (Paykull, 1798) Atomaria rubida Reitter, 1875 Atomaria rubricollis Brisout, 1863 Atomaria scutellaris Motschulsky, 1849 Atomaria testacea Stephens, 1830 Atomaria zetterstedti (Zetterstedt, 1838) Atomaria barani Brisout, 1863 Atomaria punctithorax Reitter, 1887 Atomaria fimetarii (Fabricius, 1792) Atomaria nigrirostris Stephens 1830 Atomaria linearis Stephens, 1830 Atomaria lohsei Johnson & Strand, 1968 Atomaria nigriventris Stephens, 1830 Atomaria pulchra Erichson, 1846 + + + + + + + + + * + + + + + + + + * + + ++ + + + + ++ ++ ++ ++ ++ (N) Local on dead and fungoid wood especially in Daldinia Streatley (NHJ) ? Last recorded in 1924 Local in decaying organic matter in haystacks, in wasp nests and in carrion. Also synanthropic in houses, warehouses etc. Synanthropic in vegetable refuse. [Also in birds nests, and haystacks] ) Local in heathland litter [in rabbit burrows and Bumblebee nests] Last recorded in 1907 Bradfield (NHJ) Widespread on flowers of gorse Ulex. (Nb) [Breeds in nests of Bumblebees Bombus] Local, breeds in nests of Bombus Local, breeds in nests of Bombus Bradfield (NHJ) Rare? In woodland leaf litter at Bradfield (NHJ) Rare in woodlands. Silwood Park 1999 (RGB) Widespread and frequent in compost and other decaying vegetable matter (N) Extinct? [In wetlands] Bradfield Theale (NHJ) Blackwater River SU76 2000 (PHM) Widespread and frequent in compost and other decaying vegetable matter Extinct [In wetlands in litter]. Wellington College Local in decaying vegetable matter Widespread in compost and other decaying vegetable matter Rare [In dung and decaying vegetable matter] Rare? In wetlands in marsh litter Widespread in grass cuttings etc. Local in wetlands in marsh litter ? In straw and stable debris. (N) Extinct [In farmyard debris, cellars etc.] Widespread and frequent in most habitats. (RDBK) Extinct? [In farmyard debris] ? [In compost and farmyard debris] Rare, much declined [On dung esp. in or near woodlands] (RDBIn) ? In grassland. (RDBK) Increasing in grassy habitats. Basildon Park 2000 (KNAA) Widespread in litter, compost etc. frequent in gardens. (RDBK) Extinct? [In wetlands] (N) Rare? [In wetlands in litter] (N) Rare? [In decaying vegetable matter] in horse dung (N) Rare, declining ? In grassland esp. farmland, on shaggy ink-cap fungi Coprinus cornatus Widespread in woodland in litter and moss Local in dung, compost etc. (RDBK) Rare, on conifers in woodland. Silwood Park 1999 (RGB) (N) [Compost heaps, haystack refuse.] Little Wittenham 1998 (RGB) Wytham 1994 (RGB) Local on conifers in woodlands. Windosr (PJH) 1990 56 Atomaria puncticollis Thomson, 1868 Atomaria punctithorax Reitter, 1887 Atomaria strandi Johnson, 1967 Atomaria umbrina (Gyllenhal, 1827) Ootypus globosus (Waltl, 1838) Ephistemus globulus (Paykull, 1786) Biphyllidae (False hide beetles) ++ + Rom 2 species SUB Biphyllus lunatus (Fabricius, 1787) Diplocoelus fagi Guérin-Méneville, 1844 Byturidae (Raspberry beetles) 2 species SUB Byturus ochraceus (Scriba, 1790) Byturus tomentosus (Degeer, 1774) Raspberry beetle Erotylidae (Pleasing fungus beetles) 6 species SUB Triplax aenea (Schaller, 1783) Triplax russica (Linnaeus, 1758) Tritoma bipustulata Fabricius, 1775 Dacne bipustulata (Thunberg, 1781) Dacne rufifrons (Fabricius, 1775) Phalacridae (Shining mold beetles) 14 species Phalacrus brunnipes Brisout, 1863 Phalacrus caricis Sturm, 1807 Phalacrus corruscus (Panzer, 1796) Phalacrus fimetarius (Fabricius, 1775) Phalacrus substriatus Gyllenhal, 1813 Olibrus aeneus (Fabricius, 1792) Olibrus affinis (Sturm, 1807) Olibrus corticalis (Panzer, 1796) Olibrus liquidus Erichson, 1845 Olibrus millefolii (Paykull, 1800) Olibrus pygmaeus (Sturm, 1807) Stilbus oblongus (Erichson, 1845) Stilbus testaceus (Panzer, 1796) SUB ++ ++ (RDBK) Extinct? [On hardwoods trees] Silwood Park 1999 (RGB) (N) Rare,[In woodlands esp. on conifers] (N) ? [In woodland on fungi] Rare? In herbivore dung Widespread in decaying vegetation including hay, and compost heaps. VCH 1907 2005 ++ ++ Notes VCH 1907 2005 ++ ++ Notes VCH 1907 2005 ++ ++ ++ ++ ++ Notes VCH 1907 2005 Notes ++ ++ ++ + + + * > ++ Local in the fungi Daldina concentrica especially on ash. (Nb) V.local under bark of hardwoods especially on beech. Streatley (NHJ) Local esp. on blossoms in the spring Widespread on blossoms, larvae in fruits of Rubus and a pest of cultivated raspberry etc. Local, saproxylic. Under bark and on fungi. (Nb) ? [Saproxylic on old brackets and other fungi on hardwoods] Drayton Copse 1998 (JSD) (Na) Rare. On fungi and fungoid wood. Local. Saproxylic. Under bark and on fungi. Local. Saproxylic. Under bark and on fungi. (Na) Rare on Windsor Forest 1985 (PJH) Silwood Park 1998 (RGB) in FIT Local on smutted Carex ? Rare [on smutted cereal crops] , Local on smutted grasses [On smutted Carex] see Thomson (1958) Frequent on mayweeds, tansy and mugwort in rough grassy places Rare [mainly coastal] on goat’s-beard, & other yellow composites. ? 1995 Rare ? [On Matricaria] , Tubney (JJW) (Nb) ? [On yarrow Achillea] (Nb) [On common cudweed Filago vulgaris in marshy areas] Tubney (JJW0 Rare? On reedmace Typha latifolia at margins of water. Frequent in dry grassy debris 57 Bothrideidae 4 species SUB Teredus cylindricus (Olivier, 1790) VCH 1907 2005 ++ Oxylaemus variolosus (Dufour, 1843) Anommatus diecki Reitter, 1875 Anommatus duodecimstriatus (Müller, 1821) Cerylonidae 4 species Cerylon fagi Brisout, 1867 Cerylon ferrugineum Stephens, 1830 Cerylon histeroides (Fabricius, 1792) Murmidius ovalis (Beck, 1817) Corylophidae 7 species ++ + + SUB LBrA SUB Sericoderus lateralis (Gyllenhal, 1827) Corylophus cassidoides (Marsham, 1802) Orthoperus aequalis Sharp, 1885 Orthoperus atomarius (Heer, 1841) Orthoperus atomus (Gyllenhal, 1808) Orthoperus brunnipes (Gyllenhal, 1808) Orthoperus corticalis (Redtenbacher, 1849) Orthoperus mundus Matthews, 1885 Orthoperus nigrescens Stephens, 1829 Coccinellidae (Ladybirds) 46 spec Henosepilachna argus (Geoffroy, 1785) Bryony ladybird Hippodamia variegata Goeze Hippodamia tredecimpunctata (Linnaeus, 1758) 13-spot ladybird Vibidia duodecimguttata (Poda, 1761) Subcoccinella vigintiquattuorpunctata (Linnaeus, 1758) 24-spot ladybird. Coccidula rufa (Herbst, 1783) Coccidula scutellata (Herbst, 1783) Rom |rom (RDB1) Rare, saproxylic. In bark infested with Scolytids/Anobiids. Virginia Water 1970s (AAA) Formerly placed in Colydidae. (RDB3) At roots of old trees esp. oak. Bradfield (NHJ) Formerly placed in Colydidae. (RDBK) Subterranean. Local but drastically under-recorded unless deliberately trapped for (see Owen, 1999) (Na) Subterranean, as above species (see Owen, 1999) VCH 1907 2005 ++ ++ ++ Notes VCH 1907 2005 ++ + + * + Notes + + SUB Notes VCH 1907 2005 + (Na) Rare, under bark of hardwoods esp. beech and oak Local under bark of hardwoods esp. beech and oak Local under bark of hardwoods esp. beech and oak Extinct (last UK record 1831). Formerly placed in Bothridiidae. Widespread in compost heaps under bark etc. Local in in rotting grass etc. (RDBK) Rare in timber, and on foliage of trees in ancient woodlands. Bradfield (NHJ) Rare on trees in ancient woodlands. . (RDB3) [Marsh litter in wetlands] ? ? Compost heaps. [Also comes to MV light]. Bradfield (NHJ) (Nb) Rare in woods [and a garden] Notes Recent colonist. ++ (Nb) Frilford. Theale lake (JSD) (RDBK) Extinct? Though scattered sighting in UK in recent warm summers! So may reappear. ~* + 1816 Ubiquitous ++ ++ Widespread in wetlands, on tall emergent waterside vegetation V.local in wetlands, often amongst Typha 58 Rhyzobius chrysomeloides (Herbst) Rhyzobius litura (Fabricius, 1787) Clitostethus arcuatus (Rossi, 1794) ++ ++ + Recent immigrant, Increasing on trees and bushes. Widespread and frequent, usually terrestrial (RDB1) Rare. Stethorus punctillum (Weise, 1891) Scymnus auritus Thunberg, 1795 Scymnus femoralis (Gyllenhal, 1827) Scymnus frontalis (Fabricius, 1787) Scymnus haemorrhoidalis Herbst, 1797 Scymnus limbatus Stephens, 1832 Scymnus nigrinus Kugelann, 1794 Scymnus schmidti Fürsch, 1958 Scymnus suturalis Thunberg, 1795 Nephus quadrimaculatus (Herbst, 1783) Nephus redtenbacheri (Mulsant, 1846) Hyperaspis pseudopustulata Mulsant, 1853 Platynaspis luteorubra (Goeze, 1777) Chilocorus bipustulatus (Linnaeus, 1758) Heather ladybird Chilocorus renipustulatus (Scriba, 1791) Kidney-spot ladybird Exochomus nigromaculatus (Goeze, 1777) Exochomus quadripustulatus (Linnaeus, 1758) Pine ladybird Anisosticta novemdecimpunctata (Linnaeus,1758) Water ladybird Aphidecta obliterata (L.innaeus, 1758) Larch ladybird Tytthaspis sedecimpunctata (Linnaeus, 1758) 16-spot ladybird Adalia bipunctata (Linnaeus, 1758) 2-spot ladybird Adalia decempunctata (Linnaeus, 1758) 10-spot ladybird Coccinella magnifica Redtenbacher, 1843 Scarce 7-spot ladybird Coccinella hieroglyphica Linnaeus, 1758 Hieroglyphic ladybird Coccinella septempunctata Linnaeus, 1758 7-spot ladybird Coccinella undecimpunctata Linaneus, 1758 11-spot ladybird Harmonia axyridis Harlequin ladybird Harmonia quadripunctata (Pontoppidan, 1763) ++ ++ + ++ ++ + + ++ + ++ ++ + + ++ V.local on trees and shrubs Local on foliage of oak trees (Nb) Rare. Local on ground esp in sandy areas. Local in grassland (Nb) Rare on sallows and willows. Very local on young pines (Nb) Rare on , and heaths, usually on the ground. Also can be frequent in FIT in woodland situations! Local on pines (RDB2) Local, increasing. On ivy Calcott 2003 (JSD) Local in grassland (Nb) (Na) Rare? Overlooked? In open grassy places with the ant Lasius niger sens str. Local on heathland and junipers on the ++ Widespread, on various trees esp. ash, alder and sallow. * ++ Ubiquitous on trees including ash, sycamore and confiers. ++ Local in wetlands ++ Local on larch, firs and occasionally on pines. ++ Widespread and abundant in grassland ++ Widespread and common esp. in suburbia. ++ Ubiquitous ++? (Na) Local around the nests of the wood ant Formica rufa ++ Local on heathland ++ Ubiquitous ++ Local, on low vegetation on sandy soils ++ Introduced asian speices Battersea ++ Local on pines Pinus, Rom Rom Rom Rom 59 Cream streaked ladybird Myrrha octodecimguttata (Linnaeus, 1758) 18-spot ladybird Calvia quattuorodecimguttata (Linnaeus, 1758) Cream spot ladybird Propylea quattuordecimpunctata (Linnaeus, 1758) 14-spot ladybird Rom ++ Local on pines ++ Local on Ash, Lime and other broadleaved trees ++ Ubiquitous 60 Myzia oblongoguttata (Linnaeus, 1758) Striped ladybird Anatis ocellata (Linnaeus, 1758) Eyed ladybird Halyzia sedecimguttata (Linnaeus, 1758) Orange ladybird Psyllobora vigintiduopunctata (Linnaeus, 1758) 22-spot ladybird Sphaerosomatidae 1 species SUB VCH Sphaerosoma piliferum (Müller, P.W.J., 1821) Endomychidae 6 species SUB Holoparamecus caularum (Aubé, 1843) Endomychus coccineus (Linnaeus, 1758) False ladybird Lycoperdina bovistae (Fabricius, 1792) Mycetaea hirta (Marsham, 1802) Symbiotes latus Redtenbacher, 1849 Lathridiidae (Plaster beetles) 38 species Lithostygnus serripennis Broun, 1914 Stephostethus angusticollis (Gyllenhal, 1827) Stephostethus lardarius (Degeer, 1775) Aridius bifasciatus (Reitter, 1877) Aridius nodifer (Westwood, 1839) Cartodere constricta (Gyllenhal, 1827) Lathridius anthracinus Mannerheim, 1844 Lathridius consimilis Mannerheim, 1844 Lathridius pseudominutus (Strand, 1958) Thes bergrothi (Reitter, 1880) Enicmus brevicornis (Mannerheim, 1844) Enicmus histrio Joy & Tomlin, 1910 Enicmus rugosus (Herbst, 1793) Enicmus testaceus (Stephens, 1830) Enicmus transversus (Olivier, 1790) Rom VCH ++ Local on scot’s pine on western heaths. ++ ++ Local on scot’s pine Widespread and increasing on mildewed leaves of hardwood trees. ++ Widespread and common 1907 2005 + Notes 1907 2005 ++ ++ Notes * ++ ++ VCH 1907 2005 + + ESax + ++ ++ + + ++ + + ++ + ++ + Ro/S ++ ax SUB Local. Synanthropic in