FTY 112 – Forests Through Time Discussion: Ecology of Tree Species Species Bigtooth aspen Range in State Entire state Shade Tolerance Intolerant Soil Well drained, deep Jack pine East and north of Bar Harbor Intolerant Paper birch Entire state Intolerant Quaking aspen Entire state Intolerant Excessively well drained sandy, acid, tolerates shallow, sandy/gravelly soils Moderately drained, tolerates poorly drained, nutrient poor sites Well drained, shallow to deep Ash (white) Entire state Intermediate Northern red oak Intermediate White pine South of Bangor Entire state Yellow birch Entire state Intermediate Red maple Entire state Somewhat tolerant Balsam fir North and west of Augusta Tolerant Red spruce North and west of Augusta Tolerant Northern White Cedar Entire state Tolerant to very tolerant American beech Entire state Very tolerant Sugar maple Entire state Very tolerant Hemlock Entire state Most tolerant of all tree species Intermediate Well drained, moist, good nutrients, deep Well drained, tolerates acid soils Well drained, sandy soils, low nutrients Moderately drained, tolerates poorly drained, nutrient poor sites Occupies most varied sites of all tree species Thick organic layer, poor nutrients, tolerates shallow, wet, acid soils Thick organic layer, poor nutrients, tolerates shallow, wet, acid soils Cool, moist, nutrient-rich sites, on organic soils Moderately drained, tolerates dry, shallow soil sites Well drained, moist, good nutrients Well drained to moist, acid to neutral, organic layer Competition Establish on bare soil with full sun; lasts one generation; thin crown Establish on bare soil with full sun; can require fire to open cones Establish on bare soil with full sun; lasts one generation; thin crown Establish on bare soil with full sun; lasts one generation; then crown Establish in shade, grows best in sun; thin crown Can establish in partial shade; needs full sun to survive Establish and grows in partial shade, bare soil Establish on bare soil in canopy openings Can establish in shade, ; prolific sprouting Establish and grow in shade Establish and grow in shade; grows best in sun Establish in shade, especially on rotten logs and stumps Establishes and survives under shade, grows into crown; prolific sprouting Establishes and survives under shade, grows into crown Establish and survive in shade, up to 400yr, needs exposed soil or rotten log All of these hardwood species can sprout from cut stumps or damaged roots. Conifers can not.