Chapter 15b Hearing and Equilibrium Ear anatomy • external ear external acoustic canal tympanic membrane • middle ear 3 ossicles • inner ear • malleus • incus • stapes – – – cochlea hearing vestibule equilibrium semicircular canals equilibrium HEARING cochlea anatomy • • cochlear duct – – fluid filled membrane fluid endolymph receptor Organ of Corti = spiral organ surrounded by 2 chambers : – – – fluid perilymph scala vestibuli contacts oval window scala tympani contact round window Organ of Corti • • • • = spiral organ – receptors for hearing within cochlear duct basilar membrane support hair cells receptor cells – stereocilia extensions into tectorial membrane tectorial membrane covers stereocilia ; stationary Sound • • • = alternating zones of high and low pressure sound is a vibration waves of vibrating air (or any medium) physics terms • • • • • frequency # waves per second wavelength distance betw crests amplitude height of each wave pitch = frequency – – – hertz (Hz) = cps 20 to 20000 Hz most sensitive range • 1500 – 4000 Hz loudness – – – hertz (Hz) = amplitude decibels (dB) 0 to 120 dB 0 = threshold hearing • • GOAL = change sound into action potential depolarize the receptor cells – open Na channels How? stretch their hair transmission of sound / vibration • • • • • • • sound wave (air) vibrates tympanic membrane 3 ossicles amplify the vibration stapes vibrates the oval window oval window vibrates perilymph perilymph vibrates endolymph endolymph vibrates basilar membrane basilar membrane vibrates hair cells excitation of receptor cells • • • • • • • organ of Corti basilar membrane vibrates hair cells stereocilia stretch against tectorial membrane open K channels K+ (in endolymph) rushes into cell hair cells depolarize hair cells release NT Pitch • • • • • varied thickness of basilar membrane responds to different frequency frequency ~ pitch multiple pitches possible specific areas of auditory cortex Pitch and auditory cortex auditory processing • pitch area of basilar membrane / auditory cortex • loudness number of hair cells stimulated frequency of action potentials • localization where is sound coming from – compares both ears • intensity • timing auditory pathway • to temporal lobe – – – – – cochlear nerve spiral ganglion cochlear nuclei thalamus temporal lobe • • to inferior colliculi – auditory reflex to cerebellum problems • • • deafness – – conduction deafness neural deafness tinnitus Meniere’s syndrome hearing loss tinnitus vertigo EQUILIBRIUM equilibrium anatomy • • • Vestibular apparatus = 2 structures : vestibule linear equilibrium (static) semicircular canals rotational equilibrium (dynamic) receptors • • linear acceleration – – utricle saccule macula receptor mechanism hair cells receptors rotational acceleration semicircular ducts – – ampulla receptor mechanism hair cells receptors macula anatomy • • • • • receptor mechanism in utricle , saccule hair cells stereocilia kinocilium otoliths excitation of macula • • • • position change moves otoliths - bends sterocilia toward kinocilium depolarize away from kinocilium hyperpolarize opposite sides ampulla anatomy • • receptor mechanism of semicircular ducts crista ampullaris – – hair cells cupula excitation of ampulla • • • rotation moves endolymph endolymph deforms cupula cupula bends sterocilia depolarize equilibrium pathway • • • • • hair cells vestibular nerve vestibular nuclei medulla cerebellum reflex movement cerebrum parietal lobe conscious awareness of position and movement equilibrium reflexes • • sensory input from – – – vestibular apparatus proprioception vision response – – eye movment skeletal muscle esp neck and torso • vestibulospinal tract • CN XI problems • • • vertigo nystagmus motion sickness dizziness