compost heaps etc. Local. Synanthropic in manure and compost heaps etc. Formerly placed in Merophysidae Local on fungoid wood esp. beech and birch. (RDB3) Rare in puffballs formerly Tubney (JJW) Local in fungoid wood, compost heaps rotting hay/straw (Nb) Rare in fungoid wood. Bradfield (NHJ) Notes Synanthropic Rare in decaying plant debris Widespread and eurytopic, including gardens. [Also comes to MV light] Ubiquitous, since first appearance in 1950 Frequent in fungi and litter Local on trees. ? Woodland. . Local? ? Woodland. (N) [In fungi in old woods] Silwood Park 1998 (RGB) in FIT ? ? (N) Local under bark, esp. beech, and sycamore with sooty bark disease. Local, in litter including grassheaps and old haystacks. (N) Rare in old woodlands on slime moulds and fungi. Local in woodlands Widespread in grasslands 61 Dienerella elongata (Curtis, 1830) Dienerella filiformis (Gyllenhal, 1827) Dienerella filum (Aubé, 1850) Dienerella ruficollis (Marsham, 1802) Dienerella separanda (Reitter, 1887) Adistemia watsoni (Wollaston, 1871) Corticaria alleni Johnson, 1974 Corticaria crenulata (Gyllenhal, 1827) Corticaria dubia Dajoz, 1970 Corticaria elongata (Gyllenhal, 1827) Corticaria ferruginea Marsham, 1802 Corticaria fulva (Comolli, 1837) Corticaria impressa (Olivier, 1790) Corticaria inconspicua Wollaston, 1860 Corticaria longicollis (Zetterstedt, 1838) Corticaria punctulata Marsham, 1802 + + + + + ++ + + * + Ro/S ax Corticaria serrata (Paykull, 1798) Corticaria umbilicata (Beck, 1817) Corticarina fuscula (Gyllenhal, 1827) Corticarina lambiana (Sharp, 1910) Corticarina similata (Gyllenhal, 1827) Cortinicara gibbosa (Herbst, 1793) Melanopththalma suturalis Mannerheim, 1844 Ciidae (Minute tree fungus beetles) 16 species Octotemnus glabriculus (Gyllenhal, 1827) Sulcacis affinis (Gyllenhal, 1827) Sulcacis bicornis (Mellié, 1848) Cis alni Gyllenhal, 1813 Cis coluber Abeille, 1874 Cis festivus (Panzer, 1793) Cis pygmaeus (Marsham, 1802) Cis vestitus Mellié, 1848 Cis bidentatus (Olivier, 1790) Cis bilamellatus Wood, 1884 Cis boleti (Scopoli, 1763) Cis fagi Waltl, 1839 Cis hispidus (Paykull, 1798) Cis micans (Fabricius, 1792) Cis nitidus (Fabricius, 1792) Cis setiger Mellié, 1848 ++ + ++ ++ SUB VCH 1907 2005 ++ ++ ++ ++ ++ ++ ++ ++ > ++ ++ ++ ++ ++ ++ ++ Local? Woodland. ? Synanthropic. Synanthropic. Widespread in grass heaps etc. Widespread in garss heaps etc. Synanthropic. In fungoid materials (N) Rare. Saproxylic in rotting hardwood. Hutchin’s Copse 1998 (JSD) in Lasius brunneus nest in old oak ? Rare ? In compost heaps etc. ? ? Bradfield (NHJ) Widespread and eurytopic in open habitats in moss, litter, (N) [Eurytopic and synanthropic] (RDBK) ? [Riparian] ? ? Widespread and eurytopic in moss, litter etc. (RDB1) [Old woodlands] Rare? Widespread and abundant in most habitats in moss, litter also on tree blossoms. Notes Frequent in bracket fungi esp. Trametes esp. in beech woods. Local in fungi esp. Trametes versicolor (Nb) Rare in soft brackets Trametes etc. Sulhamstead Wood 2003 (JSD) Local on fungi and in fungoid hardwood, esp. on jew’s Ear Fungus Auricularia auricula-judae esp. on Elder RDB3 Windsor ?? (Nb) Local in bracket fungi on Sallow, Aspen and birch. Local on oak under bark and on fungoid wood Local on dead oak branches [also in Polypore fungi on elm and beech] Rare, in bracket fungi. Naturalised, Australasian, appeared 20th Century. Local in Birch Polypore Piptoporus betulinus. Frequent in fungi and fungoid wood esp in soft brackets. Rare in fungoid wood esp. red rotten oak. Local on bracket fungi Trametes spp. on hardwoods. Local on fungi on hardwoods, esp. oak. Local on fungi on hardwoods Rare in bracket fungi and fungoid wood. 62 Ennearthron cornutum (Gyllenhal, 1827) Mycetophagidae (Hairy Fungus beetles) 10 species Pseudotriphyllus suturalis (Fabricius, 1801) Triphyllus bicolor (Fabricius, 1777) Litargus connexus (Fourcroy, 1785) Mycetophagus atomarius (Fabricius, 1787) Mycetophagus multipunctatus Fabricius, 1792 Mycetophagus piceus (Fabricius, 1777) Mycetophagus quadriguttatus Müller, 1821 Mycetophagus quadripustulatus (Linnaeus, 1761) Eulagius filicornis (Reitter, 1887) * Typhaea stercorea (Linnaeus, 1758) Hairy fungus beetle Colydidae (Cylindrical Bark beetles) 11 sp Aglenus brunneus (Gyllenhal, 1813) Orthocerus clavicornis (Linnaeus, 1758) Synchita humeralis (Fabricius, 1792) Synchita separanda (Reitter, 1882) Cicones undatus Guérin-Méneville, 1829 Cicones variegatus (Hellwig, 1792) Bitoma crenata (Fabricius, 1775) Langelandia anophthalma Aubé, 1843 Colydium elongatum (Fabricius, 1787) Aulonium trisulcum (Fourcroy, 1785) Pycnomerus fuliginosus Erichson, 1842 Tenebrionidae (Darkling beetles) 29 species + 1 sub-fossil Eledona agricola (Herbst, 1783) Scaphidema metallicum (Fabricius, 1792) Alphitophagus bifasciatus (Say, 1823) Two-banded fungus beetle Gnatocerus cornutus (Fabricius, 1798) Broad-horned flour beetle Tribolium castaneum (Herbst, 1797) Rust-red flour beetle ++ SUB VCH ++ ++ LRom SUB 1907 2005 ++ ++ ++ ++ ++ ++ ++ ++ VCH LRom ESax 1907 2005 + ++ ++ ++ ++ ++ ++ ++ + ++ SUB VCH 1907 2005 ++ ++ Widespread on fungi and fungoid wood. Notes Local in fungi and fungoid hardwoods (Nb) Rare in fungi and fungoid hardwoods. Local in fungi and fungoid wood including gorse. Bradfield (NHJ) Local in fungi on trees, and under bark. Local in fungi on trees, and under bark. (Nb) Local in fungi on trees, and under bark. (Na) Rare in fungi on trees, and under bark. ,Branbourne Park, 1988 (ABDrane) Local in fungi on trees, and under bark. Naturalised, on fungi and fungoid hardwood Reading TDH) Southcote 2004 (JSD) Widespread in haystack debris and occasionally under hardwood bark. Notes 1975 (Nb) Extinct? [Sandy ground with Peltigera lichens] Hitchcopse Pit (DJM) (Nb) Local on fungoid wood. (RDB3) V.local increasing under bark esp. Local, increasing under bark, esp. Sycamore with sooty bark disease. (Na) Rare on fungi and dry deaying fungoid hardwoods Bradfield (NHJ) Widespread under bark of hardwoods, and conifers. (RDB3) Local but increasing? Subterranean in gardens and parks >20 sites (RDB3) Local but increasing? In dead under bark esp. beech, birch & oak etc. (Na) Rare, declining. On elm predating Scolytids in bark. Naturalised, Australasian species. Increasingly common under bark incl. pine. Spreading esp.in west of county Notes (Nb) Local in fungi, usually Chicken-of-the-woods Laetiporus sulphureus esp. on oak and cherry. (Nb) Local on fungoid wood esp. elder. Also in old stables. [Synanthropic in stored grain etc., also in fungi, compost and haystack refuse] [Synanthropic in stored grain etc.] ++ [Synanthropic in stored grain etc.] 63 Tribolium confusum du Val, 1863 Confused flour beetle Latheticus oryzae Waterhouse, 1880 Palorus ratzeburgi (Wissmann, 1848) Small-eyed flour beetle Alphitobius diaperinus (Panzer, 1796) Lesser mealworm beetle Alphitobius laevigatus (Fabricius, 1781) Black fungus beetle Corticeus bicolor (Olivier, 1790) Corticeus fraxini (Kugelann, 1794) Corticeus linearis (Fabricius, 1790) Myrmechixenus vaporariorum Guérin-Méneville, 1843 Blaps lethifera Marsham, 1802 Blaps mucronata Latreille, 1804 Churchyard or Cellar beetle Tenebrio molitor Linnaeus, 1758 Yellow mealworm beetle Tenebrio obscurus Fabricius, 1792 Dark mealworm beetle Helops caeruleus (Linnaeus, 1758) Nalassus (=Cylindrinotus) laevioctostriatus (Goeze, 1777) False ground beetle! Opatrum sabulosum (Linnaeus, 1758) Melanimon tibialis (Fabricius, 1781) Crypticus quisquilius (Linnaeus, 1761) Prionychus ater (Fabricius, 1775) Prionychus melanarius (Germar, 1813) Gonodera luperus (Herbst, 1783) Pseudocistela ceramboides (Linnaeus, 1758) Isomira murina (Linnaeus, 1758) Mycetochara humeralis (Fabricius, 1787) Cteniopus sulphureus (Linnaeus, 1758) Lagria hirta (Linnaeus, 1758) Tetratomidae 3 species [Synanthropic in stored grain etc.] [Synanthropic in stored grain etc.] [Synanthropic in stored grain etc.] LRom Local, in Manure heaps also on fungi in wild, also comes readily to MV light. + Local in manure heaps esp. deep litter debris from chicken farms + > + + Rare under ash and elm bark with Scolytids esp. Scolytus multistriatus & S. scolytus in elm bark. Local under pine bark, with Scolytids esp. Ips sexdentatus. Rare under pine bark with Pityogenes bidentatus. (RDB3) Rare in compost heaps and grass cuttings. . Formerly placed in the Colydidae. ++ Extinct? [Synanthropic] Synathropic in cellars and stables, declining? ++ Synanthropic. In stored flour etc. Often attracted to light. ++ Synanthropic in stored flour etc. also in stables ++ (Nb) ? [Saproxylic in a variety of dead wood esp. oak] Frequent esp. under bark on trees esp. pine, also on heather on open heath. Grazes Pleurococcus algae. ++ ++ ++ ++ ++ ++ ++ ++ ++ ++ SUB Tetratoma ancora Fabricius, 1791 Tetratoma desmaresti Latreille, 1807 Tetratoma fungorum Fabricius, 1790 Salpingidae (Narrow-waisted bark beetles) 9 species ++ SUB (Nb) Rare on disturbed sandy ground at roots of plants. V.local in open disturbed sandy areas on heathland. Frilford Tubney on Corallian sands (Nb) Local in rotting wood esp. red-rotten oak. (RDB2) [Saproxylic] Local, Saproxylic. Adults on vegetation in rides and woodland edge habitats. (Nb) Rare, saproxylic on hardwoods, comes to light. Widespread in woods and commons on flowers. (Na) Rare, Saproxylic on oak etc. Windsor Ffrilford on the Corallian (JSD etc Local on hedgerows, also in open situations in grassland VCH 1907 2005 + ++ ++ Notes VCH 1907 - Notes (Nb) ? [On oak, hazel etc., on branches encrusted in canker fungi Phlebia merismoides, and other fungi]. (Na) Rare on shaded oak boughs on old trees. Local on fungi and fungoid wood, including birch polypore Piptoporus betulinus. 64 2005 * ++ Lissodema cursor (Gyllenhal, 1813) Lissodema quadripustulata (Marsh, 1802) Rabocerus gabrieli Gerhardt, 1901 Salpingus castaneus (Panzer, 1796) Salpingus reyi (Abeille, 1874) Vincenzellus ruficollis (Panzer, 1794) Rhinosimus planirostris (Fabricius, 1787) Rhinosimus ruficollis (Linnaeus, 1761) Pyrochroidae (Cardinal beetles) 2 species + + ++ + + SUB Pyrochroa coccinea (Linnaeus, 1761) Black-headed cardinal Pyrochroa serraticornis (Scopoli, 1763) Red-headed cardinal beetle Rom Melandryidae (False darkling beetles) 13 species SUB ++ Hallomenus binotatus (Quensel, 1790) Orchesia micans (Panzer, 1793) Orchesia minor Walker, 1836 Orchesia undulata Kraatz, 1853 Anisoxya fuscula (Illiger, 1798) Abdera biflexuosa (Curtis, 1829) Abdera flexuosa (Paykull, 1799) * Abdera quadrifasciata (Curtis, 1829) Phloiotrya vaudoueri Mulsant, 1856 Melandrya caraboides (Linnaeus, 1761) Conopalpus testaceus (Olivier, 1790) Osphya bipunctata (Fabricius, 1775) Scraptidae 12 species Scraptia dubia Scraptia fuscula Müller, 1821 Scraptia testacea Allen, 1940 Anaspis costai Emery, 1876 Anaspis frontalis (Linnaeus, 1758) Anaspis garneysi Fowler, 1889 Anaspis humeralis (Fabricius, 1775) Anaspis lurida Stephens, 1832 VCH 1907 2005 ++ SUB (Na) ? [Saproxylic, on Ash Fraxinus, in dying tips of branches high up on trees] (Nb) Local on dead and dying branches of hardwoods esp. hawthorn and maple (Nb) Rare on dead wood including pine and beech. Local on dead and moribund conifers. Local in burnt twigs especially birch on heathland after fires. Local on dead and moribund hardwoods, especially hawthorn and beech Widespread on dead and moribund hardwoods Local on dead and moribund hardwoods Notes (Nb) Widespread, much increased in 20th C? larvae under bark of hardwoods (and conifers?). Widespread, larvae under bark of hardwoods and conifers. >50 records VCH 1907 2005 ++ ++ ++ ++ ++ ++ ++ ++ ++ ++ ++ ++ Notes VCH Notes 1907 2005 + ++ ++ ++ ++ ++ ++ ++ (Nb) V.local in bracket fungi, and fungoid wood incl. Pine. (Nb) V.local in fungi and fungoid wood including Inonotus radiatus on Alder. (Nb) V.local