JOHN D. ZARDUS
Department of Biology
The Citadel
171 Moultrie Street
Charleston, SC 29407
phone 843-953-7511, fax 843-953-7264
john.zardus@citadel.edu
EDUCATION
1998
1991
1988
Ph.D. (Biology) – Northeastern University, Boston, Massachusetts
M.S. (Zoology) – Brigham Young University, Provo, Utah
B.S. (Zoology) – Brigham Young University, Provo, Utah
RESEARCH INTERESTS
My research interests are anchored in understanding the diversity and function of marine
invertebrates. The scope of my work encompasses aspects of larval biology, field ecology,
population genetics and molecular systematics. I am presently studying the evolution and
ecology of commensal barnacles. A number of barnacle species live attached exclusively to
hosts such as sea turtles and whales. I am exploring how this association is maintained along
with the phylogenetic relationships and life-histories of these commensal crustaceans.
TEACHING EXPERTISE
I specialize in marine biology, invertebrate zoology, and introductory biology. My philosophy
is to teach about life where it lives when possible and within the context of the scientific
method and evolutionary theory. Mentoring, in my view, is integral to teaching and I also
seek to encourage and guide students in their career objectives.
POSITIONS HELD
2005 – present
2001– 2005
1999 – 2001
Assistant Professor, Dept. of Biology, The Citadel, Charleston, SC
Research Fellow, Kewalo Marine Lab, Univ. of Hawaii, Honolulu, HI
Postdoctoral Fellow, Dept. of Biology, Univ. of Mass., Boston, MA
FIELD RESEARCH EXPERIENCE / FACULTY DEVELOPMENT
2009
2008
2001 – 2005
1992 – 1996
1990 & 1995
1994
1993
1989
1987 – 1990
Marine Biological Laboratory, Woods Hole, MA
Gay Head – Bermuda Transect, R/V Endeavor, mid-Atlantic Ocean
Kewalo Marine Lab, Univ. of Hawaii, Honolulu, HI
Friday Harbor Laboratories, Univ. of Washington, Friday Harbor, WA
Harbor Branch Oceanographic Institution, Fort Pierce, FL
Bamfield Marine Station, Univ. of British Columbia, Bamfield, BC
Marine Science Center, Northeastern Univ., Nahant, MA
Hawaii Volcanoes National Park, Volcano, HI
Hopkins Marine Station, Stanford Univ., Pacific Grove, CA
RESEARCH SUPPORT
Extramural
National Science Foundation
(2007-12)
NSF Initiative, Assembling the Tree of Life, collaborative research
Phylogeny on the half-shell - assembling the bivalve tree of life, $25,816.
Hawaii Conservation Alliance
(2002 – 2004) Introduction and dispersal of the Atlantic barnacle Chthamalus proteus in
Hawaii, $10,000.
Intramural
Citadel Foundation
(2009) Bungled barnacles: investigating the taxonomic status of barnacles displaying
morphological plasticity relative to substratum selection, $2,198.
(2008)
Reproduction and larval development of the titan acorn barnacle, Megabalanus
coccopoma, a Pacific invader resident in SC, $2,580.
(2007)
Genetic relationships among barnacles associated with marine mammals, $2,951.
(2006)
Genetic relationships among barnacles associated with sea turtles, $2,944.
(2005)
Morphological diversity among barnacles associated with sea turtles, $2,838.
Star of the West Foundation
(2008) Deep-sea sampling expedition, $1,154.