in fungi and fungoid wood Local in fungi and fungoid wood and decaying bracket fungi. Apparently very rare in 19th C!! (Na) Local on dead and dying branches of hardwoods esp. Salix and Populus (Nb) Local on dead and dying boughs of oak on large trees (Nb) Rare on bracket fungi Inonotus radiatus on Alder, at Sheffield Bottom 2003 (JSD) (Na) Local on dead and dying boughs of oak on large trees (Nb) V.local on fungoid hardwood boughs esp of beech and oak. (Nb) Local on hardwoods esp. oak. (Nb) Local on dead and moribund oak boughs on large trees. (RDB3) Rare saproxylic Windsor Forest, Hutchin’s Copes 1998 (JSD) Windspr in 19th C (RDB1) possibly the first definite record of this species in Sy. (RDB3) Rare on old oak trees in parkland esp. those with stag’s horn die back. Rare, saproxylic, adults at blossoms. Widespread and frequent, saproxylic, adults on blossoms esp. hawthorn, sloe etc Local, adults on blossoms esp. hawthorn, sloe etc Widespread and frequent, saproxylic, adults on blossoms esp. hawthorn, sloe etc. Local, saproxylic, adults on blossoms esp. hawthorn, sloe etc. 65 Anaspis maculata Fourcroy, 1785 Anaspis pulicaria Costa, 1854 Anaspis regimbarti Schilsky, 1895 Anaspis rufilabris (Gyllenhal, 1827) Anaspis thoracica (Linnaeus, 1758) Mordellidae (Tumbling Flower beetles) 13 species ++ ++ ++ ++ ++ SUB VCH Tomoxia bucephala Costa, 1854 Variimorda villosa (Schrank, 1781) 1907 2005 ++ ++ Mordella holomelaena Apfelbeck, 1914 Mordellistena acuticollis Schilsky, 1895 + ++ Mordellistena brevicauda (Boheman, 1849) Mordellistena humeralis (Linnaeus, 1758) Mordellistena neuwaldeggiana (Panzer, 1796) Mordellistena parvula (Gyllenhal, 1827) * Mordellistena pumila (Gyllenhal, 1810) Mordellistena pseudoparvula Ermisch, 1956 Mordellistena secreta Horák, 1983 Mordellistena variegata (Fabricius, 1798) Mordellochroa abdominalis (Fabricius, 1775) + ++ ++ ++ ++ + + + ++ Rhipiphoridae (Wasp-nest beetles) 1 species SUB VCH Metoecus paradoxus (Linnaeus, 1761) Oedemeridae (False blister beetles) 7 species recorded Nacerdes melanura (Linnaeus, 1758) Wharfborer Ischnomera caerulea (Linnaeus, 1758) Ischnomera cyanea (Fabricius1787) Ischnomera sanguinicollis (Fabricius, 1787) Oncomera femorata (Fabricius, 1792) Oedemera lurida (Marsham, 1802) Oedemera nobilis (Scopoli, 1763) Thick-kneed beetle SUB Rom VCH Widespread and often abundant on blossoms esp. hawthorn, sloe etc. Local, adults on blossoms esp. hawthorn, sloe etc. Widespread, adults on blossoms esp. hawthorn, sloe etc. Widespread, adults on blossoms esp. hawthorn, sloe etc. (Na) Rare, saproxylic adults on oak boughs, also at blossoms. Notes (Na) Local, saproxylic on a variety of trees including pine. (Nb) Local in woods, parks, and open grassland. Saproxylic: Females seen egg-laying in old beech & pine stumps (JSD)! (RDBK) Rare, in woodland, adults often on flowers esp. umbels. Recent immigrant?, first recorded in UK in 1985, now widespread on mugwort Artemesia vulgaris (Levey, 2002). (RDBK) Rare on chalk downland. (RDBK) Rare on hedgerows and woods. , (RDBK) Local, increasing, on a variety of hardwoods in small branches >20+ post 1980 records (RDBK) Rare [On yarrow? In open warm areas on grassland] Heron’s Nest 2003 (JSD) Local in open grasslands esp. on the (RDBK) ? Recently recognised spp. ) V.local, saproxylic in hardwood twigs etc. Local, saproxylic, adults on blossoms esp. hawthorn, sloe etc. 1907 2005 ++ Notes 1907 2005 + Notes + ++ + + ++ ++ Local in nests of social wasps Vespidae, sometimes in houses. Rare, casual import? Primarily coastal on timber in or by salt water. (RDB3) Rare, in decaying elm and oak, adults on blossoms. (Nb) V.local, in decaying hardwoods, adults at blossoms. (Nb) Rare, develops in old soft elm. Adults on blossoms. 1 (Nb) Rare in woods, saproxylic. Comes to light. Widespread and locally abundant in grasslands and rides. Widespread and locally abundant in grasslands and rides. 66 Meloidae (Oil beetles) 4 species SUB VCH 1907 2005 Notes Lytta vesicatoria (Linnaeus, 1758) Meloe proscarabaeus Linnaeus, 1758 + Meloe rugosus Marsham, 1802 Meloe violaceus Marsham, 1802 Anthicidae 6 species Notoxus monoceros (Linnaeus, 1761) Anthicus antherinus (Linnaeus, 1761) Anthicus bifasciatus (Rossi, 1792) Anthicus floralis (Linnaeus, 1758) Anthicus formicarius (Goeze, 1777) Anthicus tobias Marseul, 1879 Aderidae 3 species SUB Rom SUB Aderus brevicornis (Perris, 1869) Aderus oculatus (Paykull, 1798) Aderus populneus (Creutzer, 1796) Cerambycidae (Longhorns) 52 species Prionus coriarius (Linnaeus, 1758) The Tanner Arhopalus rusticus (Linnaeus, 1758) Dusky longhorn Arhopalus tristis (Fabricius, 1787) Asemum striatum (Linnaeus, 1758) House longhorn Tetropium gabrieli Weise, 1905 Rhagium bifasciatum Fabricius, 1775 Rhagium mordax (Degeer, 1775) Stenocorus meridianus (Linnaeus, 1758) Grammoptera ruficornis (Fabricius, 1781) Grammoptera ustulata (Schaller, 1783) Grammoptera abdominalis (Stephens, 1831) Pseudostrangalia (=Leptura) revestita (Linnaeus, 1767) Leptura (=Strangalia) quadrifasciata (Linnaeus, 1758) SUB Extinct? Hitchcopse Pit (DM) (Nb) Extinct? Last recorded in in early 20th C when reported to be common! VCH 1907 2005 ++ ++ + ++ ++ ++ Notes VCH 1907 2005 ++ ++ ++ Notes VCH Notes 19072005 ++ ++ Local on open disturbed areas on sandy soils. Frilford (JMC det JSD) Widespread. Synanthropic in compost heaps etc. (Nb) Rare in compost and manure heaps. Widespread and frequent in grass and compost heaps, and in haystack debris. Widespread in dung heaps, compost and grass heaps. Synanthropic on tips, waste ground etc. (RDB2) Extinct?? Saproxylic [In fungi and fungoid wood esp. oak, beech and elm] Windsor (JAO etc ?? (Nb) V.local on oaks, larvae in red-rot, adults on foliage [adults favour Elder Sambucus blossom] (Nb) V.local on oaks, adults often swarming on foliage of outer branches [adults at hawthorn blossom, and Salix catkins] (Na) Local, saproxylic in roots of hardwoods and pine, nocturnal, and comes to light. Local, in conifers esp. pine, much increased since 1950s and largely replaced A.tristis ++ + Rare, much declined in conifers esp. pine, Immigrant in England, increasing? Local in conifers esp. pine attracted to recently cut or felled timber > ++ ++ ++ ++ ++ ++ * Rare? Naturalised. In conifers esp. larch including felled timber. Frequent in woods and wooded heaths esp. in pine. Widespread in hardwoods esp. oak and ash. Local in woodlands, illusive despite size. Breeds in a variety of hardwoods. Frequent, adults common on blossoms esp. hawthorn, sloe etc. (RDB3) Rare on hardwoods in ancient woods and parkland. (Na) Rare though regular at sites like and Commons. (RDB1) Rare. + Local, commonest on wooded heaths, develops in deaying wood and stumps esp. birch. 67 Four-banded longhorn Stictoleptura (=Leptura) scutellata Fabricius, 1781 Pachytodes (=Judolia) cerambyciformis (Schrank, 1781) Alosterna tabacicolor (Degeer, 1775) Pseudovadonia (=Leptura) livida Fabricius, 1777 Rutpela (=Strangalia) maculata (Poda, 1761) Stenurella (= Strangalia) melanura (Linnaeus, 1758) Stenurella (= Strangalia) nigra (Linnaeus, 1758) Gracilia minuta (Fabricius, 1781) Basket longhorn Nathrius brevipennis (Mulsant, 1839) Molorchus minor (Linnaeus, 1758) Glaphyra (= Molorchus) umbellatarum (von Schreber, 1759) Aromia moschata (Linnaeus, 1758) Musk beetle Hylotrupes bajulus (Linnaeus, 1758) House longhorn Callidium violaceum (Linnaeus, 1758) Violet longhorn Poecilium (=Phymatodes) alni (Linnaeus, 1767) Phymatodes lividus Phymatodes testaceus (Linnaeus, 1758) Oak longhorn Clytus arietis (Linnaeus, 1758) Wasp beetle Anaglyptus mysticus (Linnaeus, 1758) Monochamus sutor Linnaeus, 1758 Mesosa nebulosa (Fabricius, 1781) Pogonocherus fasciculatus (Degeer, 1775) * Pogonocherus hispidulus (Piller & Mitterpacher, 1783 Pogonocherus hispidus (Linnaeus, 1758) Leiopus nebulosus (Linnaeus, 1758) Agapanthia villosoviridescens (Degeer, 1775) Saperda carcharias (L., 1758) Saperda populnea (Linnaeus, 1758) Small poplar borer Stenostola dubia (Laicharting, 1784) Lime longhorn Phytoecia cylindrica (Linnaeus, 1758) Tetrops praeusta (Linnaeus, 1758) Plum longhorn ++ (Na) Rare, increasing? Larvae in beech, oak, and birch. ++ V.local, on flowers in woodlands ++ ++ ++ ++ Local but widespread adults on flowers esp. umbels V.local, adults at flowers esp. umbels. Frequent at blossoms, breeding in hardwoods Local, but widespread perhaps commonest on wooded heaths + (Na) Rare in ancient woodland. (RDB2) ? In dry twigs and branches including bramble, and wickerwork. Much declined in 20 th C but may be recovering in last few years with recent records from Kent and Essex? Imported species, established in wide variety of timber. . Naturalised, Rare in a variety of trees esp. spruce? Adults visit blossom. (Na) Rare, breeds in dead and dying woody Rosaceae esp. hawthorn, typically on sickly looking specimens. Cow Common1998 (JSD) (Nb) Rare in willows near water. Rare in timber in houses. + ++ ++ + ? [In a variety of trees] ++ + LRom ++ ++ ++ ++ ++ ++ (Nb) Rare, declining. Introduced in wood in cooperage 1894-1905 in Reading V.local in variety of hardwoods, including Sycamore, Sweet Chestnut. Often occurs in untreated fence posts. Widespread and frequent in woods and gardens (Nb) Local, in woods and hedgeows, adults on blossoms esp. hawthorn. Alien, accidentally introduced but not established. (RDB3) Rare only modern record from Fisherlane woods (Nb) Recent immigrant/intro on pine. Bagshott 2003 (MVB) Local on wide variety of trees. ++ ++ ++ * ++ Local on a wide variety of trees, adults esp. on holly Local usually on oak preferring moribund / dead but attached branches of large trees ? [In thistles and umbellifers, increasing southward?] Dinton Pastures (GAC) Sheffield Bottom 2003 (JSD) + (Nb) Rare on lime Tilia trees in woods and parkland [comes to ligh traps] ++ ++ (Nb) V.local breeds in stems of umbellifers. Local, breeds on woody Rosaceae esp. apple, and possibly in other hardwoods. V.local on aspen on damp heathland and woodland rides 68 Bruchidae (Bean weevils) 11 species SUB Bruchus atomarius (Linnaeus, 1761) Bruchus loti Paykull, 1800 Bruchus pisorum (Linnaeus, 1758) Bruchus rufimanus Boheman, 1833 Bruchus rufipes Herbst, 1783 Bruchidius cisti (Fabricius, 1775) Bruchidius olivaceus (Germar, 1824) Bruchidius varius (Olivier, 1795) Bruchidius villosus (Fabricius, 1792) Acanthoscelides obtectus (Say, 1831) Callosobruchus chinensis (Linnaeus, 1758) Chrysomelidae (Leaf Beetles) 214 species + 3 sub-fossil Macroplea appendiculata (Panzer, 1794) Donacia aquatica (Linnaeus, 1758) Donacia bicolora Zschach, 1788 Donacia cinerea Herbst, 1784 Donacia clavipes Fabricius, 1792 Donacia crassipes Fabricius, 1775 Donacia dentata Hoppe, 1795 Donacia impressa Paykull, 1799 Donacia marginata Hoppe, 1795 Donacia obscura Gyllenhal, 1813 * Donacia semicuprea Panzer, 1796 Donacia simplex Fabricius, 1775 Donacia sparganii Ahrens, 1810 Donacia thalassina Germar, 1811 Donacia versicolorea (Brahm, 1791) Donacia vulgaris Zschach, 1788 Plateumaris affinis (Kunze, 1818) Plateumaris braccata (Scopoli, 1772) Plateumaris discolor (Panzer, 1795) Plateumaris sericea (Linnaeus, 1758) Orsodacne lineola (Panzer, 1795) Zeugophora subspinosa (Fabricius, 1781) Lema cyanella (Linnaeus, 1758) Oulema obscura (Stephens, 1831) = lichenis (Voet, 1806) Oulema melanopus (Linnaeus, 1758) Oulema rufocyanea (Suffrian) VCH 1907 2005 > ++ ++ ++ ++ + ++ ++ SUB Sax Sax VCH 19072005 Notes (Nb) ? On vetches Vicia, Lathyrus. Local on Meadow Vetchling Lathyrus pratensis ? On imported beans and pulses. Local on various legumes including cultivated beans Widespread on various vetches Vicia, and possibly clovers Trifolium spp. Rare, adults on rock-rose Helianthemum on , larvae in Lotus?? (RDB1) ? Cothill 1923 Recent immigrant, now widespread and well established on red clover. Local, but usually present wherever decent stands of Broom Cytisus grow. ? On stored beans ? Imported with beans, & lentils Notes + + + ++ ++ + + ++ ++ ++ ++ ++ ++ + + ++ + (RDB3) (RDB3) Extinct? Last recorded at 1911 [On Carex in rich wetlands] (RDB2) Local on Sparganium erectum at margins of (Nb) V.local on Typha. (Nb) V.local on reeds Phragmites at margins of lakes and ponds. Searles lane pits 2003 (JSD) (Nb) Rare on white lily Nymphaea alba, no modern record (Na) Extinct? [on arrowhead Sagittaria] (Na) Rare, on Scirpus at margins of lakes and ponds. Local on Sparganium esp. beside still water. (Na) Rare on bottle sedge Carex rostrata in bog pools. Local on Glyceria maxima at water margins esp. beside flowing water. Widespread on Sparganium at margins of ponds, lakes, rivers and canals (Na) Extinct? [On Sparganium] (Nb) Rare on Carex Local on broad-leaved pondweed Potamogeton natans in ponds and pools. Widespread on Typha, Sparganium etc. (Nb) V.local on Carex at margins of ponds and streams (Na) On Phragmites often hiding in the leaf sheaths Local in wetlands in acidic areas including bogs Local in wetlands with richer waters, fens and weedy pond margins. (Nb) Rare on Hawthorn and allied Rosaceae Local on Aspen Populus tremulae, and Willows Salix. Local on thistles esp. Cirsium arvense Widespread in damp grassland, and wetlands + ++ Local ? Recently split from next species. Scarcer than next species in my experience. Widespread? The commoner of the split in Berks, frequent in tussocks in winter + ++ ++ ++ 69 Crioceris asparagi (Linnaeus, 1758) Asparagus beetle Lilioceris lilii (Scopoli, 1763) Lily beetle Labidostomis tridentata (Linnaeus, 1758) Clytra quadripunctata (Linnaeus, 1758) Cryptocephalus aureolus Suffrian, 1847 Cryptocephalus biguttatus (Scopoli, 1763) Cryptocephalus bilineatus (Linnaeus, 1767) Cryptocephalus bipunctatus (Linnaeus, 1758) Cryptocephalus coryli (Linnaeus, 1758) Cryptocephalus frontalis Marsham, 1802 Cryptocephalus fulvus Goeze, 1777 Cryptocephalus hypochaeridis (Linnaeus, 1758) Cryptocephalus labiatus (Linnaeus, 1761) Cryptocephalus moraei (Linnaeus, 1758) Cryptocephalus nitidulus Fabricius, 1787 Cryptocephalus parvulus Müller, 1776 + Widespread but very local on asparagus in gardens, neglected allotments etc. ++ + + ?> + + + >? ++ + ++ ++ + + Local, increasing, on various garden lilies (RDB1) ? [On birch] V.local on and near wood ant Formica rufa nests, in which the larvae develop. (Nb) V.local on downland, adults on various yellow composites (RDB2) Rare, on wet heathland on Erica tetralix Bagshott 1 ). Confused with dark C. bipunctatus. (Nb) Rare on downland on kidney vetch Anthyllis vulneraria (Nb) V.local on heathland on birch?, and on the on various trees. (RDB1) Rare, on hazel and birch, restricted to (Na) ? On hawthorn. Local on sheep’s sorrel Rumex acetosella on sandy heathland. Also on Lotus corniculatus, esp. on the chalk V.local on downland, adults on various yellow composites Local on birch and oak. Local on St.John’s Worts Hypericum spp. esp. on sandy and chalky soils. (RDB1) Rare on young birch and hazel on . (Nb) V.local on birch on Lower Greensand and Tertiary heaths and commons. + ++ (Na) Rare on birch. Widespread on birch and Salix. + + (RDB2) [On hazel, birch, aspen etc.] Local in woodlands. 1992, 1995 Rare on bedstraws Galium spp. esp,. on sand and chalky soils + Local on bedstraws Galium spp. + Naturalised and increasing on rosemary Rosmarinus spp. and Lavender Lavandula spp. + + ? [On black horehound Ballota nigra and plantains] Rare on Hypericum esp. H.tetrapterum in damp rides and heaths. Rare On labiates [hemp-nettle in damp areas] (Na) Extinct? Late Neolithic, Runnymede Robinson (1991) [on Tansy] Cryptocephalus primarius Harold, 1872 Cryptocephalus punctiger Paykull, 1799 Cryptocephalus pusillus Fabricius, 1777 Cryptocephalus querceti Suffrian, 1848 Cryptocephalus sexpunctatus (Linnaeus, 1758) Lamprosoma concolor (Sturm, 1807) Timarcha goettingenis (Linnaeus, 1758) Lesser bloody-nosed beetle. Timarcha tenebricosa (Fabricius, 1775) Bloody nosed beetle Chrysolina americana (Linnaeus, ) Rosemary beetle Chrysolina banksi (Fabricius, 1775) Chrysolina brunsvicensis (Gravenhorst, 1807) Chrysolina fastuosa (Scopoli, 1763) Chrysolina graminis (Linnaeus, 1758) Tansy leaf beetle Chrysolina haemoptera (Linnaeus, 1758) Chrysolina hyperici (Forster, 1771) Chrysolina herbacea (Duftschmid, 1825) = menthastri (Suffrian, 1851) Mint leaf beetle Chrysolina oricalcia (Müller, 1776) Chrysolina polita (Linnaeus, 1758) Chrysolina sanguinolenta (Linnaeus, 1758) Chrysolina staphylaea (Linnaeus, 1758) Chrysolina varians (Schaller, 1783) Chrysolina violacea (Müller, 1776) LRom LRom Rom Rom ESax ++ ++ (Nb) ? [On plantains] Purley Local on Hypericum esp. on sandy and chalky soils. Local on Mentha aquatica & hybrids in wetlands, increasingly occurring in gardens on cultivated mints. + ++ + + + ++ (Nb) Widespread on cow parsley Anthriscus Widespread in damp areas on labiates (Na) Rare on common toadflax Linaria vulgaris. Rare? On buttercups? Local on Hypericum esp. on sandy and chalky soils. (Nb) Rare on ground-ivy Glechoma hederacea. Frilford 1987 70 Gastrophysa polygoni (Linnaeus, 1758) Ro/Sax Gastrophysa viridula (Degeer, 1775) Sax Phaedon armoraciae (Linnaeus, 1758) Phaedon cochleariae (Fabricius, 1792) Phaedon tumidulus (Germar, 1824) Hydrothassa glabra (Herbst, 1783) BrAge Hydrothassa marginella (Linnaeus, 1758) ESax Prasocuris junci (Brahm, 1790) Prasocuris phellandrii (Linnaeus, 1758) LRom Plagiodera versicolora (Laicharting, 1781) Cjhrysomela aenea L. Sax Chrysomela populi Linnaeus, 1758 Poplar leaf beetle Chrysomela tremula Fabricius, 1787 Phytodecta decemnotata (Marsham, 1802) Phytodecta olivacea (Forster, 1771) Phytodecta pallida (Linnaeus, 1758) Phytodecta viminalis (Linaneus, 1758) Phyllodecta laticollis Suffrian, 1851 Phratora (=Phyllodecta) vitellinae (Linnaeus, 1758) Phratora (=Phyllodecta) vulgatissima (Linnaeus, 1758) Galerucella calmariensis (Linnaeus, 1767) Galerucella lineola (Fabricius., 1781) Galerucella nymphaeae (Linnaeus, 1758) Galerucella sagittariae (Gyllenhal, 1813) Galerucelal pusilla Galerucella tenella (Linnaeus., 1761) Pyrrhalta viburni (Paykull, 1799) Galeruca tanaceti (Linnaeus, 1758) BrAg e Lochmaea caprea (Linnaeus, 1758) Lochmaea crataegi (Forster, 1771) Lochmaea suturalis (Thomson, 1866) Heather beetle Phyllobrotica quadrimaculata (Linnaeus, 1758) Luperus flavipes (Linnaeus, 1767) Luperus longicornis (Fabricius., 1781) Calomicrus circumfusus (Marsham, 1802) Agelastica alni (Linnaeus, 1758) Alder leaf beetle Sermylassa halensis (Linnaeus, 1767) Phyllotreta punctulata (Marsham, 1802) =aerea Allard, 1859 ++ Local on knotgrass Polygonum in disturbed open areas. Local but increasing on docks Rumex, and Persicaria, esp. along , Local on brooklime Veronica beccabungae in ponds and streams Local on Nasturtium officinale and Rorippa in wetlands including streams Widespread on hogweed Heracleum sphondylium Rare. On Ranunculus repens at Radley Local on buttercups Ranunculus in wetlands and damp grasslands Local on brooklime Veronica beccabungae in ponds and streams Local on wetland Oenanthe and occasionally on other plants. Widespread on narrow leaved Salix esp. near water Burghfield (Robinson, 1992) Local on low growing Populus tremula and Salix repens esp. on damp heaths and rides. + + ++ + ++ ++ + (RDB1) Extinct in UK? Last seen in 1950s (Nb) Local on aspen on heaths and commons, 12 post 1980 records Local, but usually present wherever good stands of broom occur. V.local on hazel, and rowan Local on sallow and other Salix esp. on damp heathy areas. Local on Populus esp. aspen P.tremulae Local on Salix and Populus ++ V.local on Salix and Populus ++ ++ ++ ++ ++ + + Widespread on purple loosestrife Lythrum salicaria at river, stream and canal margins on Local on sallows Salix. Local on waterlillies esp. Nymphaea alba Widespread on various wetland plants Rumex hydrolapathi, Lysimachia vulgaris (‘G.grisescens’ form) Ox district Local on meadowsweet Filipendula ulmaria esp in damp meadows Local on wayfaring tree Viburnum lantana esp. on the chalk, and guelder-rose V.opulus. Local on devil’s-bit cabious in damp grassland Radley ++ ++ ++ Widespread on sallows and other Salix Widespread on hawthorn Locally but can be very abundant on ling Calluna vulgaris. + + ++ V.local on skullcap Scutellaria galericulata in damp grassland and wetlands. (Nb) Local on aspen, birch, [oak] etc. Locally frequent on birch esp. on the heaths and commons (Na) ? [On gorse Ulex, broom Cytisus, and dyer’s greenweed Genista] , (RDBK) Local on bedstraws Galium spp. esp. on downland (Nb) Rare on radish Raphanus, and horse-radish Armoracia rusticana ++ ++ ++ ++ ++ ++ ++ ++ ++ ++ ? ++ ++ 71 Phyllotreta atra (Fabricius, 1775) ++ Frequent on various crucifers + + + + ++ Local on various crucifers (Nb) V.local on various crucifers. Local on cuckooflower Cardamine pratense ? On various crucifers Frequent on crucifers ++ Local on crucifers and weld Reseda lutea + + + ++ Scarce on wild mignonette Reseda luteola on the chalk Local on crucifers Local on Cardamine and Nasturtium Widespread on crucifers + ? Local on crucifers Aphthona atrocaerulea (Stephens, 1831) Aphthona atrovirens Foerster, 1849 Aphthona euphorbiae (Schrank, 1781) Aphthona herbigrada (Curtis, 1837) Aphthona lutescens (Gyllenhal 1813) Aphthona melancholica Weise, 1888 Aphthona nonstriata (Goeze, 1777) Longitarsus aeruginosus (Foudras, 1860) Longitarsus agilis (Rye, 1868) Longitarsus anchusae (Paykull, 1799) Longitarsus atricillus (Linnaeus, 1761) Longitarsus ballotae (Marsham, 1802) Longitarsus brunneus (Duftschmid, 1825) Longitarsus curtus (Allard, 1860) Longitarsus dorsalis (Fabricius, 1781) Longitarsus exoletus (Linnaeus, 1758) Longitarsus ferrugineus (Foudras, 1860) Mint flea beetle Longitarsus flavicornis (Stephens, 1831) Longitarsus fowleri Allen, 1967 Longitarsus ganglbaueri Heikertinger, 1912 Longitarsus gracilis Kutschera, 1864 Longitarsus holsaticus (Linnaeus, 1758) Longitarsus kutscherae (Rye, 1872) + + ++ + On spurges Euphorbia Local on marjoram Origanum vulgare primarily on the Widespread and frequent, wintering on evergreens Local on the with rock-rose. ? [On purple loosestrife] ? On spurges especially wood spurge Euphorbia amygdaloides Widespread on yellow flag Iris pseudacorus (RDB1) ? [on hemp agrimony Eupatorium cannabinum] (Na) ? On figworts Scrophularia sp. (Nb) Rare on viper’s bugloss and hound’s-tongue. Hitchcopse Pit Widespread on legumes esp. Trifolium spp. (Nb) Rare on black horehound Ballota nigra (Nb) ? [On meadow-rue Thalictrum and sea aster Aster tripolium] last recorded in 1940? (Na) Rare on Boraginaceae. (Nb) Local, increasing on ragwort,, also wintering in ground layer in woodlands. Local on viper’s bugloss and hound’s-tongue (RDB1) ? [On water mint Mentha aquatica] Last recorded in 1945? + Frequent on ragworts Senecio spp. (Na) Rare on teasel Dipsacus esp. on (Na) ? [On Ragwort] ? On ragwort ? [On louseworts Pedicularis] ? On plantains. Longitarsus luridus (Scopoli, 1763) Longitarsus lycopi (Foudras, 1860) Longitarsus melanocephalus (Degeer, 1775) ++ + ++ Widespread and frequent on various composites Rare on gipsywort Lycopus europaeus Local on plantains Phyllotreta consobrina (Curtis, 1837) Phyllotreta cruciferae (Goeze, 1777) Phyllotreta diademata Foudras, 1860 Phyllotreta exclamationis (Thunberg, 1784) Phyllotreta nemorum (Linnaeus, 1758) Turnip flea beetle or Black dolphin Phyllotreta nigripes (Fabricius, 1775) Phyllotreta nodicornis (Marsham, 1802) Phyllotreta ochripes (Curtis, 1837) Phyllotreta tetrastigma (Comolli, 1837) Phyllotreta undulata Kutschera, 1860 Phyllotreta vittata (Fabricius, 1801) Phyllotreta vittula (Redtenbacher, 1849) Ro/ Sax Sax Ro/ Sax