TEACHING EXPERIENCE
The Citadel, Department of Biology, Charleston, SC
spring 2006-09
fall 2005-09
fall 2005, 08
fall 2006, 09
fall 2007
May 2005-09
Introductory Biology II & Introductory Biology Lab II (for majors)
General Biology (for non-majors)
Marine Biology (for majors)
Freshwater Biology (for majors)
Invertebrate Zoology (for majors)
Tropical Rainforest & Reef Ecology, field course in Central America
College of Charleston, Grice Marine Lab, Charleston, SC
spring 2008
Deep Sea Biology (graduate seminar)
Cornell University–Shoals Marine Lab, Appledore Island, ME
spring 2000, 01
Field Marine Biology & Ecology (undergrad majors)
spring 1995
Field Marine Science II (undergrad majors)
Wheelock College, Boston, MA
spring 1997
Intro. Oceanography (undergraduate non-majors)
PROFESSIONAL AFFILIATIONS
Society for Integrative and Comparative Biology (since 1991)
American Microscopical Society (since 1996)
The Crustacean Society (since 2003)
PROFESSIONAL SERVICE
2009-present
2008
2004-present
2008, 09
2006, 09
2005, 06
Secretary, American Microscopical Society
Member-at-Large, American Microscopical Society
Subject Editor, Marine Biology Research
Proposal Judge, Libbie H. Hyman Scholarship Committee, SICB
Presentation Judge, Grice Marine Lab Graduate Student Colloquium
Student Award Judge, Society for Integrative and Comparative Biology
Publication Peer Reviews:
Aquaculture
Biofouling
Biological Bulletin
Biological Invasions
Fisheries Biology
Invertebrate Biology
Invertebrate Reproduction & Development
Invertebrate Zoology
Journal of Conchology
J. of the Marine Biol. Assoc. of the U.K.
Journal of Morphology
Marine Biology
Marine Biology Research
Marine Ecology Progress Series
Nautilus
INVITED PRESENTATIONS
Fort Johnson Marine Science Seminar Series, James Island, SC (Oct. 2009)
Winthrop University, Department of Biology, Rock Hill, SC (Feb. 2009)
University of Vermont, Department of Biology, Burlington , VT (Oct. 2007)
Grice Marine Lab, Department of Biology, Charleston, SC (Feb. 2005)
Hawaii Institute of Marine Biology, Univ. of Hawaii, Kaneohe, HI (Jun. 2004)
SELECTED PUBLICATIONS
Frick, M.G. and J.D. Zardus (in press). First report of the barnacle Cylindrolepas
darwiniana Pilsbry (Cirripedia: Balanomorpha: Coronuloidea) since it description in 1916
Journal of Crustacean Biology.
Zardus, J.D., B.T. Nedved, Y. Huang, C. Tran, and M.G. Hadfield (2008). Microbial biofilms
facilitate adhesion in biofouling invertebrates. Biological Bulletin 214:91-98.
Zardus, J.D. and G.H. Balazs (2007). Two previously unreported barnacles commensal with the
green sea turtle, Chelonia mydas (Linnaeus, 1758), in Hawaii and a comparison of their attachment
modes. Crustaceana 80:1303-1315.
Zabin, C.J., J.D. Zardus, F. Pitombo, V. Fread and M.G. Hadfield (2007). A tale of three
seas: consistency of natural history traits in a Caribbean-Atlantic barnacle introduced to
Hawaii Biological Invasions 9:523-544.
Zardus, J.D., R.J. Etter, M.R. Chase, M.A. Rex and E.E. Boyle (2006). Bathymetric and
geographic population structure in the pan-Atlantic deep-sea bivalve Deminucula atacellana
(SCHENCK 1939). Molecular Ecology 15:639-651.
Zardus, J.D. and M.G. Hadfield (2005). Multiple origins and incursions of the Atlantic
barnacle Chthamalus proteus in the Pacific. Molecular Ecology 14:3719-3733.
Zardus, J.D. and M.G. Hadfield (2004). Larval development and complemental males in
Chelonibia testudinaria, a barnacle commensal with sea turtles. Journal of Crustacean
Biology 24:409-421.
Boyle, E.E., J.D. Zardus, M.R. Chase, R.J. Etter and M.A. Rex (2004). Strategies for molecular
genetic studies of preserved deep-sea macrofauna. Deep-Sea Research 51:1319-1336.
Zardus, J.D. (2002). Protobranch bivalves. Advances in Marine Biology 42:1-65.
Zardus, J.D. and A. Martel (2001). Phylum Mollusca: Bivalvia. In, Atlas of Marine
Invertebrate Larvae. C.M. Young, M. Sewell & M. Rice eds., pp. 289-325. Academic Press.
Zardus, J.D. and M.P. Morse (1998). Embryogenesis, morphology and ultrastructure of the
pericalymma larva of Acila castrensis (Bivalvia: Protobranchia: Nuculoida). Invertebrate
Biology 117:221-244.
Morse, M.P. and J.D. Zardus (1997). Bivalvia. In, Microscopic Anatomy of Invertebrates
Vol. 6A: Mollusca II. F.W. Harrison and A.J. Kohn eds., pp. 7-118. Wiley-Liss, Inc.