Ro/ Sax ++ + + + ++ + + ++ + ++ + + 72 Longitarsus membranaceus (Foudras, 1860) Longitarsus nasturtii (Fabricius, 1792) Longitarsus nigrofasciatus (Goeze, 1777) Longitarsus obliteratus (Rosenhauer, 1847) Longitarsus ochroleucus (Marsham, 1802) Longitarsus parvulus (Paykull, 1799) Longitarsus pellucidus (Foudras, 1860) Longitarsus pratensis (Panzer, 1794) Longitarsus reichei (Allard, 1860) Longitarsus rubiginosus (Foudras, 1860) Longitarsus rutilus (Illiger, 1807) Longitarsus succineus (Foudras, 1860) Longitarsus suturalis (Marsham, 1802) Longitarsus suturellus (Duftschmid, 1825) Longitarsus tabidus (Fabricius, 1775) Altica brevicollis Foudras, 1860 Altica carinthiaca Weise, 1898 Altica longicollis (Allard, 1860) =britteni Sharp, 1914 Altica ericeti (Allard, 1859) Altica lythri Aubé, 1843 Altica oleracea (Linnaeus, 1758) Altica palustris Weise, 1888 Altica helianthemi (Allard, 1859) = pusilla Duftschmid, 1825 Hermaeophaga mercurialis (Fabricius, 1792) Batophila aerata (Marsham, 1802) Batophila rubi (Paykull, 1799) Lythraria salicariae (Paykull, 1800) Ochrosis ventralis (Illiger, 1807) Crepidodera ferruginea (Scopoli, 1763) Crepidodera transversa (Marsham, 1802) Derocrepis rufipes (Linnaeus, 1758) Hippuriphila modeeri (Linnaeus, 1761) Chalcoides aurata (Marsham, 1802) Chalcoides aurea (Fourcroy, 1785) Chalcoides fulvicornis (Fabricius, 1792) Chalcoides nitidula (L.innaeus, 1758) Chalcoides plutus (Latreille, 1804) Epitrix atropae Foudras, 1860 Epitrix pubescens (Koch, 1803) Podagrica fuscicornis (L.innaeus, 1767) Podagrica fuscipes (Fabricius, 1775) Mantura matthewsi (Curtis, 1833) Mantura obtusata (Gyllenhal, 1813) ++ + Local on wood sage Teucrium scorodonia (Nb) Rare on comfrey an viper’s bugloss (Na) Rare on figworts and mulleins Local on the chalk on thyme, and marjoram (Nb) ? [On ragwort] (Nb!?) On flax Linum. Once rare now very frequent across the county Local on field bindweed Convolvulus arvensis Widespread on Plantains ? Widespread on bindweed Calystegia, riddling the leaves with small holes (Na) Rare on water figwort Scrophularia auriculata., Widespread on a variety of herbs esp. yarrow and hemp agrimony Eupatorium cannabinum (Nb) ? [on common gromwell Lithospermum officinale] , Frequent on ragworts (Nb) Rare on mulleins esp. great mullein Verbascum thapsus. (Na) ? [On hazel] Recently recognised species in Britain, probably widespread on Lathyris. etc. Windsor Forest area (PMH) Rare, on heather, Erica, and ling Calluna, and on heathland + ++ + + + (Nb) Rare on cross-leaved heath Erica tetralix on wet heathland Frequent on willowherbs and purple loosestrife ? on heathers Calluna, Erica Local on willowherbs and purple loosestrife Local on the on rock-rose and salad burnet +_ + + ++ Widespread and often abundant on dog’s mercury in woods and hedgebanks. (Nb) Rare on Rubus sp. Widespread on Rubus sp. (Nb) Local on yellow loosestrife Lysimachia vulgaris and yellow pimpernel L. nemorum. (RDB3) ? [On bittersweet] Widespread on thistles Widespread on thistles ? On Vetches Vicia Local on horsetails Equisetum esp. in damp grassland Widespread on Salix and Populus Widespread on Salix and Populus Widespread on Salix and Populus (Nb) Rare on aspen on damp heath and commons. Local on Salix and Populus (Nb) V.local on deadly nightshade Atropa bell-donna on disturbed chalky sites on the Widespread on bittersweet. (Nb) Rare on musk mallow Malva moschata. Frilford (JMC) (Na) ? Extinct? [On mallows Malva species] V.local on downland with rock-rose Helianthemum nummularium (Nb) Rare on docks Rumex. + + + + ++ + ++ + ++ ++ + ++ + LRom Rom ++ ++ ++ ++ ++ ++ ++ ++ ++ ++ ++ + + + 73 Mantura rustica (Linnaeus, 1767) Chaetocnema aerosa (Letzner, 1847) Chaetocnema arida Foudras, 1860 Chaetocnema concinna (Marsham, 1802) Chaetocnema confusa (Boheman, 1851) Chaetocnema hortensis (Fourcroy, 1785) Chaetocnema picipes Stephens, 1831 Chaetocnema subcoerulea (Kutschera, 1864) Sphaeroderma rubidum (Graells, 1858) Sphaeroderma testaceum (Fabricius, 1775) Apteropeda globosa (Illiger, 1794) Apteropeda orbiculata (Marsham, 1802) Mniophila muscorum (Koch, 1803) Psylliodes affinis (Paykull, 1799) Potato flea beetle Psylliodes attenuata (Koch, 1803) Hop flea beetle Psylliodes chalcomera (Illiger, 1807) Psylliodes chrysocephala (Linnaeus, 1758) Psylliodes cuprea (Koch, 1803) Psylliodes dulcamarae (Koch, 1803) Psylliodes hyoscyami (Linnaeus, 1758) Henbane flea beetle Psylliodes luteola (Müller, 1776) * Psylliodes napi (Fabricius, 1792) Psylliodes picina (Marsham, 1802) Psylliodes weberi Lohse, 1955 Pilemostoma fastuosa (Schaller, 1783) Cassida flaveola Thunberg, 1794 Cassida hemisphaerica Herbst, 1799 Cassida murraea Linnaeus, 1767 Cassida nebulosa Linnaeus, 1758 Cassida nobilis Linnaeus, 1758 Cassida prasina Illiger, 1798 Cassida rubiginosa Müller, 1776 Cassida vibex Linnaeus, 1767 Cassida viridis Linnaeus, 1758 Green tortoise beetle Cassida vittata de Villers, 1789 Nemonychidae 1 species Cimberis attelaboides (Fabricius, 1787) Ro/Sax > > + ++ + ++ + ++ + ++ + ++ (Nb) Rare on docks Rumex (RDBK) ? Grassy rides? Rare in grasslands. taken by many recorders from 1980s-1996 at least. Widpearead on various Polygonaceae Rare. Grassy rides. Widespread in open grasslands Recently separated spp. (Nb) Local on Juncus in damp grassland >10 post 1980 recs. Widespread on thistles Widespread on thistles (Nb) ? [On Rubus?] Widespread in woods and scrubby places (Nb) ? [In moss in woods, possibly on foxglove??] Widespread on bittersweet (RDB1) ? [On hop Humulus lupulus ] SUB + ++ + ++ (Nb) Local on thistles Carduus and possibly Cirsium sp. Widespread on various crucifers Local on various crucifers Widespread on bittersweet Solanum dulcamarae (RDB1) Extinct? Associated with henbane Hyoscyamus niger. , ++ + + + + ++ + + + ++ ++ ++ (RDBK) Recent arrival (On cereal crops, also on trees in autumn) Also comes to light. Local on various crucifers ? On purple loosestrife. ? On Rorippa (Na) Rare on ploughman’s spikenard Inula conyza on downland V.local, in open often damp places. Host plant(s) unknown? (Na) Rare on campion Silene. ? [On common fleabane Pulicaria dysenterica] (RDBIn) ? [On Chenopodiaceae] (Nb) Rare on spurreys on disturbed sandy soils (Nb) Local on yarrow in open grassland including road verges, and ruderal sites. Widespread on thistles and knapweeds Widespread on thistles and knapweeds Local on water mint Mentha aquatica + Rare on spurreys VCH 1907 2005 + Notes Local on pine Pinus spp., often on the flowers 74 Anthribidae (Orthocerous Weevils) 8 species SUB Platyrhinus resinosus (Scopoli, 1763) Dissoleucus (=Tropideres) niveirostris (Fabricius, 1798) Platystomos albinus (Linnaeus, 1758) Bird-dropping weevil Anthribus fasciatus Forster, 1771 Anthribus nebulosus Forster, 1771 Araecerus fasciculatus (Degeer, 1775) Choragus sheppardi Kirby, 1819 Bruchela rufipes (Olivier, 1790) Rhynchitidae (Orthocerous Weevils) 16 species Involvulus (=Rhynchites) caeruleus (Degeer, 1775) Twig-cutting weevil Involvulus (=Rhynchites) cupreus (Linnaeus, 1758) Lasiorhynchites (=Rhynchites) cavifrons Gyllenhal, 1833 Lasiorhynchites (=Rhynchites) olivaceus Gyllenhal, 1833 Neocoenorrhinus (=Rhynchites) aeneovirens (Marsham, 1802) Neocoenorrhinus (=Rhynchites) aequatus (Linnaeus., 1767) Neocoenorrhinus (=Rhynchites) germanicus Herbst, 1797 Neocoenorrhinus (=Rhynchites) interpunctatus Stephens, 1831 Neocoenorrhinus (=Rhynchites) longiceps Thomson, 1888 Neocoenorrhinus (=Rhynchites) pauxillus Germar, 1839 Temnocerus (=Rhynchites) nanus (Paykull, 1792) Temnocerus (=Rhynchites) tomentosus Gyllenhal, 1839 Byctiscus betulae (Linaneus, 1758) Byctiscus populi (Linnaeus, 1758) ??? Deporaus betulae (Linnaeus, 1758) Deporaus mannerheimi (Hummel, 1823) VCH 1907 2005 + (Nb) Rare in Daldinia concentrica on ash, (RDB2) Rare on hardwood branches in woods and old hedgerows. + (Nb) Local on hardwoods, alder, beech etc. seems to have increased in abundance in past few years,!? + + (Na) V.local on trees esp. oak in open parkland, preying on bark coccids. (Nb) Local on trees including pines, preying on Coccinae. 5 post 1980 sites Casual introduction, Synanthropic. (Na) Rare on old dead ivy Helix hederae. Rare, increasing. On wild mignonette Reseda lutea. Unhill Downs 1997 (PJH) + ++ SUB Notes VCH 1907 2005 ++ Notes V.local on rosaceuos trees esp. Cratageus, Prunus and Malus. + (Nb) V.local on rowan Sorbus aucuparia and whitebeam S.aria. + (Nb) V.local on Quercus in one-year old twigs. ++ (Na) V.local on Quercus in one-year old twigs. + Local on oak ++ Frequent on hawthorn, also on other Rosaceae Malus, Prunus. + Local, on wide range of shrubby, and herbaceus Rosaceae. + (Nb) Rare on oak. + (Nb) V.local on birch, Salix and Pyrus. + Rare. [On wide range of shrubby and herbaceus Rosaceae, esp. Cratageus, Prunus] + + Local on birch, also Salix and Alnus (Nb) Local on Salix and Populus esp. P. tremulae + (Nb) Rare, declining? On birch and hazel. (RDB3) Rare on Populus esp. aspen P.tremulae. Frequent on Betula less common on Alnus, Corylus, etc.. Rare on Betula in wetter sites than betulae. ++ + 75 Attelabidae (Leaf Roller weevils) 2 species SUB Attelabus nitens (Scopoli, 1763) Red oak-roller Apoderus coryli (Linnaeus, 1758) Apionidae (= Brentidae)( Seed Weevils) 74 species Apion cruentatum Walton, 1844 Apion frumentarium (Linnaeus, 1758) Apion haematodes (Linnaeus, 1758) Apion rubens Stephens, 1839 Apion rubiginosum Grill, 1893 Aizobius sedi Germar, 1818 Perapion affine Kirby, 1808 Perapion curtirostre Germar, 1817 Perapion hydrolapathi (Marsham, 1802) Perapion lemoroi Brisout, 1880 Perapion marchicum Herbst, 1797 Perapion violaceum Kirby, 1808 Aspidapion aeneum (Fabricius, 1775) Aspidapion radiolus (Marsham, 1802) Ceratapion carduorum Kirby, W., 1808 Ceratapion onopordi Kirby, 1808 Ceratapion gibbirostre Gyllenhal Acentrotypus brunnipes Boheman 1905. SUB Notes VCH 19072005 Notes + ++ ++ + ++ + ++ ++ ++ ++ ++ Ro/S ax Rom ++ + ++ + + Diplapion confluens Kirby, 1808 Diplapion stolidum Germar, 1817 Omphalapion hookerorum Kirby, 1808 Exapion difficile Herbst, 1797 Exapion fuscirostre (Fabricius, 1775) Exapion genistae Kirby, 1811 Exapion ulicis (Forster, 1771) Kalcapion pallipes Kirby, 1808 Kalcapion semivittatum Gyllenhal, 1833 Melanapion minimum Herbst, 1797 Squamapion atomarium Kirby, 1808 Squamapion cineraceum Wencker, 1864 Squamapion flavimanum Gyllenhal, 1833 Squamapion vicinum Kirby, 1808 Taeniapion urticarium (Herbst, 1784) VCH 1907 2005 + + ++ ++ ++ ++ + + ++ + Ro/ + V.local on oak foliage especially on young ‘scrub’ oaks. Local on hazel V.local on Rumex acetosa esp. in unimproved grasslands. Local on broad leaved docks Rumex in grassland on verges etc. Local on Rumex acetosella on disturbed sandy soils Local on Rumex acetosella on disturbed sandy soils (RDB3) Rare on Rumex acetosella on disturbed sandy soils. Frilford (JMC det JSD) (Nb) ? [On stonecrops] (Na) Rare on sorrel Rumex acetosa. Frequent on various docks and sorrels Local on Rumex incl. R. acetosa Rare on Polygonum aviculare Local on Rumex acetosella on disturbed sandy soils Local on Rumex incl. R. acetosa Locally abundant on mallows Malva spp. Locally abundant on mallows Malva spp. ?Rare? on thistles Local on thistles, knapweeds, and burdocks ?Rare on thistles RDB1 Extinct formerly on cudweeds Tubney 1905 (JJW) ? [On mayweeds and chamomile] (Nb) V.local on ox-eye daisy. Herons Nest 2003 (JSD) Local on mayweeds Tripleurospermum spp. (Na) ? On dyer’s greenweed Genista tinctoria. Local on broom Cytisus (Na) ? [On petty whin Genista anglica] Widespread and frequent on gorse Ulex Local on dog’s mercury Mercurialis perennis, in woodlands (Na) Local on annual mercury Mercurialis annua esp. on derelict /abandoned sites in London area. (RDB3) ? [On Sallows in sawfly galls made by Pontania spp.] V.local on thyme Thymus, mostly on the chalk. (Na) Rare on self-heal Prunella vulgaris. Kirtons farm 2003 (JSD) (Na) V.local on on marjoram. (Nb) ? [On mints Mentha spp.] Rare on nettles Urtica spp. 76 Malvapion malvae (Fabricius., 1775) Pseudapion rufirostre (Fabricius., 1775) Sax Ro m Ro m ++ Locally abundant on mallows Malva spp. ++ Locally abundant on mallows Malva spp. Cynapion afer Gyllenhal, 1833 Cynapion gyllenhali Kirby, 1808 Cynapion spencii Kirby, 1808 Eutrichapion ervi Kirby, 1808 Eutrichapion scutellare Kirby, 1811 Eutrichapion viciae (Paykull, 1800) Eutrichapion vorax Herbst, 1797 Eutrichapion punctigerum (Paykull, 1792) Hemitrichapion reflexum Gyllenhal, 1833 Hemitrichapion waltoni Stephens, 1839 Holotrichapion aethiops Herbst, 1797 Holotrichapion ononis Kirby, 1808 Holotrichapion pisi (Fabricius, 1801) Oxystoma cerdo Gerstaecker, 1854 Oxystoma craccae (Linnaeus, 1767) Oxystoma pomonae (Fabricius, 1798) ? [On Lathyrus] , (Nb) Local on vetches Vicia esp. V.cracca. ++ + ++ ++ ++ Rom + Rom Ro/S ax + ++ + ++ + ++ + Oxystoma subulatum Kirby, 1808 Pirapion immune Kirby, 1808 Catapion curtisi Stephens, 1831 Catapion pubescens Kirby, 1811 Catapion seniculus Kirby, 1808 Ischnopterapion loti Kirby, 1808 Ischnopterapion modestum Germar, 1817 Ischnopterapion virens Herbst, 1797 Protopirapion atratulum (Marsham, 1802) Stenopterapion meliloti Kirby, 1808 Stenopterapion tenue Kirby, 1808 Synapion ebeninum Kirby, 1808 Trichapion simile Kirby, 1811 ++ + Protapion apricans Herbst, 1797 Protapion assimile Kirby, 1808 Protapion dichroum Bedel, 1886 Protapion difforme Ahrens, 1817 Protapion dissimile Germar, 1817 Protapion filirostre Kirby, 1808 Protapion laevicolle Kirby, 1811 Protapion nigritarse Kirby, 1808 ++ + ++ + + ++ + ++ ++ + + + ++ + + + ? [Damp places on Vicia] , Frequent on various vetches V.local on heathlands on gorse, esp. Ulex minor, and possibly U.gallii. Frequent on various vetches Local on vetches V.local on various vetches (Na) Rare? On Onobrychis vicifolia on calcareous grassland. Rare on horseshoe vetch Hippocrepis comosa on the chalk. ?On Vicia . V.local on restharrows Ononis repens and O. spinosa, esp. on the chalk. Widespread on medicks Medicago (Nb) Local on vetches esp. V.cracca Local on various vetches Vicia Local on various vetches Vicia and Lathyrus Local but widespread on meadow vetchling Lathyrus pratensis Local on broom Cytisus esp. on sandy soils. (Na) ? [On Trifolium] (Nb) ? [On Trifolium especially small yellow flowered species] Rare on Medicago [and Trifolium hybridium ] Widespread, on Lotus corniculatus Local on large bird’s-foot trefoil Lotus uliginosus in damp meadows and marshlands. Local on a wide variety of clovers Trifolium in open grassy places. Local on broom Cytisus esp. on sandy soils. Local on Melilotus. Local on medicks Medicago esp. lupulina Rare, on large bird’s-foot trefoil Lotus uliginosus. Local but common on the heaths on Betula. Frequent on red clover Local on various Trifolium spp. Widespread and often abundant on clovers esp. Trifolium repens (Nb) ? Usually found on Polygonum spp. (Nb) V.local on hare’s-foot clover T.arvense on sandy soils. Frilford etc (JSD) (Nb) ? [On black medick Medicago lupulina] (Na) ? [On subterranean clover T.subterraneum] Frequent on various clovers 77 Protapion schoenherri Boheman, 1839 Protapion trifolii (Linnaeus, 1768) Protapion varipes Germar, 1817 Nanophyidae (formerly part of Apionidae) 2 species ++ ++ SUB Nanophyes marmoratus (Goeze, 1777) Dieckmanniellus (=Nanophyes) gracilis (Redtenbacher, 1849) Dryophthoridae 2 species SUB Dryophthorus corticalis (Paykull, 1792). Sitophilus granarius (L.innaeus, 1758) Grain weevil Sitophilus oryzae (Linnaeus, 1763) Small grain weevil Erirhinidae 10 species Grypus equiseti (Fabricius., 1775) Notaris acridulus (L.innaeus, 1758) BrA ge Notaris scirpi (Fabricius, 1792) Tournotaris (=Notaris) bimaculatus (Linnaeus, 1787) Thryogenes festucae (Herbst, 1795) Thryogenes nereis (Paykull, 1800) Thryogenes scirrhosus (Gyllenhal, 1836) Stenopelmus rufinasus Gyllenhal, 1836 Raymondionymidae 1 species SUB Ferreria (=Raymondionymus) marqueti (Aubé, 1863) Curculionidae (True weevils) 308 species + 3 sub-fossil VCH 1907 2005 ++ ++ Notes VCH 1907 2005 ++ ++ Notes ++ SUB SUB (Na) ? [On hares-foot clover] Frilford JJW Frequent on red and white clover (Nb) ? [On clovers] VCH 1907 2005 ++ ++ Widespread on purple loosestrife, in damp grassland and pond and river margins. (Na) V.local on water Purslane Lythrum portula . Snelsmore Common 2003 (JSD) (RDB ) Windsor Forest (PMH) Syananthropic in grain, occasionally found outdoors. Hutchins Copse 1998 (JSD) Syananthropic in grain occasionally found outdoors Hutchins Copse 1998 (JSD) Notes (Nb) V.local on horsetail Equisetum (arvense and palustre) damp places. Sheffield Bottom 2003 (JSD) Local in wetlands on grasses esp. Glyceria maxima ++ Local on large pond-sedge Carex acutiformis and other wetland plants including Typha (Nb) [On large wetland grasses and sedges] + + + + Local in wetlands on sedges? Local on spike-rush Eleocharis palustris (Nb) Rare on bur-reeds Sparganium spp. Very local on water fern Azolla, but usually present on good colonies of the host plant. VCH 1907 2005 + Notes 1902 1951 1950 2004 Notes Subterranean 78 Archarius (=Curculio) pyrrhoceras Marsham, 1802 Archarius (=Curculio) salicivorus Paykull, 1792 Archarius (=Curculio) villosus Fabricius, 1781 Curculio betulae (Stephens, 1831) Curculio glandium Marsham, 1802 Curculio nucum Linnaeus, 1758 Curculio rubidus (Gyllenhal, 1836) Curculio venosus (Gravenhorst, 1807) Acalyptus carpini (Fabricius, 1792) Anoplus plantaris (Naezen, 1794) Anoplus roboris Suffrian, 1840 Anthonomus bituberculatus Thomson, 1868 Anthonomus brunnipennis (Curtis, 1840) Anthonomus chevrolati Desbrochers, 1868 Anthonomus humeralis (Panzer, 1795) Anthonomus pedicularius (Linnaeus, 1758) Anthonomus piri Kollar, 1837 Anthonomus pomorum (Linnaeus, 1758) Apple blossom weevil Anthonomus rubi (Herbst, 1795) Anthonomus ulmi (Degeer, 1775) Cionus alauda (Herbst, 1784) Cionus hortulanus (Fourcroy, 1785) Cionus nigritarsis Reitter, 1904 Cionus scrophulariae (Linnaeus, 1758) Cionus tuberculosus (Scopoli, 1763) Cleopus pulchellus (Herbst, 1795) Ellescus bipunctatus (Linnaeus, 1758) Dorytomus dejeani Faust, 1882 Dorytomus hirtipennis Bedel, 1884 Dorytomus ictor (Herbst, 1795) Dorytomus longimanus (Forster, 1771) Dorytomus melanophthalmus (Payk, 1792) Dorytomus rufatus (Bedel, 1888) Dorytomus taeniatus (Fabricius, 1781) Dorytomus tortrix (Linnaeus, 1761) Dorytomus tremulae (Fabricius, 1787) Cleopomiarus (=Miarus) graminis (Gyllenhal, 1813) Cleopomiarus (=Miarus) plantarum (Germar, 1824) Gymnetron beccabungae (L., 1761) Gymnetron melanarium (Germar, 1821) Gymnetron rostellum (Herbst, 1795) ++ Widespread on oak, developing in currant galls made by the wasp Neuroterus quercusbaccarum (Hym., Cynipidae). Widespread on Salix, developing in galls made by the Pontania saw-flies. (Nb) Rare on oak, in galls made by Biorhiza pallida (Hym., Cynipidae) (Nb) Rare on birch. Windsor 1991 Widespread on oak Local on hazel. Declining? Possibly due to paucity of nuts, due to competition with grey squirrel ?? (Nb) Rare on birch. Local on oak (Nb) Rare on Salix in wetlands. Widespread on birch (Nb) Local on alder. Rare on hawthorn. , Rare on tormentil Potentilla erecta. (RDBK) ? [On hawthorn and possibly Sorbus] (RDBK) Rare on crab apple Malus. Widespread on hawthorn (RDB3) ? [On fruit trees esp. Apple] Local on Apple Malus sp. ++ + ++ ++ * ++ ++ ++ + + * + ++ + ++ ++ + + + Widespread on Rosaceae (Nb) Rare on elm Widespread on figworts Scrophularia nodosa and S.auriculata. Also on Mulleins including V.blattaria Widespread on mulleins and figworts [occasionally on Buddleja] (Na) Rare on dark mullein Verbascum nigrum on the chalk. Local on figworts, including introduced garden spp. and occasionally on Buddleja. Local on water figwort S.auriculata [and occasionally on S.nodosa] Local on figworts and mulleins. Abundant on moth mullein Verbascum blattaria (Nb) ? On willows in damp areas esp. heaths. Local on aspen. (Na) [On willows in wet woods] (Nb) Rare on poplars esp. hybrid black poplars. Local on poplars Populus. Rare? on Salix. Local on Salix spp. Widespread on Salix Rare on aspen (Nb) Rare on aspen esp. large trees. Also on White Poplar P.alba. (Nb) Rare on Campanula on the ++ ++ ++ + ++ ++ + ++ ++ + + + + ++ * (RDBK) [On Campanula] + + (Na) ? Burghfield (Robinson 1992 ) (Nb) Rare on germander speedwell Veronica chamaedrys. (Na) ? [On a variety of plants including Matricaria, Filago and possibly Veronica chaemaedrys] Sax 79 ++ (Nb) Rare on brooklime Veronica beccabungae. (Nb) Local on water speedwells [and hybrid]. (Nb) Rare on plantains. Rare on plantains. Widespread on plantains Plantago spp. ++ Local on plantains in open dry areas. Miarus campanulae (Linnaeus, 1767) + Rare breeds on Campanula esp. C. glomerata] on the , [harebell C. rotundifolia ], Adults also at yellow composites. Rhinusa (=Gymnetron) antirrhini (Paykull, 1800) Rhinusa (=Gymnetron) collina (Gyllenhal, 1813) Rhinusa (=Gymnetron) linariae (Panzer, 1795) Isochnus (=Rhynchaenus) populicola (Silfverberg, 1977) * Orchestes (=Rhynchaenus) alni (Linnaeus, 1758) Orchestes (=Rhynchaenus) fagi (Linnaeus, 1758) Orchestes (=Rhynchaenus) iota (Fabricius,, 1787) Orchestes (=Rhynchaenus) pilosus (Fabricius, 1781) Orchestes (=Rhynchaenus) quercus (Linnaeus, 1758) Orchestes (=Rhynchaenus) rusci (Herbst, 1795) Orchestes (=Rhynchaenus) signifer (Creutzer, 1799) =avellanae (Donovan, 1797) Pseudorchestes (=Rhynchaenus) pratensis (Germar, 1821) Ramphus oxyacanthae (Marsham, 1802) Ramphus pulicarius (Herbst, 1795) Tachyerges (=Rhynchaenus) pseudostigma Tempere, 1982 Tachyerges (=Rhynchaenus) salicis (Linnaeus, 1758) Tachyerges (=Rhynchaenus) stigma (Germar, 1821) Smicronyx jungermanniae (Reich, 1797) Smicronyx reichi (Gyllenhal, 1836) Orthochaetes insignis (Aubé, 1863) Orthochaetes setiger (Beck, 1817) ++ * + Local on common toadflax Linaria vulgaris (Na) rare on common toadflax Linaria vulgaris (Na) Rare on common toadflax Linaria vulgaris Rare? On crack willow Salix fragilis. ++ ++ + ++ Local on elm Ulmus Widespread and frequent on beech, wintering in holes and crevices in trees. (Nb) Rare, but locally abundant on bog myrtle Myrica gale on the Tertiary heaths. Widespread on oak ++ Widespread on oak ++ + Widespread on birch Local on hazel and oak + (Nb) Rare on common knapweed Centaurea nigra. ++ ++ + Widespread on hawthorn and other Rosaceae Local on Salix (RDBK) Rare leaf-miner on Salix. + Local on Salix + Local on willows and alder + + (Nb) V.local on dodder Cuscuta on heaths on Calluna, and on and richer grasslands on Lotus etc. (RDB3) Rare on centaury Centarium erythraea esp.on chalk grassland. (Nb) Rare in litter/moss on open grassland esp. downland (Nb) Local in litter/moss on open grassland esp. downland, [but also in woodlands with bluebells] Gymnetron veronicae (Germar, 1821) Gymnetron villosulum Gyllenhal, 1838 Mecinus circulatus (Marsham, 1802) Mecinus (=Gymnetron) labile (Herbst, 1795) Mecinus (= Gymnetron) pascuorum (Gyllenhal, 1813) Mecinus pyraster (Herbst, 1795) Pseudostyphlus pillumus (Gyllenhal, 1836) Sibinia potentillae Germar, 1824 Sibinia primitus (Herbst, 1795) Tychius junceus (Reich, 1797) Tychius lineatulus Stephens, 1831 LRom Rom Rom Ro/Sa x LRo m ++ + ++ + ++ ++ * (Na) On scented mayweed Matricaria recutita] (Nb) Rare on spurreys. (Nb) Local on spurreys etc. esp. on disturbed sandy grounds. Snelsmore Common, Herons Nest 2003 (JSD) Local on various small legumes (Na) Rare on zig-zag clover Trifolium medium. 80 Tychius meliloti Stephens, 1831 Tychius parallelus (Panzer, 1794) Tychius picirostris (Fabricius, 1787) Tychius polylineatus (Germar, 1824) Tychius pusillus Germar, 1842 Tychius schneideri (Herbst, 1795) Tychius squamulatus (Gyllenhal, 1836) Tychius stephensi Gyllenhal, 1836 Bagous binodulus (Herbst, 1795) Bagous limosus (Gyllenhal, 1827) Bagous longitarsis Thomson, 1868 Bagous glabrirostris (Herbst, 1795) Bagous lutulentus (Gyllenhal, 1813) Bagous tempestivus (Herbst, 1795) Bagous (=Hydronomus) alismatis (Marsham, 1802) Baris lepidii Germar, 1824 Baris picicornis (Marsham, 1802) Limnobaris dolobrosa (Goeze, 1777) Limnobaris pilistriata (Stephens, 1831) Limnobaris t-album (Linnaeus., 1758) Amalorrhynchus melanarius (Stephens, 1831) Amalus scortillum (Herbst, 1795) Calosirus terminatus (Herbst, 1795) Ceutorhynchus alliariae Brisout, 1860 Ceutorhynchus assimilis (Paykull, 1792) Cabbage seed weevil Ceutorhynchus atomus Boheman, 1845 Ceutorhynchus chalybaeus Germar, 1824 Ceutorhynchus cochleariae (Gyllenhal, 1813) Ceutorhynchus constrictus (Marsham, 1802) Ceutorhynchus contractus (Marsham, 1802) Ceutorhynchus erysimi (Fabricius, 1787) Ceutorhynchus pallidactylus (Marsham, 1802) Cabbage stem weevil Ceutorhynchus pectoralis Weise, 1895 Ceutorhynchus pervicax Weise, 1883 Ceutorhynchus picitarsis Gyllenhal, 1837 Ceutorhynchus pleurostigma (Marsham,1802) Cabbage gall weevil Ceutorhynchus pyrrhorhynchus (Marsham, 1802) Ceutorhynchus rapae Gyllenhal, 1837 Ceutorhynchus resedae (Marsham, 1802) Ceutorhynchus sulcicollis (Paykull, 1800 ++ * ++ * ++ + + + + > * + + + ++ ++ ++ + ++ LBr A Ro m * Local on melilots (Na) ? [On Broom] , Widespread and frequent on clovers (RDBK) Rare [on downland on red clover] (Nb) Local on disturbed groun on small Trifolium sp. Herons Nest 2003 (JSD) (Nb) Rare on downland on Kidney Vetch (Nb) Rare on the on bird’s-foot trefoil, and possibly Kidney Vetch. Rare on Trifoliums. Extinct in UK? [On water soldier Stratiotes aloides] (Nb) Rare on pondweed Potamogeton natans. (RDB1) ? [On Myriophyllum] (Nb) [On hornworts Ceratophyllum sp.] (Nb) Rare on water horsetail Equisetum fluviatile from which it can occasionally be swept. (Nb) Rare on creeping buttercup Ranunculus repens. (Nb) Rare on water Plantain Alisma plantago-aquatica (Na) Rare on various crucifers in wetland areas esp. riverbanks as at (Nb) Rare on wild mignonette Reseda lutea on chalky soils Extinct in UK. Local on sedges Carex in wetlands Local on sedges Carex in wetlands Local on water-cress Nasturtium spp. esp. on plants growing over flowing water Local on knotgrass P.aviculare in open disturbed areas including old industrial sites. Heron’s Nest 2003 (JSD) (Nb) Rare on wild carrot Daucus carota. Heron’s Nest 2003 (JSD) Local on garlic mustard Alliaria petiolata on hedgebanks, verges etc Widespread on various crucifers esp. Brassica, Sisymbrium etc. (Na) Rare on thale cress Arabidopsis thaliana on disturbed sandy soils. Tubney (JEC) + + + + ++ ? On hedge mustard Sisymbrium officinale. ? On Cardamine sp. (Nb) Rare on garlic mustard Alliaria petiolata ? On hedge mustard Sisymbrium officinale Widespread on crucifers ++ Widespread on crucifers > ? [On crucifers] (Na) Rare on cuckooflower Cardamine pratense. ? On hedge mustard Sisymbrium officinale ? [On brassicas] > ++ + + Local on hedge mustard Sisymbrium officinale (Nb) Rare on various crucifers. (Nb) ? [On weld Reseda luteola] Rare? On various crucifers 81 Ceutorhynchus thomsoni Kolbe, 1900 Ceutorhynchus turbatus Schultze, 1903 Ceutorhynchus typhae (Herbst, 1795) = floralis (Paykull, 1792) Coeliodes rana (Fabricius, 1787) =dryados (Gmelin, 1790) Coeliodes transversealbofasciatus (Goeze, 1777) = erythroleucos (Gmelin, 1790) Coeliodes ruber (Marsham, 1802) Coeliodinus (=Coeliodes) rubicundus (Herbst, 1795) Datonychus melanostictus (Marsham, 1802) Datonychus urticae Boheman, 1845 Drupenatus nasturtii (Germar, 1824) Glocianus distinctus (Brisout, 1870) Glocianus moelleri Thomson, 1868 Glocianus pilosellus Gyllenhal, 1837 Glocianus punctiger Sahlberg, 1835 Hadroplontus litura (Fabricius, 1775) Hadroplontus trimaculatus (Fabricius, 1775) Micrelus ericae (Gyllenhal, 1813) Microplontus campestris Gyllenhal, 1837 Microplontus rugulosus (Herbst, 1795) Microplontus triangulum Boheman, 1845 Mogulones asperifoliarum (Gyllenhal, 1813 Mogulones euphorbiae Brisout, 1866 Mogulones geographicus (Goeze, 1777) Nedyus quadrimaculatus (Linnaeus, 1758) Parethelcus pollinarius (Forster, 1771) Trichosirocalus rufulus (Dufour, 1851) Trichosirocalus troglodytes (Fabricius, 1787) Zacladus exiguus (Olivier, 1807) (RDBK) ? [On garlic mustard Alliaria petiolata] Rare on hoary cress Cardaria draba Local on crucifers. Until recently known in Britain as C.floralis ++ Local on oak + (Nb) Rare on oak + + Local on oak (Nb) Rare on birch ++ + + ++ + ++ Local on gypsywort Lycopus europaeus, and water mint Mentha aquatica and hybrids near water (RDB3) [On woundwort Stachys] , (Nb) Local on water-cress Nasturtium spp. esp. on plants growing over flowing water Local on small yellow composites in open grassland (RDBK) Rare. (RDB2) Extinct? [Open sandy grassland] Hitchcopse Pit (DC) (Nb) Rare on dandelions Taraxacum spp. Local on thistles Cirsium spp. (Nb) Extinct? [On musk thistle Carduus natans] , , + LRo m Ro m Poophagus sisymbrii (Fabricius, 1777) Sirocalodes depressicollis (Gyllenhal, 1813 Sirocalodes mixtus (Mulsant & Rey, 1858) Stenocarus umbrinus (Gyllenhal, 1837) Tapeinotus sellatus (Fabricius, 1794) Thamiocolus viduatus (Gyllenhal, 1813) Trichosirocalus barnevillei (Brisout, 1866) Trichosirocalus horridus (Panzer, 1801) + + ++ ++ + + + + * + ++ ++ Widespread on heathers esp. Calluna but also on Erica. spp. (Nb) Rare on ox-eye daisy Leucanthemum vulgare Rare on chamomile. (Nb) Rare on yarrow Achillea. Local on Boraginaceae esp. Cynoglossum officinale, and Echium vulgare (Na) Rare, polyphagous? especially on Veronica? on downland. Tubney 1918 (Nb) Rare on viper’s bugloss on the . Frilford, Tubney (JMC /JSD) Widespread and frequent wherever nettles Urtica spp. grow. Common on nettles Urtica ++ ++ + + ++ ++ ++ Local on water-cress Nasturtium spp. and yellocress Rorippa spp. esp. on plants growing over flowing water Local on fumitorys Fumaria sp. in open grassland, and ruderal sites. Herons Nest 2003 (JSD) (Nb) V.local on climbing corydalis Corydalis clavicula on wooded former heathland and commons. (Nb) Rare on poppies Papaver spp. Frilford (JMC) (Na) Local on yellow loosestrife. Virginia Water 1998 (JSD) (Nb) Extinct ? [On marsh woundwort Stachys palustris] (Nb) Local on yarrow Achillea millefolium in open grassy places including road verges. Herons Nest 2003 (JSD) (Na) Rare on spear thistle Cirsium vulgare and musk thistle Carduus nutans. Moulsford Downs (SU5682) 1996 (PJH) (Na) Rare on plantains Plantago spp. on disturbed sandy and chalky sites. Herons Nest 2003 (JSD) Widespread in disturbed open habitats on plantains Plantago + (Nb) Rare on small flowered Geranium spp. ++ Ro m BrA ge Ro m ++ 82 Zacladus geranii (Paykull, 1800) ++ ? [On Geraniums] Heron’s Nest 2003 (JSD) Eubrychius velutus (Beck, 1817) Neophytobius quadrinodosus (Gyllenhal, 1813) Pelenomus canaliculatus Fahraeus, 1843 Pelenomus comari (Herbst, 1795) Pelenomus olssoni Israelson, 1972 Pelenomus quadricorniger Colonelli, 1986 Pelenomus quadrituberculatus (Fabricius, 1787) Pelenomus waltoni Boheman, 1843 Phytobius leucogaster (Marsham, 1802) Rhinoncus albicinctus Gyllenhal, 1837 * Rhinoncus bruchoides (Herbst, 1784) * + + + * + + + ++ ++ (Nb) Rare, aquatic, submerged on pond-weeds. (Na) Rare in grassland [on Polygonum?] (Nb) Rare, aquatic on Myriophyllum in ponds. (Na) Rare on marsh cinquefoil Potentila palustris. [ also on Purple loosestrife L. salicaria] (RDB3) Rare on water purslane Lythrum portula in ponds, rut-pools esp on sandy soils (Na) [Wetlands on pale persicaria Polygonum lapathifolum] Rare? On Polygonums [in dry grassland] (Nb) Local on water pepper Polygonum hydropiper esp. along damp woodland rides beside wheel ruts. (Nb) Rare, aquatic on Myriophyllum in ponds. (RDB1) Rare on amphibious bistort P.amphibium esp. on floating leaves. Virginia Water 2003 (JSD) Local on redshank Polygonum persicaria in rough grassland esp. near rivers. Ro m Rhinoncus castor (Fabricius, 1792) Rhinoncus inconspectus (Herbst, 1795) Rhinoncus pericarpius (Linnaeus, 1758) Rhinoncus perpendicularis (Reich, 1797) Rutidosoma globulus (Herbst, 1795) Cossonus parallelepipedus (Herbst, 1795) Stereocorynes truncorum (Germar, 1824) Euophryum confine (Broun, 1881) Euophryum rufum (Broun, 1880) Pentarthrum huttoni Wollaston, 1854 Rhyncolus ater (Linnaeus, 1758) Phloeophagus lignarius (Marsham, 1802) Cryptorhynchus lapathi (Linnaeus, 1758) Acalles misellus Boheman, 1844 Acalles ptinoides (Marsham, 1802) Acalles roboris Curtis, 1835 Gronops lunatus (Fabricius, 1775) Graptus (=Alophus) triguttatus (Fabricius, 1775) Strophosoma capitatum Degeer, 1775 Neliocarus (=Strophosoma) faber (Herbst, 1784) Neliocarus (=Strophosoma) nebulosum (Stephens, 1831) Neliocarus (=Strophosoma) sus (Stephens, 1831) Strophosoma melanogramma (Forster, 1771) Attactagenius (=Cneorhinus) plumbeus (Marsham, 1802) Philopedon plagiatus (Schaller, 1783) Old lady weevil Barynotus moerens (Fabricius, 1792) ++ Local on sheep’s sorrel Rumex acetosella Local on amphibious bistort Polygonum amphibium Local on Rumex. Spp. Local on Polygonums including amphibious bistort P.amphibium (Na) [On aspen] (Nb) Rare, saproxylic, developing in heartwood of hardwoods. (Na) [Dead wood in ancient woodland] Naturalised. Widespread and frequent. Immigrant from New Zealand, found in rotting wood and building timber. Synanthropic. Rare on rotting wood, esp. structural timber. Much declined possibly at expense of above spp. Extinct? [In dead wood esp. pine] Local in dead wood esp. oak and ash, also Sycamore. (Nb) [On willows] Widespread on dead wood esp. small branches and twigs in woods and hedgerows. ++ ++ + + (Nb) V.local in woodland and on heaths in litter, rotting wood and ant nests. Rare in woods in oak leaf litter and in dead wood. (Nb) Rare on spurreys Spergularia. on diturbed sandy soils. Frilford etc (JMC) Local in grassland with Plantains. + ++ ++ Local on a variety oftrees and bushes esp. in heathy habitats (Nb) Local on disturbed sandy ground on plantains Local on a wide variety of trees and herbs including heathers. Frequent on wooded heaths. ++ ++ + Local on heather and esp. ling. Frequent on a wide variety of trees and herbs. Very abundant on heathsand commons (Nb) Rare on open often sandy ground + Rare on disturbed open sandy ground + Local in woods esp with dog’s mercury Mercurialis perennis esp. ++ ++ ++ ++ * + ++ + ++ Ro m Ro/ Sax Sax 83 Barynotus obscurus (Fabricius, 1775) + ? Open grassy places Radley Bronze age + + + + + ++ Recent Introducuction. Recent Introducuction. Local, polyphagous on trees and on the ground esp. on downland. Recent import, spreading? On lilac and privet hedges. (Nb) ? [Sandy area and heaths] V.local in disturbed open areas on sand and chalk Local on disturbed sandy ground including gardens + + (Nb) Local often in parks and gardens, ( (Nb) Rare disturbed sandy or chalky ground. Hatford 2005 (JMC det JSD) ++ ? [Coastal] Frequent. Polyphagous in many habitats including gardens. + ++ Recent Introduction. Widespread, a serious garden pest + Frequent and largely synanthropic. A serious garden pest. + ++ ++ (Nb) ? [Ground dwelling on heathlands under heather and ling] Rare ground dwelling esp.on heathlands under heather and ling, Widespread and frequent on hedges and in woods on a variety of hardwood trees Local on hedges and in woods on a variety of hardwood trees esp. alder + Rare ? On trees and shrubs esp. elm. Phyllobius oblongus (Linnaeus, 1758) Phyllobius pomaceus Gyllenhal, 1834 Phyllobius pyri (Linnaeus, 1758) Phyllobius roboretanus Gredler, 1882 Phyllobius viridiaeris (Laicharting, 1781) Liophloeus tessulatus (Müller, 1776) ++ ++ ++ ++ ++ ++ Widespread and frequent on hedges and in woods on a variety of hardwood trees Widespread and frequent on nettles Widespread and frequent on hedges and in woods on a variety of hardwood trees Widespread and frequent on a variety of herbage and on trees Widespread and frequent on a variety of herbage and on trees Local on Ivy clad trees, larvae in Umbellifers. Polydrusus cervinus (Linnaeus, 1758) Polydrusus confluens Stephens, 1831 Polydrusus flavipes (Degeer, 1775) Polydrusus mollis (Stroem, 1768) Polydrusus pterygomalis Boheman, 1840 Polydrusus splendidus (Herbst, 1784) Polydrusus tereticollis (DeGeer, 1775) = undatus (Fabricius, 1781) Barypeithes araneiformis (Schrank, 1781) ++ + + + + ++ Widespread and frequent on hedges and in woods on a variety of hardwood trees (Nb) Rare on broom and gorse esp. on heaths. (Nb) Rare on a variety of trees esp. aspen (Nb) ? On a variety of trees and bushes. Widespread and frequent on hedges and in woods on a variety of hardwood trees (Na) Rare on a variety of trees esp. hazel and oak. Widespread and frequent on hedges and in woods on a variety of hardwood trees esp. birch ++ Widespread on ground in grassy places Otiorhynchus armadillo (Rossi, 1792) Otiorhynchus aurifer Boheman, 1843 Otiorhynchus clavipes (Bonsdorff, 1785) Otiorhynchus crataegi Germar, 1824 Otiorhynchus desertus Rosenhauer, 1847 Otiorhynchus ligneus (Olivier, 1807) Otiorhynchus ovatus (Linnaeus, 1758) Strawberry root weevil Otiorhynchus porcatus (Herbst, 1795) Otiorhynchus raucus (Fabricius., 1777) BrA ge Ro m Otiorhynchus rugifrons (Gyllenhal, 1813) Otiorhynchus rugosostriatus (Goeze, 1777) Rough strawberry weevil Otiorhynchus salcicola Heyden, 1908 Otiorhynchus singularis (Linnaeus, 1767) Clay-coloured weevil Otiorhynchus sulcatus (Fabricius. 1775) Black vine weevil Caenopsis fissirostris (Walton, J., 1847) Caenopsis waltoni (Boheman, 1843) Phyllobius argentatus (Linnaeus, 1758) Phyllobius glaucus (Scopoli, 1793) = calcaratus (Fabricius, 1792) Phyllobius maculicornis Germar, 1824 BrA 84 hge Barypeithes pellucidus (Boheman, 1834) Brachysomus echinatus (Bonsdorff, 1785) Sciaphilus asperatus (Bonsdorff, 1785) Sitona cambricus Stephens, 1831 Sitona cylindricollis (Fahraeus, 1840) Sitona griseus (Fabricius, 1775) Sitona hispidulus (Fabricius, 1777) Sitona humeralis Stephens, 1831 Sitona lepidus Gyllenhal, 1834 Sitona lineatus (Linnaeus, 1758) Pea weevil Sitona macularius (Marsham, 1802) Sitona puberulus Reitter, 1903 Sitona puncticollis Stephens, 1831 Sitona regensteinensis (Herbst, 1797) Sitona striatellus Gyllenhal, 1834 Sitona sulcifrons (Thunberg, 1798) Sitona suturalis Stephens, 1831 Sitona waterhousei Walton, 1846 Tanymecus palliatus (Fabricius, 1787) Trachyphloeus angustisetulus Hansen, 1935 Trachyphloeus alternans Gyllenhal, 1834 Trachyphloeus aristatus (Gyllenhal, 1827) Trachyphloeus asperatus Boheman, 1843 Trachyphloeus bifoveolatus (Beck, 1817) Trachyphloeus digitalis (Gyllenhal, 1827) * Trachyphloeus scabriculus (Linnaeus, 1771) Trachyphloeus spinimanus Germar, 1824 Tropiphorus elevatus (Herbst, 1795) Tropiphorus terricola (Newman, 1838) Hypera arator (Linnaeus, 1758) Hypera fuscocinerea (Marsham, 1802) Hypera meles (Fabricius, 1792) Hypera nigrirostris (Fabricius, 1775) Hypera plantaginis (Degeer, 1775) Hypera pollux (Fabricius, 1801) Hypera postica (Gyllenhal, 1813) Hypera punctata (Fabricius, 1775) Hypera rumicis (Linnaeus, 1758) Hypera suspiciosa (Herbst, 1795) Ro m Sax ++ + + Widespread on ground in grassy places esp. on sand, also in gardens, and common at base of trees. (Nb) Rare, ground dwelling in open grassy places. Local on hernage and trees in many habitats + + + + ++ ++ ++ Local in damp grassland and wetlands on large bird’s-foot trefoil Lotus uliginosus ? On melilots V.local in sandy areas on [Restharrow, birdsfoot, and broom] Frilford etc Widespread in grasslands on clovers and other legumes Local in dry grassland esp. on sand and chalk on legumes esp. medicks Frequent on clovers Very frequent in virtually all situations including gardens (Nb) ? [On legumes] (RDBK) Rare? Local on clovers Locally frequent on gorse and broom Locally frequent on gorse and broom ? On clovers Frequent on vetches Vicia and Lathyrus, in garssalnd and hedgebanks etc. (Nb) Local on bird’s-foot trefoil Lotus corniculatus (Nb) Rare? + ++ ++ ++ + ++ + ++ + + + + + + + > > ++ +++ ++ ++ ++ ++ ++ Local on open disturbed ground esp. on a variety of soils. (Nb) ? On ground in disturbed open areas . (Nb) Rare on open disturbed ground on a variety of dry substrates. (Nb) Rare on open disturbed ground on a variety of dry substrates. Hitchcopse Pit Rare on open disturbed ground on a variety of dry substrates. (Na) Rare in open chalk grassland Rare on open disturbed ground on a variety of dry substrates. Frilfrord (JMC) (Nb) Rare on open disturbed ground on a variety of dry substrates. (Nb) Rare, (Nb) ? [Woodlands often near dog’s mercury Mercurialis perennis] , Rare, declining? On Caryophyllaceae (Stellaria, Cerastium, Spergularia etc.) (Nb) Rare on Medicks. (Na) Rare but increasing on white clover T.repens Local on clovers esp. red clover T.pratense in grassland including roadside verges. Widespread on bird’s-foot trefoil Lotus corniculatus in grassland including roadside verges. V.Local on umbellifers Oenanthe, Apium in wetlands Widespread on various clovers Trifolium spp. and medicks Medicago spp. in grassland including ruderal sites. Local on various clovers Trifolium spp. In grassland (ground dwelling) including ruderal sites. Widespread on docks Rumex spp. ? [On Lotus, Lathyrus and Vicia] 85 Hypera venusta (Fabricius, 1781) + Rare on gorse Ulex esp. U.minor. Limobius borealis (Paykull, 1792) Larinus planus (Fabricius, 1792) Cleonus pigra (Scopoli, 1763) + (Na) Rare? On Geranium species esp. G. pratense. (Nb) ? Extinct? (Nb) Tubney (JJW) Coniocleonus nebulosus (Linnaeus, 1758) Rhinocyllus conicus (Frölich, 1792) Magdalis armigera (Fourcroy, 1785) Magdalis barbicornis (Latreille, 1804) Magdalis carbonaria (Linnaeus, 1758) Magdalis cerasi (Linnaeus, 1758) Magdalis memnonia (Gyllenhal, 1837) Magdalis ruficornis (Linnaeus, 1758) Plum weevil Ro m > Ro m Liparus coronatus (Goeze, 1777) Leiosoma deflexum (Panzer, 1795) Leiosoma oblongulum Boheman, 1842 Leiosoma troglodytes Rye, 1873 = pyrenaeum Brisout, 1866 Hylobius abietis (Linnaeus, 1758) Large pine weevil Lepyrus capucinus (Schaller, 1783) Pissodes castaneus (Degeer, 1775) Small banded pine weevil Trachodes hispidus (Linnaeus, 1758) * Orobitus cyaneus (Linnaeus, 1758) Tanysphyrus lemnae (Paykull, 1792) Duckweed weevil Scolytidae (Bark Beetles) 47 species + 1 sub-fossil Scolytus intricatus (Ratzeburg, 1837) Scolytus mali (Bechstein, 1805) Large fruit tree bark beetle Scolytus multistriatus (Marsham, 1802) Small elm bark beetle Scolytus pygmaeus (Fabricius, 1787) Pygmy elm bark beetle Scolytus rugulosus (Müller, 1818) Small fruit tree bark beetle Scolytus scolytus (Fabricius, 1775) * ++ ++ + ++ ++ ++ + ++ + (Nb) Rare? On cow parsley esp. on chalky soils.) Widespread on Ranunculacae in woods on wood anemone and in damp grasslands on (Nb) Rare on Ranunculacae (wood anemone?). (RDB2) [Downland, woods] ++ Widespread on scot’s pine Pinus sylvestris * ESa x (RDB3) Rare at roots of heather. (Na) Sub fossil only. Neolithic Runnymede Local on elm esp. wych elm Ulmus glabra in hedgerows (Na) Rare on twigs of Rosaceae. (Nb) Rare on birch Betula, boring in dead trunks and branches. (Nb) Local, saproxylic developing in Rosaceae, and oak. Rare on pine including planted amenity trees. Local on Prunus esp. Sloe + (Extinct) [On sallows] not seen in UK since 1897. Local on pine. An important pest species in commercial plantations2 + + (Nb) Rare on dead wood esp. small branches in shady places in old woods. Rare (but easily overlooked?) on violets Viola. ++ Widespread on duckweed Lemna spp. ++ ++ Widespread in branches of hardwoods esp.oak and beech often on recently broken off boughs (Nb) Rare in Prunus and Malus including orchard trees. ++ Local in elm Ulmus + Recent colonist to UK (Heal, 2003), ++ Local in branches and twigs of woody rosaceae esp. sloe. ++ Local in elm 86 Common elm bark beetle Hylesinus crenatus (Fabricius, 1787) Large ash bark beetle Hylesinus oleiperda (Fabricius, 1792) Leperisinus varius (Fabricius, 1775) Common ash bark beetle Leperisinus orni (Fuchs, 1906) Pteleobius vittatus (Fabricius, 1787) Kissophagus hederae (Schmitt, 1843) Hylastinus obscurus (Marsham, 1802) Phloeophthorus rhododactylus (Marsham, 1802) Phloeosinus thujae (Perris, 1855) Hylurgops palliatus (Gyllenhal, 1813) Hylastes angustatus (Herbst, 1793) Hylastes ater (Fabricius, 1792) Hylastes attenuatus Erichson, 1836 Hylastes cunicularius Erichson, 1836 Hylastes opacus Erichson, 1836 Tomicus piniperda (Linnaeus, 1758) Pine shoot beetle Dryocoetinus alni (Georg, 1856) * Dryocoetes autographus (Ratzeburg, 1837) Dryocoetinus villosus (Fabricius, 1792) Xylocleptes bispinus (Duftschmid, 1825) Taphrorychus bicolor (Herbst, 1793) Xyloterus domesticus (Linnaeus, 1758) Ambrosia beetle Xyloterus signatus (Fabricius, 1792) Cryphalus abietis (Ratzeburg, 1837) Cryphalus piceae (Ratzeburg, 1837) Ernoporus caucasicus Lindemann, 1876 Ernoporus fagi (Fabricius, 1798) Ernoporus tiliae (Panzer, 1793) Trypophloeus granulatus (Ratzeburg, 1837) Xyleborus dispar (Fabricius, 1792) Shot-hole borer Xyleborus dryographus (Ratzeburg, 1837) Xyleborus saxeseni (Ratzeburg, 1837) Pityophthorus pubescens (Marsham, 1802) Pityogenes bidentatus (Herbst, 1783) Pityogenes chalcographus (Linnaeus., 1761) Pityogenes trepanatus (Noerdlinger, 1848) Ips sexdentatus (Boerner, 1767) Orthotomicus laricis (Fabricius, 1792) Ro m ++ Local on ash, mainly in bark on larger branches and trunks, [Also on oak and walnut Juglans] ++ ++ Local on ash, in recently dead branches and twigs. Widespread on ash, in slender branches. ++ ++ ++ ++ ++ + ++ + + + + ++ (Nb) Rare on ash, in twigs and small branches. Maybe conspecific with L.. varius (Fabricius, 1775) V.local in twigs of elm Ulmus. (Nb) V.local in dead ivy Hedera helix. Local in dead broom Cytisus and occasionally in gorse stems Local in dead broom Cytisus and occasionally in gorse stems Rare, naturalized on cypress and cedars first found in in 1921. Local on conifers esp. pine ? On pine and spruce Local on pine ? On pine ? [On spruce Picea] Local on pine and occasionally on ash and elm. Frequent on pine, and occasionally Norway spruce Picea. ++ + ++ ++ (Na) [On alder, beech and hazel] Sheffield Bottom 2003 (JSD) Naturalised, in dead and dying spruce Picea. Local in oak Local in traveller’s joy Clematis vitalba, frequent on the chalk. ++ ++ (Nb) V.local in beech, oak and other hardwoods. Local in oak and beech + (Nb) Rare on oak. 1994 ? [On conifers] ? [On conifers] (RDB1) Sub-fossil only. [In limes Tilia ] (Na) Local but widespread on Beech esp. in bark on small twigs. (RDB1) In bark of small-leaved lime Tilia cordata. 1910 Extinct in UK last recorded at in 1867 (Nb) Local in oak esp. sappy exposed heartwood. ++ + + + + + * ++ + (Nb) Rare in hardwoods esp. oak. Local in oak and Beech Local in pine and spruce bark Local in pine and spruce bark ? [In bark of a variety of conifers] , Local in pine bark Local in pine bark esp. on large fallen or felled trees. Padworth Common 2004 (JSD) Local in pine and larch bark 87 Orthotomicus suturalis (Gyllenhal, 1827) Platypodidae 1 Species ++ SUB VCH Platypus cylindrus (Fabricius, 1792) Oak pinhole-borer Local in pine bark 1907 2005 ++ Notes 1907 2005 ++ Notes 1907 2005 Notes (Nb) Local, increasing, in oak, beech, ash, and sweet chestnut esp. large felled trunks. >20 post 1980 recs. STREPSIPTERA Stylopidae SUB VCH Stylops melittae Kirby, W 1802 Halictophagidae Elenchus tenuicornis (Kirby, W 1815) SUB VCH Parasitizes solitary bees, Andrena etc. Parasitizes solitary bees, Andrena etc. 88